51:
29:
557:
The closest relatives of the
Piesmatidae remain rather insufficiently determined. After the ash-grey leaf bugs were recognized as Pentatomomorpha, they were most often placed in the
2674:
2713:
2648:
2687:
731:
703:
Cassis, G., & R.T. Schuh (2010) Systematic methods, fossils, and relationships within
Heteroptera (Insecta). Cladistics 26:262-280.
2692:
2772:
2700:
2557:
2534:
50:
2718:
2627:
2640:
661:
679:
724:
476:
was for some time placed in this family, but it has more recently been suggested that it is a member of the
652:(2007): An Unusual, Primitive Piesmatidae (Insecta: Heteroptera) in Cretaceous Amber from Myanmar (Burma).
2487:
37:
2562:
2739:
666:
251:
526:
but given its distribution it might more comfortably be considered closely related to the ancestor of
2767:
2596:
231:
204:
1629:
2350:
2091:
1851:
1676:
183:
2360:
2049:
2023:
219:
and the firm part of the wings are extensively dimpled. This resembles the similar pattern of the
1650:
1527:
1518:
717:
538:
532:
45:
2744:
460:
305:
582:
499:
472:
436:
419:
409:
334:
2726:
2583:
1310:
1270:
1032:
551:
494:
489:
399:
316:
2679:
2525:
554:) and was initially considered to be a primitive piesmatid but this has since been rejected.
2731:
2393:
2335:
2330:
2164:
2075:
1838:
1616:
1608:
1535:
1318:
1180:
570:
522:
477:
362:
283:
168:
2388:
2355:
2294:
2198:
2193:
2130:
2105:
1846:
1699:
1540:
1504:
1494:
1476:
1412:
1067:
800:
246:
28:
2635:
2622:
2414:
2172:
2054:
1817:
1788:
1590:
1499:
1484:
1355:
1342:
1249:
1220:
1100:
1004:
888:
847:
598:
242:
2761:
2462:
2450:
2422:
2281:
2221:
2203:
2159:
2143:
2028:
2013:
1911:
1660:
1645:
1637:
1371:
1294:
1286:
1196:
1072:
1009:
974:
935:
922:
857:
837:
504:
195:
2588:
2539:
2383:
2234:
2059:
2018:
1980:
1963:
1949:
1937:
1890:
1882:
1728:
1715:
1681:
1489:
1456:
1402:
1278:
1241:
1204:
1159:
984:
909:
901:
812:
784:
341:
227:
2705:
2661:
2519:
2458:
2312:
2247:
2229:
2211:
2180:
2008:
1809:
1557:
1449:
1407:
1397:
1392:
1302:
1233:
1225:
1188:
1057:
1052:
994:
966:
949:
345:
112:
2510:
2345:
2304:
2286:
2067:
1972:
1869:
1861:
1825:
1742:
1384:
1257:
1175:
1062:
1047:
1042:
1019:
927:
914:
852:
842:
821:
558:
543:
466:
447:
372:
323:
277:
224:
143:
122:
2609:
2365:
2325:
2317:
2257:
2242:
2216:
2114:
1988:
1767:
1759:
1723:
1574:
1425:
1417:
1363:
1262:
1149:
979:
772:
760:
741:
709:
603:
590:
586:
566:
562:
516:
352:
267:
191:
180:
172:
102:
82:
62:
2570:
2549:
2504:
2430:
2373:
2273:
2265:
2185:
2122:
2083:
1996:
1903:
1780:
1707:
1595:
1582:
1334:
1326:
1212:
1120:
1112:
1027:
1014:
999:
989:
778:
619:
574:
456:
327:
262:
220:
199:
2575:
230:, and was initially taken to signify a close relationship. It is due to
2378:
2340:
2041:
2033:
1916:
1796:
1037:
547:
2666:
596:
593:. For some time, the Psamminae were even included in the Piesmatidae.
295:
Three subfamilies and eleven genera belong to the family
Piesmatidae:
883:
867:
766:
754:
578:
511:
484:
451:
382:
257:
216:
212:
187:
151:
92:
72:
2653:
2481:
2614:
2454:
1133:
507:
238:
2485:
1446:
1097:
797:
713:
2601:
565:
analysis, with their relatives variously presumed to be the
493:
from the earliest Eocene (about 55 mya) is still a very
1281:(cottony cushion scales, giant coccids, ground pearls)
2461:. Heteroptera with Coleorrhyncha were referred to as
618:
2494:
2413:
2303:
2256:
2142:
2113:
2104:
1971:
1962:
1936:
1902:
1881:
1860:
1837:
1808:
1779:
1758:
1741:
1698:
1659:
1628:
1607:
1573:
1556:
1526:
1517:
1475:
1383:
1354:
1158:
1132:
1111:
965:
948:
900:
866:
820:
811:
179:. The Piesmatidae are distributed mostly in the
2070:(assassin bugs, wheel bugs, thread-legged bugs)
680:"family Piesmatidae Amyot & Serville, 1843"
470:were split off into separate genera. The genus
271:have been found on many host plants but mostly
725:
8:
678:Dellapé, Pablo M.; Henry, Thomas J. (2020).
2482:
2110:
1968:
1933:
1755:
1695:
1570:
1523:
1472:
1468:
1443:
1108:
1094:
962:
817:
808:
794:
732:
718:
710:
650:Grimaldi, David A. & Engel, Michael S.
27:
18:
645:
643:
641:
639:
637:
635:
614:. However a cladistic analysis rejected
198:. A common species found throughout the
631:
215:, some 2–4 mm overall. The head,
7:
2641:8b1c5ee9-1081-4d95-a531-21113a6c2c7d
2224:(jewel bugs or metallic shield bugs)
664::10.1206/0003-0082(2008)36112.0.CO;2
2036:(plant bugs, leaf bugs, grass bugs)
1983:(minute pirate bugs or flower bugs)
1640:(marsh treaders or water measurers)
446:Many species were formerly in the
137:Amyot & Audinet-Serville, 1843
14:
2175:(giant shield bugs and relatives)
930:(typical treehoppers, thorn bugs)
514:might be a northern relative of
321:Nel, Waller & De Ploeg, 2004
49:
2368:(milkweed bugs, true seed bugs)
2276:(squash bugs, leaf-footed bugs)
1543:(unique-headed bugs, gnat bugs)
387:Lepelitier & Serville, 1825
157:Lepelitier & Serville, 1825
1799:(water scorpions, needle bugs)
536:was placed within the family.
1:
211:Ash-grey leaf bugs are small
2425:(red bugs, cotton stainers)
404:Breddin In Schumacher, 1913
2789:
2446:
1932:
1694:
1471:
1467:
1442:
1252:(pupillarial palm scales)
1107:
1093:
807:
793:
749:
654:American Museum Novitates
550:and lived about 100 mya (
186:, with some occurring in
149:
142:
46:Scientific classification
44:
35:
26:
21:
1366:(woolly conifer aphids)
684:Lygaeoidea Species File
2289:(scentless plant bugs)
1289:(Mediterranean scales)
610:with the discovery of
497:member of the family.
2740:Paleobiology Database
2433:(bordered plant bugs)
2206:(includes ebony bugs)
1872:(creeping water bugs)
281:. The host plants of
2773:Heteroptera families
2636:Fauna Europaea (new)
2404:(ash-grey leaf bugs)
2396:(atypical seed bugs)
2062:(old world bat bugs)
1991:(bed bugs, bat bugs)
1919:(pygmy backswimmers)
1828:(velvety shore bugs)
1420:(jumping plant lice)
530:. From 2007 to 2009
232:convergent evolution
38:Parapiesma quadratum
2268:(broad-headed bugs)
1619:(velvet water bugs)
1207:(ornate pit scales)
577:, or the peculiar,
255:have been found on
184:Northern Hemisphere
1791:(giant water bugs)
1718:(spiny shore bugs)
1651:Paraphrynoveliidae
1564:(semiaquatic bugs)
1519:Enicocephalomorpha
1313:(phenacoleachiids)
1311:Phenacoleachiidae
1273:(false pit scales)
1271:Lecanodiaspididae
177:ash-grey leaf bugs
175:, commonly called
2755:
2754:
2727:Open Tree of Life
2488:Taxon identifiers
2479:
2478:
2475:
2474:
2471:
2470:
2466:
2442:
2441:
2434:
2426:
2405:
2397:
2394:Rhyparochromidae
2369:
2321:
2290:
2277:
2269:
2238:
2225:
2207:
2189:
2176:
2168:
2165:Acanthosomatidae
2152:
2134:
2126:
2100:
2099:
2087:
2079:
2078:(royal palm bugs)
2076:Thaumastocoridae
2071:
2063:
2045:
2037:
2004:
1992:
1984:
1958:
1957:
1928:
1927:
1920:
1894:
1873:
1829:
1821:
1800:
1792:
1771:
1751:
1749:(true water bugs)
1737:
1736:
1719:
1711:
1690:
1689:
1669:
1641:
1620:
1599:
1586:
1566:
1552:
1551:
1544:
1513:
1512:
1462:
1438:
1437:
1434:
1433:
1421:
1375:
1367:
1346:
1338:
1337:(giant mealybugs)
1330:
1322:
1319:Phoenicococcidae
1314:
1306:
1298:
1290:
1282:
1274:
1266:
1253:
1245:
1237:
1229:
1216:
1208:
1200:
1192:
1184:
1181:Asterolecaniidae
1168:
1142:
1124:
1089:
1088:
1085:
1084:
1081:
1080:
1033:Hypochthonellidae
1023:
958:
944:
943:
931:
918:
892:
876:
830:
552:million years ago
542:was found in mid-
455:, from which the
252:Piesma linnavouri
162:
161:
138:
16:Family of insects
2780:
2748:
2747:
2735:
2734:
2722:
2721:
2709:
2708:
2706:NBNSYS0000160715
2696:
2695:
2683:
2682:
2670:
2669:
2657:
2656:
2644:
2643:
2631:
2630:
2618:
2617:
2605:
2604:
2592:
2591:
2579:
2578:
2566:
2565:
2553:
2552:
2543:
2542:
2530:
2529:
2528:
2515:
2514:
2513:
2483:
2453:are a different
2448:
2432:
2424:
2403:
2395:
2367:
2336:Cryptorhamphidae
2331:Colobathristidae
2319:
2288:
2275:
2267:
2236:
2223:
2205:
2188:(burrowing bugs)
2187:
2174:
2166:
2153:
2151:
2148:
2132:
2131:Termitaphididae
2124:
2111:
2085:
2077:
2069:
2061:
2043:
2035:
2001:Curalium cronini
1998:
1990:
1982:
1969:
1942:
1934:
1918:
1892:
1871:
1839:Aphelocheiroidea
1827:
1819:
1798:
1790:
1769:
1756:
1752:
1750:
1747:
1717:
1709:
1696:
1670:
1668:
1667:(water treaders)
1665:
1639:
1618:
1597:
1585:(water striders)
1584:
1571:
1567:
1565:
1562:
1542:
1541:Enicocephalidae
1536:Aenictopecheidae
1528:Enicocephaloidea
1524:
1473:
1469:
1461:
1455:
1452:
1444:
1419:
1373:
1365:
1344:
1336:
1328:
1320:
1312:
1304:
1296:
1288:
1280:
1272:
1264:
1251:
1243:
1236:(armored scales)
1235:
1227:
1214:
1206:
1198:
1190:
1182:
1169:
1167:
1164:
1143:
1141:
1138:
1122:
1109:
1095:
1021:
963:
959:
957:
954:
929:
916:
890:
877:
875:
872:
831:
829:
826:
818:
809:
795:
734:
727:
720:
711:
704:
701:
695:
694:
692:
690:
675:
669:
665:
651:
647:
571:Colobathristidae
478:Thaumastocoridae
442:
425:
415:
405:
388:
378:
368:
358:
340:
322:
311:
287:remain unknown.
136:
54:
53:
31:
19:
2788:
2787:
2783:
2782:
2781:
2779:
2778:
2777:
2758:
2757:
2756:
2751:
2743:
2738:
2730:
2725:
2717:
2712:
2704:
2699:
2691:
2686:
2678:
2673:
2665:
2660:
2652:
2647:
2639:
2634:
2626:
2621:
2613:
2608:
2600:
2595:
2587:
2582:
2574:
2569:
2561:
2556:
2548:
2546:
2538:
2533:
2524:
2523:
2518:
2509:
2508:
2503:
2490:
2480:
2467:
2438:
2409:
2389:Pachygronthidae
2356:Heterogastridae
2299:
2295:Stenocephalidae
2252:
2199:Parastrachiidae
2194:Thaumastellidae
2173:Tessaratomidae
2149:
2147:
2146:
2138:
2106:Pentatomomorpha
2096:
1954:
1941:(Coleorrhyncha)
1940:
1939:
1938:Peloridiomorpha
1924:
1898:
1877:
1856:
1847:Aphelocheiridae
1833:
1818:Gelastocoridae
1804:
1789:Belostomatidae
1775:
1770:(water boatmen)
1748:
1746:
1745:
1733:
1700:Leptopodomorpha
1686:
1666:
1664:
1663:
1655:
1624:
1603:
1563:
1561:
1560:
1548:
1509:
1505:Stemmocryptidae
1495:Hypsipterygidae
1477:Dipsocoromorpha
1463:
1459:
1453:
1430:
1413:Phacopteronidae
1379:
1350:
1345:(stictococcids)
1343:Stictococcidae
1327:Pseudococcidae
1305:(ensign scales)
1250:Halimococcidae
1166:(scale insects)
1165:
1163:
1162:
1154:
1139:
1137:
1136:
1128:
1103:
1077:
1068:Tettigometridae
955:
953:
952:
940:
896:
891:(hairy cicadas)
889:Tettigarctidae
873:
871:
870:
862:
827:
825:
824:
803:
801:Auchenorrhyncha
789:
745:
738:
708:
707:
702:
698:
688:
686:
677:
676:
672:
660:
649:
648:
633:
628:
602:or placed in a
440:
423:
413:
403:
395:
386:
376:
366:
356:
338:
320:
309:
301:
293:
247:Caryophyllaceae
241:sap, mostly of
205:Piesma cinereum
158:
155:
135:
48:
17:
12:
11:
5:
2786:
2784:
2776:
2775:
2770:
2760:
2759:
2753:
2752:
2750:
2749:
2736:
2723:
2710:
2697:
2684:
2671:
2658:
2645:
2632:
2623:Fauna Europaea
2619:
2606:
2593:
2580:
2567:
2554:
2544:
2531:
2516:
2500:
2498:
2492:
2491:
2486:
2477:
2476:
2473:
2472:
2469:
2468:
2447:
2444:
2443:
2440:
2439:
2437:
2436:
2428:
2423:Pyrrhocoridae
2419:
2417:
2415:Pyrrhocoroidea
2411:
2410:
2408:
2407:
2399:
2391:
2386:
2381:
2376:
2371:
2363:
2358:
2353:
2348:
2343:
2338:
2333:
2328:
2323:
2315:
2309:
2307:
2301:
2300:
2298:
2297:
2292:
2284:
2279:
2271:
2262:
2260:
2254:
2253:
2251:
2250:
2245:
2240:
2232:
2227:
2222:Scutelleridae
2219:
2214:
2209:
2204:Thyreocoridae
2201:
2196:
2191:
2183:
2178:
2170:
2162:
2156:
2154:
2140:
2139:
2137:
2136:
2133:(termite bugs)
2128:
2119:
2117:
2108:
2102:
2101:
2098:
2097:
2095:
2094:
2089:
2081:
2073:
2065:
2057:
2055:Plokiophilidae
2052:
2047:
2039:
2031:
2026:
2021:
2016:
2011:
2006:
1994:
1986:
1977:
1975:
1966:
1960:
1959:
1956:
1955:
1953:
1952:
1946:
1944:
1930:
1929:
1926:
1925:
1923:
1922:
1914:
1908:
1906:
1900:
1899:
1897:
1896:
1893:(backswimmers)
1887:
1885:
1879:
1878:
1876:
1875:
1866:
1864:
1858:
1857:
1855:
1854:
1849:
1843:
1841:
1835:
1834:
1832:
1831:
1823:
1814:
1812:
1806:
1805:
1803:
1802:
1794:
1785:
1783:
1777:
1776:
1774:
1773:
1764:
1762:
1753:
1739:
1738:
1735:
1734:
1732:
1731:
1726:
1721:
1713:
1704:
1702:
1692:
1691:
1688:
1687:
1685:
1684:
1679:
1673:
1671:
1657:
1656:
1654:
1653:
1648:
1643:
1638:Hydrometridae
1634:
1632:
1630:Hydrometroidea
1626:
1625:
1623:
1622:
1613:
1611:
1605:
1604:
1602:
1601:
1593:
1591:Hermatobatidae
1588:
1579:
1577:
1568:
1554:
1553:
1550:
1549:
1547:
1546:
1538:
1532:
1530:
1521:
1515:
1514:
1511:
1510:
1508:
1507:
1502:
1500:Schizopteridae
1497:
1492:
1487:
1485:Ceratocombidae
1481:
1479:
1465:
1464:
1447:
1440:
1439:
1436:
1435:
1432:
1431:
1429:
1428:
1423:
1415:
1410:
1405:
1400:
1395:
1389:
1387:
1381:
1380:
1378:
1377:
1374:(phylloxerans)
1372:Phylloxeridae
1369:
1360:
1358:
1356:Phylloxeroidea
1352:
1351:
1349:
1348:
1340:
1332:
1324:
1316:
1308:
1300:
1297:(giant scales)
1295:Monophlebidae
1292:
1287:Micrococcidae
1284:
1276:
1268:
1260:
1255:
1247:
1239:
1231:
1226:Dactylopiidae
1223:
1221:Conchaspididae
1218:
1210:
1202:
1199:(carayonemids)
1197:Carayonemidae
1194:
1186:
1178:
1172:
1170:
1156:
1155:
1153:
1152:
1146:
1144:
1130:
1129:
1127:
1126:
1117:
1115:
1105:
1104:
1101:Sternorrhyncha
1098:
1091:
1090:
1087:
1086:
1083:
1082:
1079:
1078:
1076:
1075:
1070:
1065:
1060:
1055:
1050:
1045:
1040:
1035:
1030:
1025:
1022:(lanternflies)
1017:
1012:
1007:
1005:Dictyopharidae
1002:
997:
992:
987:
982:
977:
971:
969:
960:
956:(planthoppers)
946:
945:
942:
941:
939:
938:
933:
925:
920:
912:
906:
904:
898:
897:
895:
894:
886:
880:
878:
864:
863:
861:
860:
855:
850:
848:Clastopteridae
845:
840:
834:
832:
815:
805:
804:
798:
791:
790:
788:
787:
781:
775:
769:
763:
757:
750:
747:
746:
739:
737:
736:
729:
722:
714:
706:
705:
696:
670:
630:
629:
627:
624:
599:incertae sedis
483:There are two
444:
443:
441:Kormilev, 1969
432:
431:
427:
426:
424:Bergroth, 1921
416:
406:
394:
391:
390:
389:
379:
377:Pericart, 1974
369:
359:
349:
330:
312:
310:Pericart, 1974
300:
297:
292:
289:
243:Chenopodiaceae
160:
159:
156:
147:
146:
140:
139:
130:
126:
125:
120:
116:
115:
110:
106:
105:
100:
96:
95:
90:
86:
85:
80:
76:
75:
70:
66:
65:
60:
56:
55:
42:
41:
33:
32:
24:
23:
15:
13:
10:
9:
6:
4:
3:
2:
2785:
2774:
2771:
2769:
2766:
2765:
2763:
2746:
2741:
2737:
2733:
2728:
2724:
2720:
2715:
2711:
2707:
2702:
2698:
2694:
2689:
2685:
2681:
2676:
2672:
2668:
2663:
2659:
2655:
2650:
2646:
2642:
2637:
2633:
2629:
2624:
2620:
2616:
2611:
2607:
2603:
2598:
2594:
2590:
2585:
2581:
2577:
2572:
2568:
2564:
2559:
2555:
2551:
2545:
2541:
2536:
2532:
2527:
2521:
2517:
2512:
2506:
2502:
2501:
2499:
2497:
2493:
2489:
2484:
2464:
2463:Prosorrhyncha
2460:
2456:
2452:
2451:Coleorrhyncha
2445:
2435:
2429:
2427:
2421:
2420:
2418:
2416:
2412:
2406:
2400:
2398:
2392:
2390:
2387:
2385:
2382:
2380:
2377:
2375:
2372:
2370:
2364:
2362:
2359:
2357:
2354:
2352:
2351:Henicocoridae
2349:
2347:
2344:
2342:
2339:
2337:
2334:
2332:
2329:
2327:
2324:
2322:
2316:
2314:
2311:
2310:
2308:
2306:
2302:
2296:
2293:
2291:
2285:
2283:
2282:Hyocephalidae
2280:
2278:
2272:
2270:
2264:
2263:
2261:
2259:
2255:
2249:
2246:
2244:
2241:
2239:
2235:Pentatomidae
2233:
2231:
2228:
2226:
2220:
2218:
2215:
2213:
2210:
2208:
2202:
2200:
2197:
2195:
2192:
2190:
2184:
2182:
2179:
2177:
2171:
2169:
2167:(shield bugs)
2163:
2161:
2160:Urostylididae
2158:
2157:
2155:
2150:(shield bugs)
2145:
2144:Pentatomoidea
2141:
2135:
2129:
2127:
2121:
2120:
2118:
2116:
2112:
2109:
2107:
2103:
2093:
2092:Velocipedidae
2090:
2088:
2082:
2080:
2074:
2072:
2066:
2064:
2060:Polyctenidae
2058:
2056:
2053:
2051:
2048:
2046:
2044:(damsel bugs)
2040:
2038:
2032:
2030:
2029:Microphysidae
2027:
2025:
2022:
2020:
2017:
2015:
2014:Lasiochilidae
2012:
2010:
2007:
2005:
2002:
1995:
1993:
1987:
1985:
1981:Anthocoridae
1979:
1978:
1976:
1974:
1970:
1967:
1965:
1961:
1951:
1948:
1947:
1945:
1943:
1935:
1931:
1921:
1915:
1913:
1912:Helotrephidae
1910:
1909:
1907:
1905:
1901:
1895:
1891:Notonectidae
1889:
1888:
1886:
1884:
1883:Notonectoidea
1880:
1874:
1868:
1867:
1865:
1863:
1859:
1853:
1852:Potamocoridae
1850:
1848:
1845:
1844:
1842:
1840:
1836:
1830:
1824:
1822:
1816:
1815:
1813:
1811:
1807:
1801:
1795:
1793:
1787:
1786:
1784:
1782:
1778:
1772:
1766:
1765:
1763:
1761:
1757:
1754:
1744:
1740:
1730:
1727:
1725:
1722:
1720:
1716:Leptopodidae
1714:
1712:
1706:
1705:
1703:
1701:
1697:
1693:
1683:
1680:
1678:
1677:Madeoveliidae
1675:
1674:
1672:
1662:
1661:Mesovelioidea
1658:
1652:
1649:
1647:
1646:Macroveliidae
1644:
1642:
1636:
1635:
1633:
1631:
1627:
1621:
1615:
1614:
1612:
1610:
1606:
1600:
1598:(riffle bugs)
1594:
1592:
1589:
1587:
1581:
1580:
1578:
1576:
1572:
1569:
1559:
1555:
1545:
1539:
1537:
1534:
1533:
1531:
1529:
1525:
1522:
1520:
1516:
1506:
1503:
1501:
1498:
1496:
1493:
1491:
1488:
1486:
1483:
1482:
1480:
1478:
1474:
1470:
1466:
1458:
1457:Coleorrhyncha
1451:
1445:
1441:
1427:
1424:
1422:
1416:
1414:
1411:
1409:
1406:
1404:
1401:
1399:
1396:
1394:
1391:
1390:
1388:
1386:
1382:
1376:
1370:
1368:
1362:
1361:
1359:
1357:
1353:
1347:
1341:
1339:
1333:
1331:
1325:
1323:
1321:(palm scales)
1317:
1315:
1309:
1307:
1301:
1299:
1293:
1291:
1285:
1283:
1279:Margarodidae
1277:
1275:
1269:
1267:
1261:
1259:
1256:
1254:
1248:
1246:
1244:(felt scales)
1242:Eriococcidae
1240:
1238:
1232:
1230:
1224:
1222:
1219:
1217:
1215:(soft scales)
1211:
1209:
1205:Cerococcidae
1203:
1201:
1195:
1193:
1187:
1185:
1179:
1177:
1174:
1173:
1171:
1161:
1157:
1151:
1148:
1147:
1145:
1135:
1131:
1125:
1119:
1118:
1116:
1114:
1110:
1106:
1102:
1096:
1092:
1074:
1073:Tropiduchidae
1071:
1069:
1066:
1064:
1061:
1059:
1056:
1054:
1051:
1049:
1046:
1044:
1041:
1039:
1036:
1034:
1031:
1029:
1026:
1024:
1018:
1016:
1013:
1011:
1010:Eurybrachidae
1008:
1006:
1003:
1001:
998:
996:
993:
991:
988:
986:
983:
981:
978:
976:
975:Acanaloniidae
973:
972:
970:
968:
964:
961:
951:
950:Fulgoromorpha
947:
937:
936:Myerslopiidae
934:
932:
926:
924:
923:Melizoderidae
921:
919:
917:(leafhoppers)
915:Cicadellidae
913:
911:
908:
907:
905:
903:
899:
893:
887:
885:
882:
881:
879:
869:
865:
859:
858:Machaerotidae
856:
854:
851:
849:
846:
844:
841:
839:
838:Aphrophoridae
836:
835:
833:
828:(froghoppers)
823:
819:
816:
814:
810:
806:
802:
796:
792:
786:
782:
780:
776:
774:
770:
768:
764:
762:
758:
756:
752:
751:
748:
743:
735:
730:
728:
723:
721:
716:
715:
712:
700:
697:
685:
681:
674:
671:
668:
663:
658:
655:
646:
644:
642:
640:
638:
636:
632:
625:
623:
621:
617:
613:
609:
605:
601:
600:
594:
592:
588:
584:
580:
576:
572:
568:
564:
560:
555:
553:
549:
545:
541:
540:
535:
534:
529:
525:
524:
519:
518:
513:
509:
506:
502:
501:
496:
492:
491:
486:
481:
479:
475:
474:
469:
468:
463:
462:
458:
454:
453:
449:
439:
438:
434:
433:
429:
428:
422:
421:
417:
412:
411:
407:
402:
401:
397:
396:
392:
385:
384:
380:
375:
374:
370:
365:
364:
360:
355:
354:
350:
347:
343:
337:
336:
331:
329:
325:
319:
318:
313:
308:
307:
303:
302:
298:
296:
290:
288:
286:
285:
280:
279:
274:
270:
269:
264:
260:
259:
254:
253:
248:
244:
240:
237:They feed on
235:
233:
229:
226:
222:
218:
214:
209:
207:
206:
201:
197:
196:South America
193:
189:
185:
182:
178:
174:
170:
166:
154:
153:
148:
145:
141:
134:
131:
128:
127:
124:
121:
119:Superfamily:
118:
117:
114:
111:
108:
107:
104:
101:
98:
97:
94:
91:
88:
87:
84:
81:
78:
77:
74:
71:
68:
67:
64:
61:
58:
57:
52:
47:
43:
40:
39:
34:
30:
25:
20:
2495:
2402:Piesmatidae
2401:
2384:Oxycarenidae
2361:Idiostolidae
2320:(stilt bugs)
2237:(stink bugs)
2050:Pachynomidae
2024:Medocostidae
2019:Lyctocoridae
2000:
1964:Cimicomorpha
1950:Peloridiidae
1729:Aepophilidae
1710:(shore bugs)
1682:Mesoveliidae
1490:Dipsocoridae
1403:Carsidaridae
1303:Ortheziidae
1265:(lac scales)
1234:Diaspididae
1228:(cochineals)
1191:(beesoniids)
1189:Beesoniidae
1183:(pit scales)
1123:(whiteflies)
1121:Aleyrodidae
1113:Aleyrodoidea
985:Achilixiidae
928:Membracidae
910:Aetalionidae
902:Membracoidea
813:Cicadomorpha
785:Paraneoptera
783:Superorder:
777:Infraclass:
699:
687:. Retrieved
683:
673:
667:PDF fulltext
656:
653:
615:
611:
608:Piesmatoidea
607:
606:superfamily
597:
595:
556:
537:
531:
527:
521:
515:
498:
488:
482:
471:
465:
459:
450:
445:
435:
418:
414:Slater, 1970
408:
398:
381:
371:
361:
351:
342:Baltic amber
333:
315:
304:
294:
282:
276:
272:
266:
256:
250:
236:
228:Cimicomorpha
210:
203:
176:
164:
163:
150:
132:
36:
22:Piesmatidae
2768:Piesmatidae
2662:iNaturalist
2540:PIESMATIDAE
2526:Piesmatidae
2520:Wikispecies
2496:Piesmatidae
2459:Heteroptera
2313:Artheneidae
2287:Rhopalidae
2248:Megarididae
2230:Plataspidae
2212:Lestoniidae
2181:Dinidoridae
2125:(flat bugs)
2086:(lace bugs)
2068:Reduviidae
2009:Joppeicidae
1997:Curaliidae
1870:Naucoridae
1862:Naucoroidea
1826:Ochteridae
1820:(toad bugs)
1810:Ochteroidea
1558:Gerromorpha
1450:Heteroptera
1408:Homotomidae
1398:Calophyidae
1393:Aphalaridae
1329:(mealybugs)
1058:Nogodinidae
1053:Meenoplidae
1020:Fulgoridae
995:Delphacidae
967:Fulgoroidea
822:Cercopoidea
616:Cretopiesma
612:Cretopiesma
546:amber from
539:Cretopiesma
533:Cretopiesma
503:, found in
430:Thaicorniae
367:Drake, 1948
357:Drake, 1924
346:Rovno amber
339:Popov, 2001
299:Piesmatinae
167:is a small
165:Piesmatidae
133:Piesmatidae
113:Heteroptera
2762:Categories
2366:Lygaeidae
2346:Geocoridae
2318:Berytidae
2305:Lygaeoidea
1989:Cimicidae
1973:Cimicoidea
1768:Corixidae
1760:Corixoidea
1743:Nepomorpha
1418:Psyllidae
1385:Psylloidea
1364:Adelgidae
1263:Kerriidae
1258:Kermesidae
1176:Aclerdidae
1134:Aphidoidea
1063:Ricaniidae
1048:Lophopidae
1043:Kinnaridae
868:Cicadoidea
853:Epipygidae
843:Cercopidae
771:Subclass:
761:Arthropoda
626:References
559:Lygaeoidea
544:Cretaceous
467:Parapiesma
461:Afropiesma
448:type genus
373:Parapiesma
326:, France,
324:Oise amber
306:Afropiesma
278:Proteaceae
225:infraorder
144:Type genus
123:Lygaeoidea
109:Suborder:
83:Arthropoda
2431:Largidae
2326:Blissidae
2274:Coreidae
2266:Alydidae
2258:Coreoidea
2243:Canopidae
2217:Phloeidae
2186:Cydnidae
2123:Aradidae
2115:Aradoidea
2084:Tingidae
1724:Omaniidae
1708:Saldidae
1617:Hebridae
1609:Hebroidea
1596:Veliidae
1583:Gerridae
1575:Gerroidea
1448:Suborder
1426:Triozidae
1335:Putoidae
1213:Coccidae
1160:Coccoidea
1150:Aphididae
1099:Suborder
980:Achilidae
884:Cicadidae
874:(cicadas)
799:Suborder
773:Pterygota
753:Kingdom:
742:Hemiptera
689:29 August
604:monotypic
591:Lygaeidae
587:subfamily
583:Psamminae
567:Berytidae
563:cladistic
561:based on
517:Mcateella
510:from the
500:Heissiana
473:Thaicoris
457:subgenera
437:Thaicoris
420:Sympeplus
410:Saxicoris
393:Psamminae
353:Mcateella
335:Heissiana
268:Mcateella
234:however.
192:Australia
181:temperate
173:true bugs
103:Hemiptera
69:Kingdom:
63:Eukaryota
2571:BugGuide
2547:BioLib:
2511:Q1950799
2505:Wikidata
2374:Malcidae
2042:Nabidae
2034:Miridae
1917:Pleidae
1904:Pleoidea
1797:Nepidae
1781:Nepoidea
1140:(aphids)
1028:Gengidae
1015:Flatidae
1000:Derbidae
990:Cixiidae
779:Neoptera
759:Phylum:
755:Animalia
744:families
659:: 1-17.
620:Aradidae
575:Malcidae
528:Eopiesma
490:Eopiesma
487:genera.
400:Psammium
348:, Eocene
328:Ypresian
317:Eopiesma
291:Taxonomy
263:Fabaceae
221:Tingidae
200:Americas
129:Family:
79:Phylum:
73:Animalia
59:Domain:
2379:Ninidae
2341:Cymidae
1038:Issidae
767:Insecta
765:Class:
740:Extant
589:of the
548:Myanmar
223:of the
213:insects
99:Order:
93:Insecta
89:Class:
2745:218539
2732:568266
2719:236392
2693:107629
2680:100726
2667:343428
2615:1PIESF
2576:247450
2563:478458
2449:Note:
581:-like
579:beetle
523:Miespa
512:Eocene
505:Baltic
485:fossil
452:Piesma
383:Piesma
363:Miespa
284:Miespa
273:Acacia
258:Acacia
217:thorax
188:Africa
169:family
152:Piesma
2675:IRMNG
2628:12841
2589:8NKPC
2550:17128
2457:from
2455:clade
1454:(with
508:amber
495:basal
239:plant
2714:NCBI
2688:ITIS
2654:9651
2649:GBIF
2610:EPPO
2558:BOLD
691:2020
657:3611
585:, a
573:and
520:and
464:and
275:and
245:and
194:and
2701:NBN
2602:625
2597:EoL
2584:CoL
2535:AFD
662:DOI
265:).
202:is
171:of
2764::
2742::
2729::
2716::
2703::
2690::
2677::
2664::
2651::
2638::
2625::
2612::
2599::
2586::
2573::
2560::
2537::
2522::
2507::
682:.
634:^
622:.
569:,
480:.
344:,
332:†
314:†
249:.
208:.
190:,
2465:.
2003:)
1999:(
1460:)
733:e
726:t
719:v
693:.
261:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
