66:
44:
5007:
5255:
5243:
5279:
5267:
529:
467:. The amoebas started dying and when examined were found to contain giant virus specimens. The authors said they got the idea to probe permafrost samples for new viruses after reading about an experiment that revived a similar aged seed of
820:
744:
814:
Legendre, M.; Bartoli, J.; Shmakova, L.; Jeudy, S.; Labadie, K.; Adrait, A.; Lescot, M.; Poirot, O.; Bertaux, L.; Bruley, C.; Coute, Y.; Rivkina, E.; Abergel, C.; Claverie, J.-M. (March 2014).
1106:
517:
of the genome has been estimated to be 2.23 × 10 substitutions/site/year. The authors have suggested that the two known
Alphapithoviruses diverged around two hundred thousand years ago.
494:
drilling operations could lead to previously undiscovered and potentially pathogenic viruses being unearthed. However, other scientists dispute that this scenario poses a real threat.
947:
479:
1099:
5348:
5376:
5381:
752:
1092:
5173:
4931:
939:
227:
767:
552:
5283:
273:) in length and 0.5 μm (500 nm) in diameter, making it the largest virus yet found. It is 50% larger in size than the
793:
5386:
65:
962:
486:
Although the virus is harmless to humans, its viability after being frozen for millennia has raised concerns that global
2892:
988:
5353:
1069:
4924:
4636:
4141:
3215:
4965:
218:
179:
1115:
456:
208:
186:
52:
4960:
5339:
816:"Thirty-thousand-year-old distant relative of giant icosahedral DNA viruses with a pandoravirus morphology"
425:(ORFs), which translate into 467 different proteins. The genome encodes all the proteins needed to produce
5301:
5166:
4917:
3403:
340:
genome sequence reveals that it is barely related to that virus, but more closely resembles members of
3877:
5218:
5213:
4454:
3976:
3171:
1450:
1119:
828:
572:
60:
4273:
3941:
3884:
3856:
3828:
3498:
3182:
3113:
2373:
2366:
1457:
1415:
1408:
322:, more than a typical virus, but far fewer than the 2556 putative protein-coding sequences found in
5271:
5208:
5193:
4955:
4468:
4322:
4101:
3849:
3842:
2662:
1860:
1478:
1464:
1422:
1335:
5242:
4560:
4511:
4461:
4440:
3891:
1401:
4280:
3353:
3346:
2419:
2054:
2040:
1632:
641:
469:
422:
114:
875:
5183:
5028:
4576:
3535:
3410:
3389:
3282:
2938:
2596:
2238:
1801:
1394:
1321:
1039:
895:
856:
718:
687:
Levasseur A, Andreani J, Delerce J, Bou Khalil J, Robert C, La Scola B, Raoult D (July 2016).
547:
43:
689:"Comparison of a modern and fossil Pithovirus reveals its genetic conservation and evolution"
5259:
5079:
5015:
4366:
4294:
4187:
3762:
3704:
3667:
3396:
3127:
3085:
3033:
2996:
2966:
2927:
2781:
2707:
2564:
1886:
1849:
1823:
1781:
1582:
1572:
1387:
1239:
1210:
1029:
1019:
887:
846:
836:
708:
700:
633:
342:
328:. Thus, its genome is far less densely packed than any other known virus. Two-thirds of its
1060:
5089:
5074:
5069:
5053:
5006:
4826:
4697:
4532:
4480:
4393:
4349:
4338:
4176:
4028:
3969:
3793:
3514:
3444:
3332:
3315:
3250:
2847:
2802:
2523:
2030:
1812:
1723:
1643:
1539:
662:
557:
303:
has a thick, oval wall with an opening at one end. Internally, its structure resembles a
832:
666:
5247:
5178:
5146:
5033:
4991:
4869:
4847:
4815:
4780:
4686:
4651:
4587:
4430:
4419:
4382:
4224:
4108:
3998:
3927:
3917:
3678:
3470:
3433:
3339:
3134:
2718:
2691:
2338:
2327:
2231:
2196:
2189:
2175:
2168:
2117:
2089:
2019:
2008:
1973:
1757:
1560:
1429:
1366:
1300:
1278:
1200:
1034:
1007:
913:
874:
Queiroz, Victória F.; Rodrigues, Rodrigo A. L.; Abrahão, Jônatas Santos (1 June 2024).
851:
815:
713:
688:
534:
487:
275:
138:
459:. The virus was discovered buried 30 m (100 ft) below the surface of a late
5370:
5198:
5110:
5043:
4996:
4837:
4732:
4718:
4546:
4539:
4518:
4359:
4308:
4301:
4165:
4061:
4039:
4012:
3988:
3948:
3905:
3870:
3863:
3835:
3821:
3814:
3715:
3651:
3622:
3601:
3580:
3569:
3549:
3481:
3303:
3272:
3261:
3239:
3106:
2973:
2959:
2949:
2916:
2840:
2792:
2735:
2728:
2575:
2475:
2349:
2263:
2245:
2224:
2210:
2110:
2047:
1997:
1919:
1897:
1870:
1833:
1735:
1702:
1550:
1528:
1328:
1311:
1289:
1267:
1189:
1178:
514:
126:
645:
233:. The 2014 discovery was made when a viable specimen was found in a 30,000-year-old
5161:
5156:
5120:
5105:
5084:
4940:
4859:
4804:
4739:
4708:
4675:
4525:
4497:
4315:
4287:
4266:
4252:
4245:
4235:
4071:
4005:
3934:
3782:
3752:
3741:
3688:
3644:
3634:
3615:
3591:
3542:
3454:
3367:
3325:
3164:
3096:
3074:
3054:
3044:
2854:
2770:
2681:
2652:
2640:
2630:
2618:
2608:
2586:
2544:
2534:
2507:
2486:
2464:
2440:
2430:
2408:
2305:
2256:
2217:
2203:
2182:
1951:
1929:
1907:
1774:
1676:
1664:
1621:
1610:
1517:
1506:
1471:
1377:
1229:
1084:
562:
354:
324:
281:
102:
90:
5324:
5333:
5151:
5130:
5048:
5023:
4725:
4504:
4490:
4403:
4259:
4049:
3962:
3955:
3898:
3804:
3725:
3608:
3525:
3491:
3417:
3379:
3360:
3296:
3289:
3120:
2980:
2669:
2497:
2397:
2292:
2142:
2124:
2100:
1963:
1941:
1767:
1745:
1713:
1654:
1443:
1355:
1217:
1167:
1143:
502:
463:
sediment. It was found when riverbank samples collected in 2000 were exposed to
460:
359:
348:
203:
891:
5228:
5203:
4597:
4553:
4447:
4208:
3017:
3007:
2759:
2359:
1686:
1436:
637:
524:
452:
367:
238:
899:
628:
Yong, Ed (3 March 2014). "Giant virus resurrected from 69,000-year-old ice".
5188:
5125:
5115:
5038:
4793:
4664:
4198:
4154:
3228:
2905:
2877:
2316:
1156:
1128:
841:
542:
434:
418:
304:
270:
1043:
860:
722:
17:
5318:
4970:
876:"A taxonomic proposal for cedratviruses, orpheoviruses, and pithoviruses"
704:
567:
437:, rather than the more typical method of taking over the host's nucleus.
290:
234:
1024:
501:, was isolated in 2016. The core features such as the order of ORFs and
332:
are unlike those of other viruses. Despite the physical similarity with
449:
329:
242:
4909:
1008:"Estimating evolutionary rates in giant viruses using ancient genomes"
528:
5223:
4986:
491:
464:
410:
311:
258:
257:, a reference to large storage containers of ancient Greece known as
213:
5295:
4944:
414:
266:
230:
77:
374:, in contrast to greater than 61% for pandoraviruses. The family
426:
319:
5299:
4913:
4777:
4648:
4138:
3212:
2889:
2289:
1140:
1088:
592:
433:
therefore undergoes its entire replication cycle in its host's
294:
223:
794:"Pithovirus: Bigger than Pandoravirus with a smaller genome"
505:(ORFans) are well conserved between the two known species.
279:, the previous largest-known viruses, and is larger than
963:"Frozen Giant Virus Still Infectious After 30,000 Years"
745:"Ancient "Giant Virus" Revived From Siberian Permafrost"
261:, was chosen to describe the new species. A specimen of
940:"Biggest-ever virus revived from Stone Age permafrost"
429:; these proteins are present in the purified virions.
289:
has the largest viral genome, containing 1.9 to 2.5
5308:
5139:
5098:
5062:
5014:
4979:
4857:
4835:
4824:
4813:
4802:
4791:
4706:
4695:
4684:
4673:
4662:
4635:
4585:
4574:
4478:
4428:
4417:
4391:
4380:
4347:
4336:
4233:
4222:
4196:
4185:
4174:
4163:
4152:
4089:
4059:
4037:
4026:
3986:
3915:
3802:
3791:
3780:
3750:
3739:
3713:
3702:
3676:
3665:
3632:
3589:
3578:
3567:
3523:
3512:
3479:
3468:
3442:
3431:
3377:
3313:
3270:
3259:
3248:
3237:
3226:
3152:
3094:
3083:
3072:
3042:
3031:
3005:
2994:
2947:
2936:
2925:
2914:
2903:
2876:
2828:
2790:
2779:
2768:
2757:
2716:
2705:
2679:
2650:
2628:
2606:
2584:
2573:
2562:
2532:
2521:
2495:
2484:
2473:
2462:
2428:
2417:
2406:
2395:
2347:
2336:
2325:
2314:
2303:
2156:
2134:
2098:
2087:
2080:
2028:
2017:
2006:
1995:
1961:
1939:
1917:
1895:
1884:
1858:
1847:
1821:
1810:
1799:
1755:
1733:
1711:
1700:
1674:
1652:
1641:
1630:
1619:
1608:
1570:
1548:
1537:
1526:
1515:
1504:
1375:
1364:
1353:
1309:
1298:
1287:
1276:
1265:
1227:
1198:
1187:
1176:
1165:
1154:
1127:
667:"30,000-year-old giant virus 'comes back to life'"
202:, first described in a 2014 paper, is a genus of
787:
785:
783:
781:
682:
680:
480:Proceedings of the National Academy of Sciences
455:by Chantal Abergel and Jean-Michel Claverie of
809:
807:
768:"World's Biggest Virus May Have Ancient Roots"
597:International Committee on Taxonomy of Viruses
448:was discovered in a 30,000-year-old sample of
4925:
1100:
8:
5296:
4932:
4918:
4910:
4832:
4821:
4810:
4799:
4788:
4774:
4703:
4692:
4681:
4670:
4659:
4645:
4641:
4582:
4425:
4388:
4344:
4230:
4193:
4182:
4171:
4160:
4149:
4135:
4034:
3799:
3788:
3747:
3710:
3673:
3586:
3575:
3520:
3476:
3439:
3267:
3256:
3245:
3234:
3223:
3209:
3091:
3080:
3039:
3002:
2944:
2933:
2922:
2911:
2900:
2886:
2882:
2787:
2776:
2765:
2713:
2581:
2570:
2529:
2492:
2481:
2470:
2425:
2414:
2403:
2344:
2333:
2322:
2311:
2300:
2286:
2095:
2084:
2025:
2014:
2003:
1892:
1855:
1818:
1807:
1708:
1649:
1638:
1627:
1616:
1545:
1534:
1523:
1512:
1372:
1361:
1306:
1295:
1284:
1273:
1195:
1184:
1173:
1162:
1151:
1137:
1133:
1107:
1093:
1085:
657:
655:
42:
31:
1033:
1023:
850:
840:
712:
285:, the smallest eukaryotic cell, although
5174:Laboratory diagnosis of viral infections
738:
736:
734:
732:
623:
621:
619:
617:
615:
613:
961:Pappas, Stephanie (16 September 2015).
918:Swiss Institute of Bioinformatics (SIB)
584:
989:"Out of Siberian Ice, a Virus Revived"
413:is one circular, double-stranded DNA (
7:
5266:
792:Racaniello, Vincent (4 March 2014).
398:has now been ratified by the ICTV.
5377:Nucleocytoplasmic large DNA viruses
5278:
358:. These families all contain large
228:nucleocytoplasmic large DNA viruses
226:based virus and is a member of the
5382:Discovered cryopreserved organisms
950:from the original on 5 March 2014.
25:
421:(bp), encoding approximately 467
5277:
5265:
5254:
5253:
5241:
5005:
766:Brumfiel, Geoff (18 July 2013).
743:Sirucek, Stefan (3 March 2014).
553:List of longest-living organisms
527:
265:measures approximately 1.5
64:
1006:Duchêne, S; Holmes, EC (2018).
497:A modern species in the genus,
477:findings were published in the
938:Coghlan, Andy (3 March 2014).
593:"Virus Taxonomy: 2023 Release"
417:) chromosome of about 610,000
362:viruses with DNA genomes. The
1:
987:Zimmer, Carl (3 March 2014).
821:Proc. Natl. Acad. Sci. U.S.A.
499:Alphapithovirus massiliense
219:Alphapithovirus massiliense
180:Alphapithovirus massiliense
5403:
892:10.1007/s00705-024-06055-x
5237:
5003:
4966:Social history of viruses
4951:
4787:
4773:
4658:
4644:
4148:
4134:
3222:
3208:
2899:
2885:
2299:
2285:
1150:
1136:
638:10.1038/nature.2014.14801
446:Alphapithovirus sibericum
209:Alphapithovirus sibericum
187:Alphapithovirus sibericum
175:
170:
59:
53:Alphapithovirus sibericum
50:
41:
34:
457:Aix-Marseille University
206:known from two species,
842:10.1073/pnas.1320670111
473:two years earlier. The
5167:Helper dependent virus
914:"Pithovirus sibericum"
318:contains 467 distinct
5387:Unaccepted virus taxa
1078:NCBI Taxonomy Browser
772:National Public Radio
5219:Virus quantification
5214:Virus classification
1120:virus classification
1072:Pithovirus sibericum
880:Archives of Virology
573:Virus classification
61:Virus classification
5209:Virus-like particle
4102:Permutotetraviridae
1861:Orthopolintovirales
1336:Malacoherpesviridae
833:2014PNAS..111.4274L
749:National Geographic
423:open reading frames
5340:Pithovirus (genus)
2420:Malgrandaviricetes
2055:Sphaerolipoviridae
2041:Matshushitaviridae
1633:Nucleocytoviricota
993:The New York Times
705:10.1093/gbe/evw153
470:Silene stenophylla
116:Nucleocytoviricota
5364:
5363:
5302:Taxon identifiers
5293:
5292:
5184:Neurotropic virus
5029:Viral replication
4907:
4906:
4903:
4902:
4899:
4898:
4895:
4894:
4891:
4890:
4887:
4886:
4883:
4882:
4879:
4878:
4769:
4768:
4765:
4764:
4761:
4760:
4757:
4756:
4753:
4752:
4749:
4748:
4631:
4630:
4627:
4626:
4623:
4622:
4619:
4618:
4615:
4614:
4611:
4610:
4607:
4606:
4577:Yunchangviricetes
4570:
4569:
4413:
4412:
4376:
4375:
4332:
4331:
4218:
4217:
4130:
4129:
4126:
4125:
4122:
4121:
4118:
4117:
4085:
4084:
4081:
4080:
4022:
4021:
3776:
3775:
3772:
3771:
3735:
3734:
3698:
3697:
3661:
3660:
3563:
3562:
3559:
3558:
3536:Carmotetraviridae
3508:
3507:
3464:
3463:
3427:
3426:
3411:Gammaflexiviridae
3404:Deltaflexiviridae
3390:Alphaflexiviridae
3283:Alphatetraviridae
3204:
3203:
3200:
3199:
3196:
3195:
3192:
3191:
3148:
3147:
3144:
3143:
3068:
3067:
3064:
3063:
3027:
3026:
2990:
2989:
2939:Chrymotiviricetes
2872:
2871:
2868:
2867:
2864:
2863:
2824:
2823:
2820:
2819:
2816:
2815:
2812:
2811:
2753:
2752:
2749:
2748:
2745:
2744:
2701:
2700:
2597:Bacilladnaviridae
2558:
2557:
2554:
2553:
2517:
2516:
2458:
2457:
2454:
2453:
2450:
2449:
2391:
2390:
2387:
2386:
2383:
2382:
2281:
2280:
2277:
2276:
2273:
2272:
2239:Portogloboviridae
2152:
2151:
2076:
2075:
2072:
2071:
2068:
2067:
2064:
2063:
1991:
1990:
1987:
1986:
1983:
1982:
1880:
1879:
1843:
1842:
1802:Preplasmiviricota
1795:
1794:
1791:
1790:
1696:
1695:
1604:
1603:
1600:
1599:
1596:
1595:
1592:
1591:
1500:
1499:
1496:
1495:
1492:
1491:
1488:
1487:
1395:Autographiviridae
1349:
1348:
1345:
1344:
1322:Alloherpesviridae
1261:
1260:
1257:
1256:
1253:
1252:
1249:
1248:
1025:10.1093/ve/vey006
548:Largest organisms
396:Alphacedratvirus,
390:, and the family
370:, similar to the
195:
194:
16:(Redirected from
5394:
5357:
5356:
5344:
5343:
5342:
5329:
5328:
5327:
5297:
5281:
5280:
5269:
5268:
5257:
5256:
5245:
5080:Phenotype mixing
5016:Viral life cycle
5009:
4934:
4927:
4920:
4911:
4833:
4822:
4811:
4800:
4789:
4781:dsDNA-RT viruses
4775:
4704:
4693:
4682:
4671:
4660:
4652:ssRNA-RT viruses
4646:
4642:
4583:
4426:
4394:Serpentovirales
4389:
4367:Orthomyxoviridae
4345:
4295:Peribunyaviridae
4231:
4194:
4188:Chunqiuviricetes
4183:
4172:
4161:
4150:
4142:(–)ssRNA viruses
4136:
4035:
3878:Nanghoshaviridae
3800:
3789:
3763:Botourmiaviridae
3748:
3711:
3705:Howeltoviricetes
3674:
3668:Amabiliviricetes
3587:
3576:
3521:
3477:
3440:
3397:Betaflexiviridae
3268:
3257:
3246:
3235:
3224:
3216:(+)ssRNA viruses
3210:
3128:Picobirnaviridae
3092:
3086:Duplopiviricetes
3081:
3040:
3034:Vidaverviricetes
3003:
2997:Resentoviricetes
2967:Megabirnaviridae
2945:
2934:
2928:Duplornaviricota
2923:
2912:
2901:
2887:
2883:
2788:
2782:Huolimaviricetes
2777:
2766:
2714:
2708:Repensiviricetes
2582:
2571:
2565:Cressdnaviricota
2530:
2493:
2482:
2471:
2426:
2415:
2404:
2345:
2334:
2323:
2312:
2301:
2287:
2096:
2085:
2026:
2015:
2004:
1893:
1887:Tectiliviricetes
1856:
1850:Polintoviricetes
1824:Priklausovirales
1819:
1808:
1782:Marseilleviridae
1709:
1650:
1639:
1628:
1617:
1583:Papillomaviridae
1573:Zurhausenvirales
1546:
1535:
1524:
1513:
1388:Ackermannviridae
1373:
1362:
1307:
1296:
1285:
1274:
1240:Tristromaviridae
1211:Lipothrixviridae
1196:
1185:
1174:
1163:
1152:
1138:
1134:
1109:
1102:
1095:
1086:
1081:
1048:
1047:
1037:
1027:
1003:
997:
996:
984:
978:
977:
975:
973:
958:
952:
951:
935:
929:
928:
926:
924:
910:
904:
903:
871:
865:
864:
854:
844:
811:
802:
801:
789:
776:
775:
763:
757:
756:
755:on 4 March 2014.
751:. Archived from
740:
727:
726:
716:
693:Genome Biol Evol
684:
675:
674:
665:(3 March 2014).
663:Morelle, Rebecca
659:
650:
649:
625:
608:
607:
605:
603:
589:
537:
532:
531:
515:rate of mutation
483:in March 2014.
380:Alphaorpheovirus
343:Marseilleviridae
212:, which infects
69:
68:
46:
32:
27:Genus of viruses
21:
5402:
5401:
5397:
5396:
5395:
5393:
5392:
5391:
5367:
5366:
5365:
5360:
5352:
5347:
5338:
5337:
5332:
5323:
5322:
5317:
5304:
5294:
5289:
5233:
5135:
5094:
5090:Viral evolution
5075:Antigenic shift
5070:Antigenic drift
5058:
5054:Lysogenic cycle
5010:
5001:
4975:
4947:
4938:
4908:
4875:
4853:
4827:Revtraviricetes
4783:
4745:
4698:Revtraviricetes
4654:
4603:
4566:
4533:Paramyxoviridae
4481:Mononegavirales
4474:
4455:Crepuscuviridae
4409:
4372:
4350:Articulavirales
4339:Insthoviricetes
4328:
4214:
4177:Negarnaviricota
4144:
4114:
4077:
4055:
4029:Stelpaviricetes
4018:
3982:
3977:Polycipiviridae
3970:Dicistroviridae
3911:
3794:Pisoniviricetes
3768:
3731:
3694:
3657:
3628:
3555:
3515:Tolucaviricetes
3504:
3460:
3445:Amarillovirales
3423:
3373:
3333:Closteroviridae
3316:Martellivirales
3309:
3251:Kitrinoviricota
3218:
3188:
3172:Polymycoviridae
3140:
3060:
3023:
2986:
2895:
2860:
2848:Finnlakeviridae
2808:
2803:Pleolipoviridae
2741:
2697:
2675:
2646:
2624:
2602:
2550:
2524:Quintoviricetes
2513:
2446:
2379:
2295:
2269:
2148:
2130:
2060:
2031:Halopanivirales
1979:
1957:
1935:
1913:
1876:
1839:
1813:Maveriviricetes
1787:
1751:
1729:
1724:Phycodnaviridae
1692:
1670:
1644:Pokkesviricetes
1588:
1566:
1540:Papovaviricetes
1484:
1451:Rountreeviridae
1341:
1245:
1223:
1146:
1123:
1113:
1068:
1057:
1052:
1051:
1005:
1004:
1000:
986:
985:
981:
971:
969:
960:
959:
955:
937:
936:
932:
922:
920:
912:
911:
907:
873:
872:
868:
813:
812:
805:
791:
790:
779:
765:
764:
760:
742:
741:
730:
686:
685:
678:
661:
660:
653:
627:
626:
611:
601:
599:
591:
590:
586:
581:
558:List of viruses
533:
526:
523:
511:
475:Alphapithovirus
443:
431:Alphapithovirus
408:Alphapithovirus
404:
388:Alphapithovirus
366:genome has 36%
364:Alphapithovirus
338:Alphapithovirus
316:Alphapithovirus
301:Alphapithovirus
263:Alphapithovirus
255:Alphapithovirus
253:The genus name
251:
237:harvested from
199:Alphapithovirus
164:Alphapithovirus
63:
36:Alphapithovirus
28:
23:
22:
15:
12:
11:
5:
5400:
5398:
5390:
5389:
5384:
5379:
5369:
5368:
5362:
5361:
5359:
5358:
5345:
5330:
5314:
5312:
5306:
5305:
5300:
5291:
5290:
5288:
5287:
5275:
5263:
5251:
5238:
5235:
5234:
5232:
5231:
5226:
5221:
5216:
5211:
5206:
5201:
5196:
5191:
5186:
5181:
5179:Marine viruses
5176:
5171:
5170:
5169:
5159:
5154:
5149:
5147:Antiviral drug
5143:
5141:
5137:
5136:
5134:
5133:
5128:
5123:
5118:
5113:
5108:
5102:
5100:
5096:
5095:
5093:
5092:
5087:
5082:
5077:
5072:
5066:
5064:
5060:
5059:
5057:
5056:
5051:
5046:
5041:
5036:
5034:Viral shedding
5031:
5026:
5020:
5018:
5012:
5011:
5004:
5002:
5000:
4999:
4994:
4992:Viral envelope
4989:
4983:
4981:
4977:
4976:
4974:
4973:
4968:
4963:
4958:
4952:
4949:
4948:
4939:
4937:
4936:
4929:
4922:
4914:
4905:
4904:
4901:
4900:
4897:
4896:
4893:
4892:
4889:
4888:
4885:
4884:
4881:
4880:
4877:
4876:
4874:
4873:
4870:Caulimoviridae
4865:
4863:
4855:
4854:
4852:
4851:
4848:Hepadnaviridae
4843:
4841:
4830:
4819:
4816:Artverviricota
4808:
4797:
4785:
4784:
4778:
4771:
4770:
4767:
4766:
4763:
4762:
4759:
4758:
4755:
4754:
4751:
4750:
4747:
4746:
4744:
4743:
4736:
4729:
4722:
4714:
4712:
4701:
4690:
4687:Artverviricota
4679:
4668:
4656:
4655:
4649:
4639:
4633:
4632:
4629:
4628:
4625:
4624:
4621:
4620:
4617:
4616:
4613:
4612:
4609:
4608:
4605:
4604:
4602:
4601:
4593:
4591:
4588:Goujianvirales
4580:
4572:
4571:
4568:
4567:
4565:
4564:
4557:
4550:
4543:
4536:
4529:
4522:
4515:
4508:
4501:
4494:
4486:
4484:
4476:
4475:
4473:
4472:
4465:
4458:
4451:
4444:
4436:
4434:
4431:Jingchuvirales
4423:
4420:Monjiviricetes
4415:
4414:
4411:
4410:
4408:
4407:
4399:
4397:
4386:
4383:Milneviricetes
4378:
4377:
4374:
4373:
4371:
4370:
4363:
4355:
4353:
4342:
4334:
4333:
4330:
4329:
4327:
4326:
4319:
4312:
4305:
4298:
4291:
4284:
4277:
4274:Leishbuviridae
4270:
4263:
4256:
4249:
4241:
4239:
4228:
4225:Ellioviricetes
4220:
4219:
4216:
4215:
4213:
4212:
4204:
4202:
4191:
4180:
4169:
4158:
4146:
4145:
4139:
4132:
4131:
4128:
4127:
4124:
4123:
4120:
4119:
4116:
4115:
4113:
4112:
4109:Sarthroviridae
4105:
4093:
4091:
4087:
4086:
4083:
4082:
4079:
4078:
4076:
4075:
4067:
4065:
4057:
4056:
4054:
4053:
4045:
4043:
4032:
4024:
4023:
4020:
4019:
4017:
4016:
4009:
4002:
3999:Alvernaviridae
3994:
3992:
3984:
3983:
3981:
3980:
3973:
3966:
3959:
3952:
3945:
3942:Solinviviridae
3938:
3931:
3928:Picornaviridae
3923:
3921:
3918:Picornavirales
3913:
3912:
3910:
3909:
3902:
3895:
3888:
3885:Nanhypoviridae
3881:
3874:
3867:
3860:
3857:Medioniviridae
3853:
3846:
3839:
3832:
3829:Cremegaviridae
3825:
3818:
3810:
3808:
3797:
3786:
3778:
3777:
3774:
3773:
3770:
3769:
3767:
3766:
3758:
3756:
3745:
3737:
3736:
3733:
3732:
3730:
3729:
3721:
3719:
3708:
3700:
3699:
3696:
3695:
3693:
3692:
3684:
3682:
3679:Wolframvirales
3671:
3663:
3662:
3659:
3658:
3656:
3655:
3648:
3640:
3638:
3630:
3629:
3627:
3626:
3619:
3612:
3605:
3597:
3595:
3584:
3573:
3565:
3564:
3561:
3560:
3557:
3556:
3554:
3553:
3546:
3539:
3531:
3529:
3518:
3510:
3509:
3506:
3505:
3503:
3502:
3499:Sinhaliviridae
3495:
3487:
3485:
3474:
3471:Magsaviricetes
3466:
3465:
3462:
3461:
3459:
3458:
3450:
3448:
3437:
3434:Flasuviricetes
3429:
3428:
3425:
3424:
3422:
3421:
3414:
3407:
3400:
3393:
3385:
3383:
3375:
3374:
3372:
3371:
3364:
3357:
3350:
3343:
3340:Endornaviridae
3336:
3329:
3321:
3319:
3311:
3310:
3308:
3307:
3300:
3293:
3286:
3278:
3276:
3265:
3254:
3243:
3232:
3220:
3219:
3213:
3206:
3205:
3202:
3201:
3198:
3197:
3194:
3193:
3190:
3189:
3187:
3186:
3183:Botybirnavirus
3175:
3168:
3156:
3154:
3150:
3149:
3146:
3145:
3142:
3141:
3139:
3138:
3135:Partitiviridae
3131:
3124:
3117:
3114:Curvulaviridae
3110:
3102:
3100:
3089:
3078:
3070:
3069:
3066:
3065:
3062:
3061:
3059:
3058:
3050:
3048:
3037:
3029:
3028:
3025:
3024:
3022:
3021:
3013:
3011:
3000:
2992:
2991:
2988:
2987:
2985:
2984:
2977:
2970:
2963:
2955:
2953:
2942:
2931:
2920:
2909:
2897:
2896:
2890:
2880:
2874:
2873:
2870:
2869:
2866:
2865:
2862:
2861:
2859:
2858:
2851:
2844:
2832:
2830:
2826:
2825:
2822:
2821:
2818:
2817:
2814:
2813:
2810:
2809:
2807:
2806:
2798:
2796:
2785:
2774:
2763:
2755:
2754:
2751:
2750:
2747:
2746:
2743:
2742:
2740:
2739:
2732:
2724:
2722:
2719:Geplafuvirales
2711:
2703:
2702:
2699:
2698:
2696:
2695:
2692:Redondoviridae
2687:
2685:
2677:
2676:
2674:
2673:
2666:
2658:
2656:
2648:
2647:
2645:
2644:
2636:
2634:
2626:
2625:
2623:
2622:
2614:
2612:
2604:
2603:
2601:
2600:
2592:
2590:
2579:
2568:
2560:
2559:
2556:
2555:
2552:
2551:
2549:
2548:
2540:
2538:
2527:
2519:
2518:
2515:
2514:
2512:
2511:
2503:
2501:
2490:
2479:
2468:
2460:
2459:
2456:
2455:
2452:
2451:
2448:
2447:
2445:
2444:
2436:
2434:
2423:
2412:
2401:
2393:
2392:
2389:
2388:
2385:
2384:
2381:
2380:
2378:
2377:
2374:Plectroviridae
2370:
2367:Paulinoviridae
2363:
2355:
2353:
2342:
2339:Faserviricetes
2331:
2328:Hofneiviricota
2320:
2309:
2297:
2296:
2290:
2283:
2282:
2279:
2278:
2275:
2274:
2271:
2270:
2268:
2267:
2260:
2249:
2242:
2235:
2232:Polydnaviridae
2228:
2221:
2214:
2207:
2200:
2197:Globuloviridae
2193:
2190:Fuselloviridae
2186:
2179:
2176:Bicaudaviridae
2172:
2169:Ampullaviridae
2160:
2158:
2154:
2153:
2150:
2149:
2147:
2146:
2138:
2136:
2132:
2131:
2129:
2128:
2121:
2118:Hytrosaviridae
2114:
2106:
2104:
2093:
2090:Naldaviricetes
2082:
2078:
2077:
2074:
2073:
2070:
2069:
2066:
2065:
2062:
2061:
2059:
2058:
2051:
2044:
2036:
2034:
2023:
2020:Laserviricetes
2012:
2009:Dividoviricota
2001:
1993:
1992:
1989:
1988:
1985:
1984:
1981:
1980:
1978:
1977:
1974:Corticoviridae
1969:
1967:
1959:
1958:
1956:
1955:
1947:
1945:
1937:
1936:
1934:
1933:
1925:
1923:
1915:
1914:
1912:
1911:
1903:
1901:
1890:
1882:
1881:
1878:
1877:
1875:
1874:
1866:
1864:
1853:
1845:
1844:
1841:
1840:
1838:
1837:
1829:
1827:
1816:
1805:
1797:
1796:
1793:
1792:
1789:
1788:
1786:
1785:
1778:
1771:
1763:
1761:
1758:Pimascovirales
1753:
1752:
1750:
1749:
1741:
1739:
1731:
1730:
1728:
1727:
1719:
1717:
1706:
1698:
1697:
1694:
1693:
1691:
1690:
1682:
1680:
1672:
1671:
1669:
1668:
1660:
1658:
1647:
1636:
1625:
1614:
1606:
1605:
1602:
1601:
1598:
1597:
1594:
1593:
1590:
1589:
1587:
1586:
1578:
1576:
1568:
1567:
1565:
1564:
1561:Polyomaviridae
1556:
1554:
1543:
1532:
1521:
1510:
1502:
1501:
1498:
1497:
1494:
1493:
1490:
1489:
1486:
1485:
1483:
1482:
1475:
1468:
1461:
1458:Salasmaviridae
1454:
1447:
1440:
1433:
1430:Herelleviridae
1426:
1419:
1416:Drexlerviridae
1412:
1409:Demerecviridae
1405:
1398:
1391:
1383:
1381:
1370:
1367:Caudoviricetes
1359:
1351:
1350:
1347:
1346:
1343:
1342:
1340:
1339:
1332:
1325:
1317:
1315:
1304:
1301:Herviviricetes
1293:
1282:
1279:Heunggongvirae
1271:
1263:
1262:
1259:
1258:
1255:
1254:
1251:
1250:
1247:
1246:
1244:
1243:
1235:
1233:
1225:
1224:
1222:
1221:
1214:
1206:
1204:
1201:Ligamenvirales
1193:
1182:
1171:
1160:
1148:
1147:
1141:
1131:
1125:
1124:
1114:
1112:
1111:
1104:
1097:
1089:
1083:
1082:
1066:
1056:
1055:External links
1053:
1050:
1049:
998:
979:
953:
930:
905:
866:
827:(11): 4274–9.
803:
777:
758:
728:
676:
651:
609:
583:
582:
580:
577:
576:
575:
570:
565:
560:
555:
550:
545:
539:
538:
535:Viruses portal
522:
519:
510:
507:
488:climate change
442:
439:
403:
400:
276:Pandoraviridae
250:
247:
193:
192:
191:
190:
183:
173:
172:
168:
167:
160:
156:
155:
148:
144:
143:
140:Pimascovirales
136:
132:
131:
124:
120:
119:
112:
108:
107:
100:
96:
95:
88:
81:
80:
75:
71:
70:
57:
56:
48:
47:
39:
38:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
5399:
5388:
5385:
5383:
5380:
5378:
5375:
5374:
5372:
5355:
5350:
5346:
5341:
5335:
5331:
5326:
5320:
5316:
5315:
5313:
5311:
5307:
5303:
5298:
5286:
5285:
5276:
5274:
5273:
5264:
5262:
5261:
5252:
5250:
5249:
5244:
5240:
5239:
5236:
5230:
5227:
5225:
5222:
5220:
5217:
5215:
5212:
5210:
5207:
5205:
5202:
5200:
5199:Viral disease
5197:
5195:
5192:
5190:
5187:
5185:
5182:
5180:
5177:
5175:
5172:
5168:
5165:
5164:
5163:
5160:
5158:
5155:
5153:
5150:
5148:
5145:
5144:
5142:
5138:
5132:
5129:
5127:
5124:
5122:
5119:
5117:
5114:
5112:
5111:Bacteriophage
5109:
5107:
5104:
5103:
5101:
5097:
5091:
5088:
5086:
5083:
5081:
5078:
5076:
5073:
5071:
5068:
5067:
5065:
5061:
5055:
5052:
5050:
5047:
5045:
5044:Virus latency
5042:
5040:
5037:
5035:
5032:
5030:
5027:
5025:
5022:
5021:
5019:
5017:
5013:
5008:
4998:
4997:Viral protein
4995:
4993:
4990:
4988:
4985:
4984:
4982:
4978:
4972:
4969:
4967:
4964:
4962:
4959:
4957:
4954:
4953:
4950:
4946:
4942:
4935:
4930:
4928:
4923:
4921:
4916:
4915:
4912:
4872:
4871:
4867:
4866:
4864:
4862:
4861:
4856:
4850:
4849:
4845:
4844:
4842:
4840:
4839:
4838:Blubervirales
4834:
4831:
4829:
4828:
4823:
4820:
4818:
4817:
4812:
4809:
4807:
4806:
4801:
4798:
4796:
4795:
4790:
4786:
4782:
4776:
4772:
4742:
4741:
4737:
4735:
4734:
4733:Pseudoviridae
4730:
4728:
4727:
4723:
4721:
4720:
4719:Belpaoviridae
4716:
4715:
4713:
4711:
4710:
4705:
4702:
4700:
4699:
4694:
4691:
4689:
4688:
4683:
4680:
4678:
4677:
4672:
4669:
4667:
4666:
4661:
4657:
4653:
4647:
4643:
4640:
4638:
4634:
4600:
4599:
4595:
4594:
4592:
4590:
4589:
4584:
4581:
4579:
4578:
4573:
4563:
4562:
4558:
4556:
4555:
4551:
4549:
4548:
4547:Rhabdoviridae
4544:
4542:
4541:
4540:Pneumoviridae
4537:
4535:
4534:
4530:
4528:
4527:
4523:
4521:
4520:
4519:Mymonaviridae
4516:
4514:
4513:
4509:
4507:
4506:
4502:
4500:
4499:
4495:
4493:
4492:
4488:
4487:
4485:
4483:
4482:
4477:
4471:
4470:
4469:Natareviridae
4466:
4464:
4463:
4459:
4457:
4456:
4452:
4450:
4449:
4445:
4443:
4442:
4438:
4437:
4435:
4433:
4432:
4427:
4424:
4422:
4421:
4416:
4406:
4405:
4401:
4400:
4398:
4396:
4395:
4390:
4387:
4385:
4384:
4379:
4369:
4368:
4364:
4362:
4361:
4360:Amnoonviridae
4357:
4356:
4354:
4352:
4351:
4346:
4343:
4341:
4340:
4335:
4325:
4324:
4323:Wupedeviridae
4320:
4318:
4317:
4313:
4311:
4310:
4309:Phenuiviridae
4306:
4304:
4303:
4302:Phasmaviridae
4299:
4297:
4296:
4292:
4290:
4289:
4285:
4283:
4282:
4278:
4276:
4275:
4271:
4269:
4268:
4264:
4262:
4261:
4257:
4255:
4254:
4250:
4248:
4247:
4243:
4242:
4240:
4238:
4237:
4232:
4229:
4227:
4226:
4221:
4211:
4210:
4206:
4205:
4203:
4201:
4200:
4195:
4192:
4190:
4189:
4184:
4181:
4179:
4178:
4173:
4170:
4168:
4167:
4166:Orthornavirae
4162:
4159:
4157:
4156:
4151:
4147:
4143:
4137:
4133:
4111:
4110:
4106:
4104:
4103:
4098:
4095:
4094:
4092:
4088:
4074:
4073:
4069:
4068:
4066:
4064:
4063:
4062:Stellavirales
4058:
4052:
4051:
4047:
4046:
4044:
4042:
4041:
4040:Patatavirales
4036:
4033:
4031:
4030:
4025:
4015:
4014:
4013:Solemoviridae
4010:
4008:
4007:
4003:
4001:
4000:
3996:
3995:
3993:
3991:
3990:
3989:Sobelivirales
3985:
3979:
3978:
3974:
3972:
3971:
3967:
3965:
3964:
3960:
3958:
3957:
3953:
3951:
3950:
3949:Caliciviridae
3946:
3944:
3943:
3939:
3937:
3936:
3932:
3930:
3929:
3925:
3924:
3922:
3920:
3919:
3914:
3908:
3907:
3906:Tobaniviridae
3903:
3901:
3900:
3896:
3894:
3893:
3889:
3887:
3886:
3882:
3880:
3879:
3875:
3873:
3872:
3871:Mononiviridae
3868:
3866:
3865:
3864:Mesoniviridae
3861:
3859:
3858:
3854:
3852:
3851:
3850:Gresnaviridae
3847:
3845:
3844:
3843:Euroniviridae
3840:
3838:
3837:
3836:Coronaviridae
3833:
3831:
3830:
3826:
3824:
3823:
3822:Arteriviridae
3819:
3817:
3816:
3815:Abyssoviridae
3812:
3811:
3809:
3807:
3806:
3801:
3798:
3796:
3795:
3790:
3787:
3785:
3784:
3779:
3765:
3764:
3760:
3759:
3757:
3755:
3754:
3749:
3746:
3744:
3743:
3738:
3728:
3727:
3723:
3722:
3720:
3718:
3717:
3716:Cryppavirales
3712:
3709:
3707:
3706:
3701:
3691:
3690:
3686:
3685:
3683:
3681:
3680:
3675:
3672:
3670:
3669:
3664:
3654:
3653:
3652:Steitzviridae
3649:
3647:
3646:
3642:
3641:
3639:
3637:
3636:
3631:
3625:
3624:
3623:Solspiviridae
3620:
3618:
3617:
3613:
3611:
3610:
3606:
3604:
3603:
3602:Atkinsviridae
3599:
3598:
3596:
3594:
3593:
3588:
3585:
3583:
3582:
3581:Leviviricetes
3577:
3574:
3572:
3571:
3570:Lenarviricota
3566:
3552:
3551:
3550:Tombusviridae
3547:
3545:
3544:
3540:
3538:
3537:
3533:
3532:
3530:
3528:
3527:
3522:
3519:
3517:
3516:
3511:
3501:
3500:
3496:
3494:
3493:
3489:
3488:
3486:
3484:
3483:
3482:Nodamuvirales
3478:
3475:
3473:
3472:
3467:
3457:
3456:
3452:
3451:
3449:
3447:
3446:
3441:
3438:
3436:
3435:
3430:
3420:
3419:
3415:
3413:
3412:
3408:
3406:
3405:
3401:
3399:
3398:
3394:
3392:
3391:
3387:
3386:
3384:
3382:
3381:
3376:
3370:
3369:
3365:
3363:
3362:
3358:
3356:
3355:
3351:
3349:
3348:
3344:
3342:
3341:
3337:
3335:
3334:
3330:
3328:
3327:
3323:
3322:
3320:
3318:
3317:
3312:
3306:
3305:
3304:Matonaviridae
3301:
3299:
3298:
3294:
3292:
3291:
3287:
3285:
3284:
3280:
3279:
3277:
3275:
3274:
3273:Hepelivirales
3269:
3266:
3264:
3263:
3262:Alsuviricetes
3258:
3255:
3253:
3252:
3247:
3244:
3242:
3241:
3240:Orthornavirae
3236:
3233:
3231:
3230:
3225:
3221:
3217:
3211:
3207:
3185:
3184:
3179:
3176:
3174:
3173:
3169:
3167:
3166:
3161:
3158:
3157:
3155:
3151:
3137:
3136:
3132:
3130:
3129:
3125:
3123:
3122:
3118:
3116:
3115:
3111:
3109:
3108:
3107:Amalgaviridae
3104:
3103:
3101:
3099:
3098:
3093:
3090:
3088:
3087:
3082:
3079:
3077:
3076:
3071:
3057:
3056:
3052:
3051:
3049:
3047:
3046:
3041:
3038:
3036:
3035:
3030:
3020:
3019:
3015:
3014:
3012:
3010:
3009:
3004:
3001:
2999:
2998:
2993:
2983:
2982:
2978:
2976:
2975:
2974:Quadriviridae
2971:
2969:
2968:
2964:
2962:
2961:
2960:Chrysoviridae
2957:
2956:
2954:
2952:
2951:
2950:Ghabrivirales
2946:
2943:
2941:
2940:
2935:
2932:
2930:
2929:
2924:
2921:
2919:
2918:
2917:Orthornavirae
2913:
2910:
2908:
2907:
2902:
2898:
2894:
2893:dsRNA viruses
2888:
2884:
2881:
2879:
2875:
2857:
2856:
2852:
2850:
2849:
2845:
2843:
2842:
2841:Anelloviridae
2837:
2834:
2833:
2831:
2827:
2805:
2804:
2800:
2799:
2797:
2795:
2794:
2793:Haloruvirales
2789:
2786:
2784:
2783:
2778:
2775:
2773:
2772:
2767:
2764:
2762:
2761:
2756:
2738:
2737:
2736:Genomoviridae
2733:
2731:
2730:
2729:Geminiviridae
2726:
2725:
2723:
2721:
2720:
2715:
2712:
2710:
2709:
2704:
2694:
2693:
2689:
2688:
2686:
2684:
2683:
2678:
2672:
2671:
2667:
2665:
2664:
2663:Metaxyviridae
2660:
2659:
2657:
2655:
2654:
2649:
2643:
2642:
2638:
2637:
2635:
2633:
2632:
2627:
2621:
2620:
2616:
2615:
2613:
2611:
2610:
2605:
2599:
2598:
2594:
2593:
2591:
2589:
2588:
2583:
2580:
2578:
2577:
2576:Arfiviricetes
2572:
2569:
2567:
2566:
2561:
2547:
2546:
2542:
2541:
2539:
2537:
2536:
2531:
2528:
2526:
2525:
2520:
2510:
2509:
2505:
2504:
2502:
2500:
2499:
2494:
2491:
2489:
2488:
2483:
2480:
2478:
2477:
2476:Cossaviricota
2472:
2469:
2467:
2466:
2461:
2443:
2442:
2438:
2437:
2435:
2433:
2432:
2427:
2424:
2422:
2421:
2416:
2413:
2411:
2410:
2405:
2402:
2400:
2399:
2394:
2376:
2375:
2371:
2369:
2368:
2364:
2362:
2361:
2357:
2356:
2354:
2352:
2351:
2350:Tubulavirales
2346:
2343:
2341:
2340:
2335:
2332:
2330:
2329:
2324:
2321:
2319:
2318:
2313:
2310:
2308:
2307:
2302:
2298:
2294:
2293:ssDNA viruses
2288:
2284:
2266:
2265:
2264:Rhizidiovirus
2261:
2259:
2258:
2253:
2250:
2248:
2247:
2246:Thaspiviridae
2243:
2241:
2240:
2236:
2234:
2233:
2229:
2227:
2226:
2225:Plasmaviridae
2222:
2220:
2219:
2215:
2213:
2212:
2211:Halspiviridae
2208:
2206:
2205:
2201:
2199:
2198:
2194:
2192:
2191:
2187:
2185:
2184:
2180:
2178:
2177:
2173:
2171:
2170:
2165:
2162:
2161:
2159:
2155:
2145:
2144:
2140:
2139:
2137:
2133:
2127:
2126:
2122:
2120:
2119:
2115:
2113:
2112:
2111:Baculoviridae
2108:
2107:
2105:
2103:
2102:
2097:
2094:
2092:
2091:
2086:
2083:
2079:
2057:
2056:
2052:
2050:
2049:
2048:Simuloviridae
2045:
2043:
2042:
2038:
2037:
2035:
2033:
2032:
2027:
2024:
2022:
2021:
2016:
2013:
2011:
2010:
2005:
2002:
2000:
1999:
1998:Helvetiavirae
1994:
1976:
1975:
1971:
1970:
1968:
1966:
1965:
1960:
1954:
1953:
1949:
1948:
1946:
1944:
1943:
1938:
1932:
1931:
1927:
1926:
1924:
1922:
1921:
1920:Kalamavirales
1916:
1910:
1909:
1905:
1904:
1902:
1900:
1899:
1898:Belfryvirales
1894:
1891:
1889:
1888:
1883:
1873:
1872:
1871:Adintoviridae
1868:
1867:
1865:
1863:
1862:
1857:
1854:
1852:
1851:
1846:
1836:
1835:
1834:Lavidaviridae
1831:
1830:
1828:
1826:
1825:
1820:
1817:
1815:
1814:
1809:
1806:
1804:
1803:
1798:
1784:
1783:
1779:
1777:
1776:
1772:
1770:
1769:
1765:
1764:
1762:
1760:
1759:
1754:
1748:
1747:
1743:
1742:
1740:
1738:
1737:
1736:Imitervirales
1732:
1726:
1725:
1721:
1720:
1718:
1716:
1715:
1710:
1707:
1705:
1704:
1703:Megaviricetes
1699:
1689:
1688:
1684:
1683:
1681:
1679:
1678:
1673:
1667:
1666:
1662:
1661:
1659:
1657:
1656:
1651:
1648:
1646:
1645:
1640:
1637:
1635:
1634:
1629:
1626:
1624:
1623:
1618:
1615:
1613:
1612:
1607:
1585:
1584:
1580:
1579:
1577:
1575:
1574:
1569:
1563:
1562:
1558:
1557:
1555:
1553:
1552:
1551:Sepolyvirales
1547:
1544:
1542:
1541:
1536:
1533:
1531:
1530:
1529:Cossaviricota
1525:
1522:
1520:
1519:
1514:
1511:
1509:
1508:
1503:
1481:
1480:
1479:Zobellviridae
1476:
1474:
1473:
1469:
1467:
1466:
1465:Schitoviridae
1462:
1460:
1459:
1455:
1453:
1452:
1448:
1446:
1445:
1441:
1439:
1438:
1434:
1432:
1431:
1427:
1425:
1424:
1423:Guelinviridae
1420:
1418:
1417:
1413:
1411:
1410:
1406:
1404:
1403:
1399:
1397:
1396:
1392:
1390:
1389:
1385:
1384:
1382:
1380:
1379:
1374:
1371:
1369:
1368:
1363:
1360:
1358:
1357:
1352:
1338:
1337:
1333:
1331:
1330:
1329:Herpesviridae
1326:
1324:
1323:
1319:
1318:
1316:
1314:
1313:
1312:Herpesvirales
1308:
1305:
1303:
1302:
1297:
1294:
1292:
1291:
1290:Peploviricota
1286:
1283:
1281:
1280:
1275:
1272:
1270:
1269:
1268:Duplodnaviria
1264:
1242:
1241:
1237:
1236:
1234:
1232:
1231:
1226:
1220:
1219:
1215:
1213:
1212:
1208:
1207:
1205:
1203:
1202:
1197:
1194:
1192:
1191:
1190:Tokiviricetes
1186:
1183:
1181:
1180:
1179:Taleaviricota
1175:
1172:
1170:
1169:
1164:
1161:
1159:
1158:
1153:
1149:
1145:
1144:dsDNA viruses
1139:
1135:
1132:
1130:
1126:
1121:
1117:
1110:
1105:
1103:
1098:
1096:
1091:
1090:
1087:
1079:
1075:
1073:
1067:
1065:
1063:
1059:
1058:
1054:
1045:
1041:
1036:
1031:
1026:
1021:
1018:(1): vey006.
1017:
1013:
1009:
1002:
999:
994:
990:
983:
980:
968:
964:
957:
954:
949:
945:
941:
934:
931:
919:
915:
909:
906:
901:
897:
893:
889:
885:
881:
877:
870:
867:
862:
858:
853:
848:
843:
838:
834:
830:
826:
823:
822:
817:
810:
808:
804:
799:
798:Virology Blog
795:
788:
786:
784:
782:
778:
773:
769:
762:
759:
754:
750:
746:
739:
737:
735:
733:
729:
724:
720:
715:
710:
706:
702:
699:(8): 2333–9.
698:
694:
690:
683:
681:
677:
672:
668:
664:
658:
656:
652:
647:
643:
639:
635:
631:
624:
622:
620:
618:
616:
614:
610:
598:
594:
588:
585:
578:
574:
571:
569:
566:
564:
561:
559:
556:
554:
551:
549:
546:
544:
541:
540:
536:
530:
525:
520:
518:
516:
508:
506:
504:
500:
495:
493:
489:
484:
482:
481:
476:
472:
471:
466:
462:
458:
454:
451:
447:
440:
438:
436:
432:
428:
424:
420:
416:
412:
409:
401:
399:
397:
393:
392:Cedratviridae
389:
385:
382:, the family
381:
377:
376:Orpheoviridae
373:
369:
365:
361:
357:
356:
351:
350:
345:
344:
339:
335:
331:
327:
326:
321:
317:
313:
308:
306:
302:
298:
296:
292:
288:
284:
283:
278:
277:
272:
268:
264:
260:
256:
248:
246:
244:
240:
236:
232:
229:
225:
221:
220:
215:
211:
210:
205:
201:
200:
189:
188:
184:
182:
181:
177:
176:
174:
169:
166:
165:
161:
158:
157:
154:
153:
149:
146:
145:
142:
141:
137:
134:
133:
130:
129:
128:Megaviricetes
125:
122:
121:
118:
117:
113:
110:
109:
106:
105:
101:
98:
97:
94:
93:
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72:
67:
62:
58:
55:
54:
49:
45:
40:
37:
33:
30:
19:
5309:
5282:
5270:
5258:
5246:
5162:Viral vector
5157:Helper virus
5121:Human virome
5106:Animal virus
5085:Reassortment
4961:Introduction
4941:Microbiology
4868:
4860:Ortervirales
4858:
4846:
4836:
4825:
4814:
4805:Pararnavirae
4803:
4792:
4740:Retroviridae
4738:
4731:
4724:
4717:
4709:Ortervirales
4707:
4696:
4685:
4676:Pararnavirae
4674:
4663:
4596:
4586:
4575:
4561:Xinmoviridae
4559:
4552:
4545:
4538:
4531:
4526:Nyamiviridae
4524:
4517:
4512:Lispiviridae
4510:
4503:
4498:Bornaviridae
4496:
4489:
4479:
4467:
4462:Myriaviridae
4460:
4453:
4446:
4441:Aliusviridae
4439:
4429:
4418:
4402:
4392:
4381:
4365:
4358:
4348:
4337:
4321:
4316:Tospoviridae
4314:
4307:
4300:
4293:
4288:Nairoviridae
4286:
4279:
4272:
4267:Hantaviridae
4265:
4258:
4253:Arenaviridae
4251:
4246:Cruliviridae
4244:
4236:Bunyavirales
4234:
4223:
4207:
4197:
4186:
4175:
4164:
4153:
4107:
4100:
4096:
4072:Astroviridae
4070:
4060:
4048:
4038:
4027:
4011:
4006:Barnaviridae
4004:
3997:
3987:
3975:
3968:
3961:
3954:
3947:
3940:
3935:Marnaviridae
3933:
3926:
3916:
3904:
3897:
3892:Olifoviridae
3890:
3883:
3876:
3869:
3862:
3855:
3848:
3841:
3834:
3827:
3820:
3813:
3803:
3792:
3783:Pisuviricota
3781:
3761:
3753:Ourlivirales
3751:
3742:Miaviricetes
3740:
3724:
3714:
3703:
3689:Narnaviridae
3687:
3677:
3666:
3650:
3645:Blumeviridae
3643:
3635:Timlovirales
3633:
3621:
3616:Fiersviridae
3614:
3607:
3600:
3592:Norzivirales
3590:
3579:
3568:
3548:
3543:Luteoviridae
3541:
3534:
3524:
3513:
3497:
3490:
3480:
3469:
3455:Flaviviridae
3453:
3443:
3432:
3416:
3409:
3402:
3395:
3388:
3378:
3368:Virgaviridae
3366:
3359:
3352:
3345:
3338:
3331:
3326:Bromoviridae
3324:
3314:
3302:
3295:
3288:
3281:
3271:
3260:
3249:
3238:
3227:
3181:
3177:
3170:
3165:Birnaviridae
3163:
3159:
3133:
3126:
3119:
3112:
3105:
3097:Durnavirales
3095:
3084:
3075:Pisuviricota
3073:
3055:Cystoviridae
3053:
3045:Mindivirales
3043:
3032:
3016:
3006:
2995:
2979:
2972:
2965:
2958:
2948:
2937:
2926:
2915:
2904:
2855:Spiraviridae
2853:
2846:
2839:
2835:
2801:
2791:
2780:
2771:Saleviricota
2769:
2758:
2734:
2727:
2717:
2706:
2690:
2682:Recrevirales
2680:
2668:
2661:
2653:Mulpavirales
2651:
2641:Smacoviridae
2639:
2631:Cremevirales
2629:
2619:Circoviridae
2617:
2609:Cirlivirales
2607:
2595:
2587:Baphyvirales
2585:
2574:
2563:
2545:Parvoviridae
2543:
2535:Piccovirales
2533:
2522:
2508:Bidnaviridae
2506:
2496:
2487:Mouviricetes
2485:
2474:
2465:Shotokuvirae
2463:
2441:Microviridae
2439:
2431:Petitvirales
2429:
2418:
2409:Phixviricota
2407:
2396:
2372:
2365:
2358:
2348:
2337:
2326:
2315:
2306:Monodnaviria
2304:
2262:
2257:Dinodnavirus
2255:
2251:
2244:
2237:
2230:
2223:
2218:Ovaliviridae
2216:
2209:
2204:Guttaviridae
2202:
2195:
2188:
2183:Clavaviridae
2181:
2174:
2167:
2163:
2141:
2123:
2116:
2109:
2099:
2088:
2053:
2046:
2039:
2029:
2018:
2007:
1996:
1972:
1962:
1952:Adenoviridae
1950:
1940:
1930:Tectiviridae
1928:
1918:
1908:Turriviridae
1906:
1896:
1885:
1869:
1859:
1848:
1832:
1822:
1811:
1800:
1780:
1775:Iridoviridae
1773:
1766:
1756:
1744:
1734:
1722:
1712:
1701:
1685:
1677:Chitovirales
1675:
1665:Asfarviridae
1663:
1653:
1642:
1631:
1622:Bamfordvirae
1620:
1611:Varidnaviria
1609:
1581:
1571:
1559:
1549:
1538:
1527:
1518:Shotokuvirae
1516:
1507:Monodnaviria
1505:
1477:
1472:Siphoviridae
1470:
1463:
1456:
1449:
1442:
1435:
1428:
1421:
1414:
1407:
1402:Chaseviridae
1400:
1393:
1386:
1378:Caudovirales
1376:
1365:
1354:
1334:
1327:
1320:
1310:
1299:
1288:
1277:
1266:
1238:
1230:Primavirales
1228:
1216:
1209:
1199:
1188:
1177:
1166:
1155:
1077:
1071:
1064:: Pithovirus
1061:
1015:
1011:
1001:
992:
982:
972:20 September
970:. Retrieved
966:
956:
944:NewScientist
943:
933:
921:. Retrieved
917:
908:
883:
879:
869:
824:
819:
797:
771:
761:
753:the original
748:
696:
692:
670:
629:
600:. Retrieved
596:
587:
563:Microbiology
512:
503:orphan genes
498:
496:
485:
478:
474:
468:
445:
444:
430:
407:
405:
395:
391:
387:
384:Pithoviridae
383:
379:
375:
371:
363:
355:Iridoviridae
353:
347:
341:
337:
334:Pandoravirus
333:
325:Pandoravirus
323:
315:
309:
300:
299:
287:Pandoravirus
286:
282:Ostreococcus
280:
274:
262:
254:
252:
217:
207:
198:
197:
196:
185:
178:
163:
162:
152:Pithoviridae
151:
150:
139:
127:
115:
104:Bamfordvirae
103:
92:Varidnaviria
91:
84:
74:(unranked):
51:
35:
29:
5334:Wikispecies
5284:WikiProject
5152:Giant virus
5131:Plant virus
5049:Lytic cycle
5024:Viral entry
4726:Metaviridae
4505:Filoviridae
4491:Artoviridae
4404:Aspiviridae
4281:Mypoviridae
4260:Fimoviridae
4050:Potyviridae
3963:Secoviridae
3956:Iflaviridae
3899:Roniviridae
3805:Nidovirales
3726:Mitoviridae
3609:Duinviridae
3526:Tolivirales
3492:Nodaviridae
3418:Tymoviridae
3380:Tymovirales
3361:Togaviridae
3354:Mayoviridae
3347:Kitaviridae
3297:Hepeviridae
3290:Benyviridae
3121:Hypoviridae
2981:Totiviridae
2670:Nanoviridae
2498:Polivirales
2398:Sangervirae
2143:Nimaviridae
2125:Nudiviridae
2101:Lefavirales
1964:Vinavirales
1942:Rowavirales
1768:Ascoviridae
1746:Mimiviridae
1714:Algavirales
1655:Asfuvirales
1444:Podoviridae
1356:Uroviricota
1218:Rudiviridae
1168:Zilligvirae
461:Pleistocene
402:Replication
372:Megaviridae
360:icosahedral
349:Megaviridae
249:Description
204:giant virus
5371:Categories
5310:Pithovirus
5229:Virosphere
5204:Viral load
5194:Satellites
4980:Components
4598:Yueviridae
4554:Sunviridae
4448:Chuviridae
4209:Qinviridae
4090:Unassigned
3153:Unassigned
3018:Reoviridae
3008:Reovirales
2829:Unassigned
2760:Trapavirae
2360:Inoviridae
2157:Unassigned
2135:Unassigned
2081:Unassigned
1687:Poxviridae
1437:Myoviridae
1080:. 1450746.
1012:Virus Evol
967:Yahoo News
886:(6): 132.
579:References
453:permafrost
419:base pairs
394:and genus,
386:and genus
378:and genus
368:GC-content
245:, Russia.
239:permafrost
222:. It is a
18:Pithovirus
5325:Q18603785
5189:Oncovirus
5126:Mycovirus
5116:Virophage
5039:Viroplasm
4794:Riboviria
4665:Riboviria
4199:Muvirales
4155:Riboviria
3229:Riboviria
2906:Riboviria
2317:Loebvirae
1157:Adnaviria
1116:Baltimore
1062:Viralzone
900:1432-8798
602:17 August
543:DNA virus
509:Evolution
441:Discovery
435:cytoplasm
305:honeycomb
291:megabases
99:Kingdom:
5319:Wikidata
5260:Category
5063:Genetics
4971:Virology
4097:Families
3160:Families
2836:Families
2164:Families
1044:29511572
948:Archived
923:10 March
861:24591590
723:27389688
671:BBC News
646:87146458
568:Virology
521:See also
450:Siberian
330:proteins
235:ice core
171:Species
147:Family:
111:Phylum:
5354:1805626
5272:Commons
5099:By host
4956:History
1035:5829572
852:3964051
829:Bibcode
714:5010891
465:amoebas
243:Siberia
214:amoebas
159:Genus:
135:Order:
123:Class:
5248:Portal
5224:Virome
4987:Capsid
3178:Genera
2252:Genera
1042:
1032:
898:
859:
849:
721:
711:
644:
630:Nature
492:tundra
411:genome
352:, and
336:, the
312:genome
269:(1500
259:pithoi
216:, and
5140:Other
4945:Virus
4779:VII:
2891:III:
642:S2CID
415:dsDNA
320:genes
231:clade
85:Realm
78:Virus
5349:NCBI
4650:VI:
3214:IV:
2291:II:
1040:PMID
974:2015
925:2014
896:ISSN
857:PMID
719:PMID
604:2024
513:The
490:and
427:mRNA
406:The
310:The
4140:V:
2878:RNA
1142:I:
1129:DNA
1030:PMC
1020:doi
888:doi
884:169
847:PMC
837:doi
825:111
709:PMC
701:doi
634:doi
314:of
295:DNA
293:of
241:in
224:DNA
5373::
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4943::
4637:RT
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