225:
31:
213:
201:
187:
54:
249:
direction, with the apex bearing a field of setae. The valva mostly is reduced to the large, membranous sacculus, which reaches far out and ends in a setose field. A thin and elongate, often articulated fibula of curved shape emerges from the centre of the dorsal valva edge, reaching in a dorsal
240:
with a forewing length of 7.5–9.5 mm (0.30–0.37 in), or a wingspan of 20–29 mm (0.79–1.14 in). The forewing maculation is usually of a yellow colour, but often exhibits a distinctively red or orange postmedial (outer) area. The tribe is characterised by a number of
245:, particularly in the morphology of the genitalia. The male genitalia are rather unique among Pyraustinae and Crambidae in general in having the costa detached from the valva and projecting freely in a
917:
Nuss, Matthias; Landry, Bernard; Mally, Richard; Vegliante, Francesca; Tränkner, Andreas; Bauer, Franziska; Hayden, James; Segerer, Andreas; Schouten, Rob; Li, Houhun; Trofimova, Tatiana;
840:"Dynamic database for an inventory of the macrocaterpillar fauna, and its food plants and parasitoids, of Area de Conservacion Guanacaste (ACG), northwestern Costa Rica"
250:
direction. The narrow uncus without setae is often forked at its tip. In the female genitalia, the appendix bursae, a membranous pouch, emerges at the
922:
529:
relationships in this group had not been studied. A 2019 study eventually investigated the relationships among
Pyraustinae and the related
712:"The phylogenetic systematics of Spilomelinae and Pyraustinae (Lepidoptera: Pyraloidea: Crambidae) inferred from DNA and morphology"
224:
30:
804:
Meyrick (Lepidoptera: Pyraloidea: Crambidae: Pyraustinae) based on adult morphology and with description of a new species".
53:
633:
595:
251:
246:
711:
944:"Molecular phylogenetics and tribal classification of Japanese Pyraustinae and Spilomelinae (Lepidoptera: Crambidae)"
212:
242:
515:
466:
300:
205:
262:
structure in the corpus bursae, varies in shape among the members of
Portentomorphini: a four-armed star in
200:
862:
Robinson, Gaden S.; Ackery, Phillip R.; Kitching, Ian J.; Beccaloni, George W.; Hernández, Luis M. (2010).
983:
895:
Tominaga, S. (2010). "Hostplants of three pyraustine species observed in
Okinawa-jima Is., Okinawa Pref".
643:
408:
330:
229:
40:
1048:
1043:
370:
354:
306:
963:
942:
Matsui, Yuki; Mally, Richard; Kohama, Sari; Aoki, Itsuzai; Azuma, Masaaki; Naka, Hideshi (2022).
835:
710:
Mally, Richard; Hayden, James E.; Neinhuis, Christoph; Jordal, Bjarte H.; Nuss, Matthias (2019).
510:
336:
217:
48:
554:
943:
734:
396:
386:
775:
955:
918:
871:
831:
813:
771:
726:
682:
672:
658:
490:
360:
175:
168:
140:
186:
550:
461:
154:
667:
623:
609:
585:
495:
424:
348:
295:
259:
1037:
967:
817:
526:
457:
365:
863:
1006:
652:
530:
254:
end of the ductus bursae, close to where it transitions into the corpus bursae; in
444:
The species of
Portentomorphini are distributed in the tropics and subtropics of
1015:
580:
402:
324:
312:
291:
192:
161:
125:
105:
800:
Sutrisno, Hari; Horak, Marianne (2003). "Revision of the
Australian species of
959:
604:
569:
419:
380:
375:
318:
165:
738:
730:
564:
Portentomorphini currently comprises 44 species in the following six genera:
558:
553:. However, a study from 2022 found Portentomorphini nested within the tribe
546:
445:
171:
158:
115:
85:
65:
1000:
391:
342:
258:, however, it is laterally attached to the corpus bursae. The signum, a
549:, consequently reinstating the name Portentomorphini on the level of a
478:
471:
433:
414:
1020:
875:
449:
95:
75:
977:
618:
475:
223:
211:
199:
185:
453:
429:
237:
981:
525:
The tribe was long considered a synonym of
Pyraustinae, as the
839:
864:"HOSTS - a Database of the World's Lepidopteran Hostplants"
481:
province at an elevation of 1,700 m (5,600 ft).
795:
793:
776:"Crambidae of Aldabra Atoll (Lepidoptera: Pyraloidea)"
755:
Amsel, H. G. (1956). "Microlepidoptera
Venezolana I".
358:. Several non-Phyllanthaceae hosts are known, such as
470:, which was described from material collected in the
294:
990:
921:; De Prins, Jurate; Speidel, Wolfgang (2003–2023).
923:"Global Information System on Pyraloidea (GlobIZ)"
705:
703:
701:
8:
750:
748:
489:The tribe Portentomorphini was described by
274:, or two large, opposing granulose areas in
236:Adult Portentomorphini are relatively small
493:in 1956 based on the newly described genus
978:
29:
20:
912:
910:
857:
855:
697:
719:Arthropod Systematics & Phylogeny
174:. The tribe was initially erected by
7:
948:Insect Systematics & Evolution
759:. 10 (1954) (1–2). Maracay: 1–336.
509:, which is currently considered a
14:
757:Boletin de Entomologia Venezolana
818:10.1046/j.1440-6055.2003.00355.x
806:Australian Journal of Entomology
270:, an elongate ovoid sclerite in
52:
1:
557:, rendering the latter tribe
897:Japan Heterocerists' Journal
574:E. L. Martin in Marion, 1957
1065:
786:(1–2). Gainesville: 1–110.
960:10.1163/1876312X-bja10037
467:Hyalobathra intermedialis
301:Portentomorpha xanthialis
49:Scientific classification
47:
37:
28:
23:
731:10.26049/ASP77-1-2019-07
502:with its single species
206:Hyalobathra phoenicozona
868:Natural History Museum
331:Pioneabathra olesialis
233:
230:Pioneabathra olesialis
221:
209:
197:
41:Pioneabathra olesialis
227:
215:
203:
189:
780:Tropical Lepidoptera
657:J. C. Shaffer &
371:Sphaeranthus indicus
355:Phyllanthus urinaria
307:Margaritaria nobilis
836:Hallwachs, Winifred
547:monophyletic group
464:; an exception is
412:was reported from
337:Isocentris filalis
316:are reported from
280:Isocentris filalis
234:
222:
218:Isocentris filalis
210:
198:
1031:
1030:
984:Taxon identifiers
903:. Tokyo: 189–191.
832:Janzen, Daniel H.
772:Munroe, Eugene G.
686:
676:
662:
647:
637:
627:
613:
599:
589:
575:
521:
508:
501:
400:(Asteraceae) for
397:Helianthus annuus
387:Abrus precatorius
148:
147:
144:
24:Portentomorphini
1056:
1024:
1023:
1011:
1010:
1009:
992:Portentomorphini
979:
972:
971:
939:
933:
932:
930:
929:
914:
905:
904:
892:
886:
885:
883:
882:
876:10.5519/havt50xw
859:
850:
849:
847:
846:
828:
822:
821:
797:
788:
787:
770:Shaffer, J. C.;
767:
761:
760:
752:
743:
742:
716:
707:
681:
671:
656:
642:
632:
622:
608:
594:
584:
573:
519:
506:
499:
491:Hans Georg Amsel
361:Euphorbia virosa
190:an unidentified
176:Hans Georg Amsel
151:Portentomorphini
139:
136:Portentomorphini
57:
56:
33:
21:
1064:
1063:
1059:
1058:
1057:
1055:
1054:
1053:
1034:
1033:
1032:
1027:
1019:
1014:
1005:
1004:
999:
986:
976:
975:
941:
940:
936:
927:
925:
916:
915:
908:
894:
893:
889:
880:
878:
861:
860:
853:
844:
842:
830:
829:
825:
799:
798:
791:
769:
768:
764:
754:
753:
746:
714:
709:
708:
699:
694:
507:(Snellen, 1875)
487:
474:in the Central
462:Central America
442:
406:; furthermore,
288:
184:
138:
51:
17:
12:
11:
5:
1062:
1060:
1052:
1051:
1046:
1036:
1035:
1029:
1028:
1026:
1025:
1012:
996:
994:
988:
987:
982:
974:
973:
934:
919:Solis, M. Alma
906:
887:
851:
823:
812:(3): 233–248.
789:
762:
744:
725:(1): 141–204.
696:
695:
693:
690:
689:
688:
668:Portentomorpha
664:
649:
615:
601:
577:
537:together with
535:Portentomorpha
520:(Gueneé, 1854)
511:junior synonym
496:Portentomorpha
486:
483:
441:
438:
425:Mabra eryxalis
349:Mabra eryxalis
296:Phyllanthaceae
287:
284:
272:Portentomorpha
243:synapomorphies
183:
180:
146:
145:
133:
129:
128:
123:
119:
118:
113:
109:
108:
103:
99:
98:
93:
89:
88:
83:
79:
78:
73:
69:
68:
63:
59:
58:
45:
44:
35:
34:
26:
25:
16:Tribe of moths
15:
13:
10:
9:
6:
4:
3:
2:
1061:
1050:
1047:
1045:
1042:
1041:
1039:
1022:
1017:
1013:
1008:
1002:
998:
997:
995:
993:
989:
985:
980:
969:
965:
961:
957:
953:
949:
945:
938:
935:
924:
920:
913:
911:
907:
902:
898:
891:
888:
877:
873:
869:
865:
858:
856:
852:
841:
837:
833:
827:
824:
819:
815:
811:
807:
803:
796:
794:
790:
785:
781:
777:
773:
766:
763:
758:
751:
749:
745:
740:
736:
732:
728:
724:
720:
713:
706:
704:
702:
698:
691:
684:
680:
674:
670:
669:
665:
660:
655:
654:
650:
645:
641:
635:
631:
625:
621:
620:
616:
611:
607:
606:
602:
600:; 21 species)
597:
593:
592:Leucocraspeda
587:
583:
582:
578:
572:
571:
567:
566:
565:
562:
560:
556:
552:
548:
544:
540:
536:
532:
528:
523:
518:
517:
516:P. xanthialis
512:
505:
498:
497:
492:
484:
482:
480:
477:
473:
472:Qin Mountains
469:
468:
463:
459:
455:
451:
447:
439:
437:
435:
431:
427:
426:
421:
417:
416:
411:
410:
405:
404:
399:
398:
393:
389:
388:
383:
382:
377:
373:
372:
367:
366:Euphorbiaceae
363:
362:
357:
356:
351:
350:
345:
344:
339:
338:
333:
332:
327:
326:
321:
320:
315:
314:
310:, species of
309:
308:
303:
302:
297:
293:
285:
283:
281:
277:
273:
269:
265:
261:
257:
253:
248:
244:
239:
232:
231:
226:
220:
219:
214:
208:
207:
202:
195:
194:
188:
181:
179:
177:
173:
170:
167:
166:pyraloid moth
163:
160:
156:
152:
142:
137:
134:
131:
130:
127:
124:
121:
120:
117:
114:
111:
110:
107:
104:
101:
100:
97:
94:
91:
90:
87:
84:
81:
80:
77:
74:
71:
70:
67:
64:
61:
60:
55:
50:
46:
43:
42:
36:
32:
27:
22:
19:
991:
951:
947:
937:
926:. Retrieved
900:
896:
890:
879:. Retrieved
867:
843:. Retrieved
826:
809:
805:
801:
783:
779:
765:
756:
722:
718:
687:; 1 species)
678:
666:
653:Pioneabathra
651:
648:; 9 species)
639:
629:
617:
603:
591:
579:
568:
563:
559:paraphyletic
542:
538:
534:
531:Spilomelinae
527:phylogenetic
524:
514:
503:
494:
488:
465:
443:
440:Distribution
423:
413:
409:P. olesialis
407:
401:
395:
385:
379:
369:
359:
353:
347:
341:
335:
329:
323:
317:
311:
305:
299:
292:caterpillars
289:
279:
276:Pioneabathra
275:
271:
267:
263:
256:Pioneabathra
255:
235:
228:
216:
204:
191:
150:
149:
135:
39:
18:
1049:Moth tribes
1044:Pyraustinae
1016:iNaturalist
802:Hyalobathra
663:(2 species)
640:Streptobela
630:Neurophruda
614:(8 species)
581:Hyalobathra
576:(3 species)
543:Hyalobathra
500:Amsel, 1956
485:Systematics
456:as well as
403:Hyalobathra
325:Phyllanthus
313:Hyalobathra
286:Food plants
264:Hyalobathra
260:sclerotised
193:Hyalobathra
182:Description
162:Pyraustinae
126:Pyraustinae
122:Subfamily:
106:Lepidoptera
1038:Categories
1007:Q116521736
954:: 77–106.
928:2023-01-31
881:2023-01-31
870:. London.
845:2023-01-31
692:References
628:(synonyms
605:Isocentris
570:Cryptosara
555:Pyraustini
545:to form a
539:Cryptosara
533:and found
504:P. incalis
420:Solanaceae
381:Isocentris
376:Asteraceae
319:Glochidion
268:Cryptosara
86:Arthropoda
968:251511500
739:1863-7221
679:Apoecetes
677:(synonym
590:(synonym
446:Australia
304:feeds on
178:in 1956.
172:Crambidae
159:subfamily
116:Crambidae
72:Kingdom:
66:Eukaryota
38:Adult of
1001:Wikidata
838:(2009).
774:(2007).
392:Fabaceae
343:Flueggea
298:family:
252:anterior
112:Family:
82:Phylum:
76:Animalia
62:Domain:
610:Meyrick
586:Meyrick
479:Shaanxi
476:Chinese
434:Poaceae
422:), and
415:Solanum
196:species
164:in the
157:of the
132:Tribe:
102:Order:
96:Insecta
92:Class:
1021:892830
966:
737:
685:, 1956
683:Munroe
675:, 1956
661:, 2007
659:Munroe
646:, 1937
644:Turner
636:, 1896
634:Warren
626:, 1885
612:, 1887
598:, 1890
596:Warren
588:, 1885
450:Africa
394:) and
384:, and
378:) for
368:) and
346:, and
247:dorsal
169:family
143:, 1956
964:S2CID
715:(PDF)
673:Amsel
624:Moore
619:Mabra
551:tribe
458:South
428:from
352:from
340:from
238:moths
155:tribe
153:is a
141:Amsel
735:ISSN
541:and
460:and
454:Asia
452:and
430:rice
334:and
322:and
290:The
278:and
266:and
956:doi
901:258
872:doi
814:doi
727:doi
513:of
436:).
1040::
1018::
1003::
962:.
952:54
950:.
946:.
909:^
899:.
866:.
854:^
834:;
810:42
808:.
792:^
784:14
782:.
778:.
747:^
733:.
723:77
721:.
717:.
700:^
638:,
561:.
522:.
448:,
328:,
282:.
970:.
958::
931:.
884:.
874::
848:.
820:.
816::
741:.
729::
432:(
418:(
390:(
374:(
364:(
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