459:. After binding to the receptor, the virus enters the cell where it uncoats. Uncoating of the virus is a two step process. Firstly the outer membrane is removed as the particle enters the cell; secondly the virus particle (without the outer membrane) fuses with the cellular membrane to release the core into the cytoplasm. The pox viral genes are expressed in two phases. The early genes encode the non-structural protein, including proteins necessary for replication of the viral genome, and are expressed before the genome is replicated. The late genes are expressed after the genome has been replicated and encode the structural proteins to make the virus particle. The assembly of the virus particle occurs in five stages of maturation that lead to the final exocytosis of the new enveloped virion. After the genome has been replicated, the immature virion assembles the A5 protein to create the intracellular mature virion. The protein aligns and the brick-shaped envelope of the intracellular enveloped virion. These particles are then fused to the cell plasma to form the cell-associated enveloped virion, which encounters the microtubules and prepares to exit the cell as an extracellular enveloped virion. The assembly of the virus particle occurs in the cytoplasm of the cell and is a complex process that is currently being researched to understand each stage in more depth. Considering the fact that this virus is large and complex, replication is relatively quick taking approximately 12 hours until the host cell dies by the release of viruses.
301:
871:
which produces intracellular mature virions contained within an envelope membrane. The origin of the envelope membrane is still unknown. The intracellular mature virions are then transported to the Golgi apparatus where it is wrapped with an additional two membranes, becoming the intracellular enveloped virus. This is transported along cytoskeletal microtubules to reach the cell periphery, where it fuses with the plasma membrane to become the cell-associated enveloped virus. This triggers actin tails on cell surfaces or is released as external enveloped virion.
33:
46:
480:
2239:
2487:
2475:
365:
445:
585:
form of smallpox) which spread from Asia between 400 and 1,600 years ago. A second clade included both alastrim minor (a phenotypically mild smallpox) described from the
American continents and isolates from West Africa which diverged from an ancestral strain between 1,400 and 6,300 years before present. This clade further diverged into two subclades at least 800 years ago.
2511:
2499:
996:
that this rash was caused by smallpox by electron-microscopic examination of tiny pieces of tissue that had fallen on the shroud was unsuccessful. I was not permitted to excise one of the postules.) …The appearance of the larger pustules and the apparent distribution of the rash are similar to smallpox rashes I have seen in more recent victims
435:
of family member genomes differ considerably. Avipoxvirus, capripoxvirus, cervidpoxvirus, orthopoxvirus, suipoxvirus, yatapoxvirus and one
Entomopox genus (Betaentomopoxvirus) along with several other unclassified Entomopoxviruses have a low G+C content while others - Molluscipoxvirus, Orthopoxvirus,
386:
form of the virus, which contains different envelope, is also infectious. They vary in their shape depending upon the species but are generally shaped like a brick or as an oval form similar to a rounded brick because they are wrapped by the endoplasmic reticulum. The virion is exceptionally large,
995:
By special permission of the late
President Anwar el Sadat, I was allowed to examine the front upper half of Ramses V's unwrapped mummy in the Cairo Museum in 1979. …Inspection of the mummy revealed a rash of elevated "pustules," each about two to four millimeters in diameter, …(An attempt to prove
330:
showed that the less potent cowpox could be used to effectively vaccinate against the more deadly smallpox, a worldwide effort to vaccinate everyone against smallpox began with the ultimate goal to rid the world of the plague-like epidemic. The last case of endemic smallpox occurred in
Somalia in
1025:
Henderson, D. A.; Inglesby, Thomas V.; Bartlett, John G.; Ascher, Michael S.; Eitzen, Edward; Jahrling, Peter B.; Hauer, Jerome; Layton, Marcelle; McDade, Joseph; Osterholm, Michael T.; O'Toole, Tara; Parker, Gerald; Perl, Trish; Russell, Philip K.; Tonat, Kevin; For The
Working Group On Civilian
870:
Vaccinia contains three classes of genes: early, intermediate and late. These genes are transcribed by viral RNA polymerase and associated transcription factors. Vaccinia replicates its genome in the cytoplasm of infected cells, and after late-stage gene expression undergoes virion morphogenesis,
597:
the divergence date between variola from
Taterapox has been estimated to be 50,000 years ago. While this is consistent with the other published estimates it suggests that the archaeological and historical evidence is very incomplete. Better estimates of mutation rates in these viruses are needed.
584:
The date of the appearance of smallpox is not settled. It most likely evolved from a rodent virus between 68,000 and 16,000 years ago. The wide range of dates is due to the different records used to calibrate the molecular clock. One clade was the variola major strains (the more clinically severe
466:
genome because it occurs in the cytoplasm, although this is typical of other large DNA viruses. Poxvirus encodes its own machinery for genome transcription, a DNA dependent RNA polymerase, which makes replication in the cytoplasm possible. Most double-stranded DNA viruses require the host cell's
350:
in 2001, the
American and UK governments have had increased concern over the use of smallpox, or a smallpox-like disease, in bioterrorism. However, several poxviruses including vaccinia virus, myxoma virus, tanapox virus and raccoon pox virus are currently being investigated for their therapeutic
419:
genomes has shown that the central region of the genome is conserved and contains ~90 genes. The termini in contrast are not conserved between species. Of this group
Avipoxvirus is the most divergent. The next most divergent is Molluscipoxvirus. Capripoxvirus, Leporipoxvirus, Suipoxvirus and
867:, known for its role in the eradication of smallpox. The vaccinia virus is an effective tool for foreign protein expression, as it elicits a strong host immune-response. The vaccinia virus enters cells primarily by cell fusion, although currently the receptor responsible is unknown.
420:
Yatapoxvirus genera cluster together: Capripoxvirus and
Suipoxvirus share a common ancestor and are distinct from the genus Orthopoxvirus. Within the Othopoxvirus genus Cowpox virus strain Brighton Red, Ectromelia virus and Mpox virus do not group closely with any other member.
592:
at 3000–4000 years ago. This is consistent with archaeological and historical evidence regarding the appearance of smallpox as a human disease which suggests a relatively recent origin. However, if the mutation rate is assumed to be similar to that of the
300:
523:
The mutation rate in poxvirus genomes has been estimated to be 0.9–1.2 x 10 substitutions per site per year. A second estimate puts this rate at 0.5–7 × 10 nucleotide substitutions per site per year. A third estimate places the rate at 4–6 × 10.
1920:
Afonso PP, Silva PM, Schnellrath LC, Jesus DM, Hu J, Yang Y, Renne R, Attias M, Condit RC, Moussatché N, Damaso CR (2012) Biological characterization and next-generation genome sequencing of the unclassified Cotia virus SPAn232 (Poxviridae). J
261:(MCV). The most common are vaccinia (seen on the Indian subcontinent) and molluscum contagiosum, but Mpox infections are rising (seen in west and central African rainforest countries). The similarly named disease
304:
A) Electron micrograph of poxvirus particles in synovium of a big brown bat, northwestern United States. B) Negative staining of poxvirus particles in cell culture supernatant. Scale bar = 100 nm.
2549:
174:
serve as natural hosts. There are currently 83 species in this family, divided among 22 genera, which are divided into two subfamilies. Diseases associated with this family include
1997:
Smithson C, Meyer H, Gigante CM, Gao J, Zhao H, Batra D, Damon I, Upton C, Li Y (2017) Two novel poxviruses with unusual genome rearrangements: NY_014 and
Murmansk. Virus Genes
1484:
315:
who is thought to have died from smallpox circa 1150 years BCE. Smallpox was thought to have been transferred to Europe around the early 8th century and then to the
6457:
1488:
455:
Replication of the poxvirus involves several stages. The virus first binds to a receptor on the host cell surface; the receptors for the poxvirus are thought to be
2542:
2046:
6470:
1118:
Evgin, Laura; Vähä-Koskela, Markus; Rintoul, Julia; Falls, Theresa; Le Boeuf, Fabrice; Barrett, John W.; Bell, John C.; Stanford, Marianne M. (May 2010).
323:
within two years of introduction. This death toll can be attributed to the indigenous population's complete lack of exposure to the virus over millennia.
842:
Two more chordopoxviruses are NY_014 and murmansk poxvirus. They are considered closely related to a "Yoka poxvirus". ICTV classifies them under a genus
339:
2535:
1708:
Gjessing, MC; Yutin, N; Tengs, T; Senkevich, T; Koonin, E; Rønning, HP; Alarcon, M; Ylving, S; Lie, KI; Saure, B; Tran, L; Moss, B; Dale, OB (2015).
6431:
2405:
288:
is based on phenotypic characteristics; morphology, nucleic acid type, mode of replication, host organisms, and the type of disease they cause. The
308:
Diseases caused by pox viruses, especially smallpox, have been known about for centuries. One of the earliest suspected cases is that of Egyptian
2163:
436:
Parapoxvirus and some unclassified Chordopoxvirus - have a relatively high G+C content. The reasons for these differences are not known.
6483:
1807:
Esposito, JJ; Sammons, SA; Frace, AM; Osborne, JD; Olsen-Rasmussen, M; Zhang, M; Govil, D; Damon, IK; et al. (August 2006).
2515:
338:
In 1986, all virus samples were destroyed or transferred to two approved WHO reference labs: at the headquarters of the federal
1469:
National Center for Infectious Diseases, Centers for Disease Control and Prevention, 1600 Clifton Rd., Atlanta, GA 30333, USA.
1444:
498:
Based on the genome organisation and DNA replication mechanism a phylogenetic relationships may exist between the rudiviruses (
2111:
1012:
471:, and therefore most double-stranded DNA viruses carry out a part of their infection cycle within the host cell's nucleus.
1007:
Date of Ramses V's death derived from the Encyclopedia of Ancient Egypt, Margaret Bunson (New York: Facts On File, 2002)
833:
Cotia virus of 2012 is an unusual chordopoxvirus that may belong to a new genus. Cotia virus was assigned the new genus
4335:
383:
814:
45:
6079:
5584:
4658:
2156:
1949:
Hora, AS; Taniwaki, SA; Martins, NB; Pinto, NNR; Schlemper, AE; Santos, ALQ; Szabó, MPJ; Brandão, PE (April 2021).
284:, is a legacy of the original grouping of viruses associated with diseases that produced poxes on the skin. Modern
6475:
1757:
Wibbelt, Gudrun; Tausch, Simon H.; Dabrowski, Piotr W.; Kershaw, Olivia; Nitsche, Andreas; Schrick, Livia (2017).
829:
Both subfamilies also contain a number of unclassified species for which new genera may be created in the future.
2197:
549:. A second estimate of this divergence time places this event at 166,000 ± 43,000 years ago. The division of the
332:
238:
2558:
6516:
2192:
576:. A new systematic has been proposed recently after findings of a new squirrel poxvirus in Berlin, Germany.
250:
32:
6358:
2398:
770:
807:
2149:
258:
4846:
5320:
6488:
6418:
5897:
5419:
4614:
2893:
2562:
2450:
2445:
1875:
1820:
1396:
742:
700:
347:
285:
40:
5716:
5384:
5327:
5299:
5271:
4941:
4625:
4556:
3816:
3809:
2900:
2858:
2851:
2102:
756:
749:
714:
6511:
5911:
5765:
5544:
5292:
5285:
4105:
3303:
2921:
2907:
2865:
2778:
2503:
2440:
2425:
2187:
2027:
844:
835:
821:
793:
728:
672:
6003:
5954:
5904:
5883:
5334:
2844:
2474:
2050:
1600:
Babkin IV, Shchelkunov SN (2006) The time scale in poxvirus evolution. Mol Biol (Mosk) 40(1):20-24
5723:
4796:
4789:
3862:
3497:
3483:
3075:
2007:
1931:
1844:
1583:
1504:"Insights into the evolution of a complex virus from the crystal structure of vaccinia virus D13"
1340:
800:
686:
94:
990:
589:
491:
The ancestor of the poxviruses is not known but structural studies suggest it may have been an
467:
DNA-dependent RNA polymerase to perform transcription. These host polymerases are found in the
6405:
6019:
4978:
4853:
4832:
4725:
4381:
4039:
3681:
3244:
2837:
2764:
2415:
2260:
2091:
1980:
1903:
1836:
1788:
1739:
1690:
1636:
1575:
1533:
1478:
1424:
1332:
1291:
1265:
1206:
1198:
1157:
1139:
1100:
1082:
1043:
1008:
456:
346:, Georgia (the United States) and at the Institute of Virus Preparations in Moscow. After the
6462:
1864:"On the origin of smallpox: correlating variola phylogenics with historical smallpox records"
5809:
5737:
5630:
5205:
5147:
5110:
4839:
4570:
4528:
4476:
4439:
4409:
4370:
4224:
4150:
4007:
3329:
3292:
3266:
3224:
3025:
3015:
2830:
2682:
2653:
2491:
2311:
2247:
2081:
2073:
1970:
1962:
1893:
1883:
1828:
1778:
1770:
1729:
1721:
1680:
1672:
1626:
1567:
1523:
1515:
1414:
1404:
1322:
1281:
1188:
1147:
1131:
1090:
1074:
1035:
1026:
Biodefense (1999). "Smallpox as a Biological Weapon: Medical and Public Health Management".
785:
707:
650:
618:
608:
416:
269:
201:
6396:
1311:"Genus specific evolution of codon usage and nucleotide compositional traits of poxviruses"
1078:
839:
in 2019. A Brazilian porcupinepox virus discovered in 2019 is closely related to the virus.
479:
6269:
6140:
5975:
5923:
5836:
5792:
5781:
5619:
5471:
5412:
5236:
4957:
4887:
4775:
4758:
4693:
4290:
4245:
3966:
3473:
3255:
3166:
3086:
2982:
2321:
2306:
2301:
2285:
2238:
2115:
1225:
880:
512:
106:
1553:"Exceptionally diverse morphotypes and genomes of crenarchaeal hyperthermophilic viruses"
572:
An isolate from a fish – salmon gill poxvirus – appears to be the earliest branch in the
1879:
1824:
1400:
6312:
6290:
6258:
6223:
6129:
6094:
6030:
5873:
5862:
5825:
5667:
5551:
5441:
5370:
5360:
5121:
4913:
4876:
4782:
4577:
4161:
4134:
3781:
3770:
3674:
3639:
3632:
3618:
3611:
3560:
3532:
3462:
3451:
3416:
3200:
3003:
2872:
2809:
2743:
2721:
2643:
2479:
2410:
2378:
2265:
2223:
1975:
1950:
1898:
1863:
1783:
1758:
1734:
1709:
1685:
1660:
1528:
1503:
1419:
1384:
1152:
1120:"Potent oncolytic activity of raccoonpox virus in the absence of natural pathogenicity"
1119:
1095:
1062:
983:
864:
858:
693:
679:
425:
379:
2121:
2086:
2061:
904:
6505:
6280:
6175:
6161:
5989:
5982:
5961:
5802:
5751:
5744:
5608:
5504:
5482:
5455:
5431:
5391:
5348:
5313:
5306:
5278:
5264:
5257:
5158:
5094:
5065:
5044:
5023:
5012:
4992:
4924:
4746:
4715:
4704:
4682:
4549:
4416:
4402:
4392:
4359:
4283:
4235:
4178:
4171:
4018:
3918:
3792:
3706:
3688:
3667:
3653:
3553:
3490:
3440:
3362:
3340:
3313:
3276:
3178:
3145:
2993:
2971:
2771:
2754:
2732:
2710:
2632:
2621:
2430:
2342:
2275:
2228:
721:
665:
557:
diverged ~137,000 ± 35,000 years ago. This was followed by the ancestor of the genus
421:
327:
230:
183:
1848:
1587:
1552:
1344:
6410:
6302:
6247:
6182:
6151:
6118:
5968:
5940:
5758:
5730:
5709:
5695:
5688:
5678:
5514:
5448:
5377:
5225:
5195:
5184:
5131:
5087:
5077:
5058:
5034:
4985:
4897:
4810:
4768:
4607:
4539:
4517:
4497:
4487:
4297:
4213:
4124:
4095:
4083:
4073:
4061:
4051:
4029:
3987:
3977:
3950:
3929:
3907:
3883:
3873:
3851:
3748:
3699:
3660:
3646:
3625:
3394:
3372:
3350:
3217:
3119:
3107:
3064:
3053:
2960:
2949:
2914:
2820:
2672:
2527:
2393:
2388:
2352:
2337:
2316:
2172:
2136:
1470:
777:
735:
506:
492:
468:
331:
1977. Extensive searches over two years detected no further cases, and in 1979 the
234:
195:
189:
82:
70:
2108:
1809:"Genome sequence diversity and clues to the evolution of variola (smallpox) virus"
1175:
Suryawanshi, Yogesh R.; Zhang, Tiantian; Razi, Farzad; Essani, Karim (July 2020).
538:
diverged from the ancestor 249 ± 69 thousand years ago. The ancestor of the genus
527:
The last common ancestor of the extant poxviruses that infect vertebrates existed
1676:
6444:
6390:
6168:
5947:
5933:
5846:
5702:
5492:
5405:
5398:
5341:
5247:
5168:
5051:
4968:
4934:
4860:
4822:
4803:
4739:
4732:
4563:
4423:
4112:
3940:
3840:
3735:
3585:
3567:
3543:
3406:
3384:
3210:
3188:
3156:
3097:
2886:
2798:
2660:
2610:
2586:
2383:
2362:
2280:
2255:
2077:
885:
763:
658:
594:
500:
364:
266:
1951:"Genomic Analysis of Novel Poxvirus Brazilian Porcupinepox Virus, Brazil, 2019"
1473:. National Center for Biotechnology Information (NCBI), NIH, Bethesda, MD, USA.
444:
6040:
5996:
5890:
5651:
4460:
4450:
4202:
3802:
2879:
2460:
2435:
1519:
1327:
1310:
613:
432:
400:
262:
167:
6381:
1202:
1193:
1176:
1143:
1086:
1039:
6236:
6107:
5641:
5597:
4671:
4348:
4320:
3759:
2599:
2571:
2420:
2357:
2347:
2270:
1888:
1832:
1759:"Berlin Squirrelpox Virus, a New Poxvirus in Red Squirrels, Berlin, Germany"
1409:
392:
388:
171:
1984:
1966:
1907:
1862:
Li, Y; Carroll, DS; Gardner, SN; Walsh, MC; Vitalis, EA; Damon, IK (2007).
1840:
1792:
1774:
1743:
1694:
1640:
1579:
1537:
1428:
1336:
1295:
1210:
1161:
1104:
1047:
2129:
2095:
1710:"Salmon Gill Poxvirus, the Deepest Representative of the Chordopoxvirinae"
1286:
1269:
6375:
2202:
2042:
1725:
316:
312:
289:
214:
210:
175:
462:
The replication of poxvirus is unusual for a virus with double-stranded
351:
potential in various human cancers in preclinical and clinical studies.
1571:
1245:
1135:
428:
form a subgroup. Vaccinia virus is most closely related to CPV-GRI-90.
343:
309:
246:
163:
6449:
2141:
1631:
1614:
1250:
2455:
2218:
623:
544:
529:
504:) and the large eukaryal DNA viruses: the African swine fever virus (
399:
in a single, linear, double-stranded segment of DNA. By comparison,
396:
320:
218:
6436:
6352:
1808:
1358:
487:
and distribution of cGAMP nucleases across member species and genera
2176:
2038:
1177:"Tanapoxvirus: From discovery towards oncolytic immunovirotherapy"
955:
495:
or a species related to both the poxviruses and the adenoviruses.
478:
443:
363:
319:
in the early 16th century, resulting in the deaths of 3.2 million
299:
57:
1383:
Mutsafi, Y; Zauberman, N; Sabanay, I; Minsky, A (30 March 2010).
985:
The greatest killer: smallpox in history, with a new introduction
2137:
Virus Pathogen Database and Analysis Resource (ViPR): Poxviridae
222:
6356:
6220:
6091:
5581:
4655:
4332:
3732:
2583:
2531:
2145:
1224:
International Committee on Taxonomy of Viruses (15 June 2004).
929:
1385:"Vaccinia-like cytoplasmic replication of the giant Mimivirus"
463:
160:
6423:
553:
into the extant genera occurred ~14,000 years ago. The genus
2008:"Taxonomy History [Yokapox virus] - Taxonomy - ICTV"
1615:"Molecular dating in the evolution of vertebrate poxviruses"
1445:"Pithovirus: Bigger than Pandoravirus with a smaller genome"
588:
A second estimate has placed the separation of variola from
1932:"Taxonomy History [Cotia virus] - Taxonomy - ICTV"
2109:
Poxviridae at the Viral Bioinformatics Resource Center
1502:
Bahar, MW; Graham, SC; Stuart, DI; Grimes, JM (2011).
637:
The following subfamilies and genera are recognized (-
1471:"DNA-dependent RNA polymerase rpo35 (Vaccinia virus)"
1061:
Chan, Winnie M.; McFadden, Grant (1 September 2014).
1028:
JAMA: The Journal of the American Medical Association
292:
virus remains the most notable member of the family.
6365:
6300:
6278:
6267:
6256:
6245:
6234:
6149:
6138:
6127:
6116:
6105:
6078:
6028:
6017:
5921:
5871:
5860:
5834:
5823:
5790:
5779:
5676:
5665:
5639:
5628:
5617:
5606:
5595:
5532:
5502:
5480:
5469:
5429:
5358:
5245:
5234:
5223:
5193:
5182:
5156:
5145:
5119:
5108:
5075:
5032:
5021:
5010:
4966:
4955:
4922:
4911:
4885:
4874:
4820:
4756:
4713:
4702:
4691:
4680:
4669:
4595:
4537:
4526:
4515:
4485:
4474:
4448:
4437:
4390:
4379:
4368:
4357:
4346:
4319:
4271:
4233:
4222:
4211:
4200:
4159:
4148:
4122:
4093:
4071:
4049:
4027:
4016:
4005:
3975:
3964:
3938:
3927:
3916:
3905:
3871:
3860:
3849:
3838:
3790:
3779:
3768:
3757:
3746:
3599:
3577:
3541:
3530:
3523:
3471:
3460:
3449:
3438:
3404:
3382:
3360:
3338:
3327:
3301:
3290:
3264:
3253:
3242:
3198:
3176:
3154:
3143:
3117:
3095:
3084:
3073:
3062:
3051:
3013:
2991:
2980:
2969:
2958:
2947:
2818:
2807:
2796:
2752:
2741:
2730:
2719:
2708:
2670:
2641:
2630:
2619:
2608:
2597:
2570:
2371:
2330:
2294:
2246:
2211:
1389:
Proceedings of the National Academy of Sciences USA
982:
956:"Pathogenic Molluscum Contagiosum Virus Sequenced"
335:(WHO) declared the disease officially eradicated.
2060:Buller, R. Mark L.; Palumbo, Gregory J. (1991).
936:. International Committee on Taxonomy of Viruses
2047:International Committee on Taxonomy of Viruses
1659:Hughes, AL; Irausquin, S; Friedman, R (2010).
1309:Roychoudhury, S; Pan, A; Mukherjee, D (2011).
2543:
2157:
1654:
1652:
1650:
1608:
1606:
542:was next to diverge from the other clades at
181:Four genera of poxviruses may infect humans:
8:
1483:: CS1 maint: multiple names: authors list (
265:is not a true poxvirus and is caused by the
1270:"Poxvirus genomes: a phylogenetic analysis"
1181:Journal of Cancer Research and Therapeutics
6353:
6275:
6264:
6253:
6242:
6231:
6217:
6146:
6135:
6124:
6113:
6102:
6088:
6084:
6025:
5868:
5831:
5787:
5673:
5636:
5625:
5614:
5603:
5592:
5578:
5477:
5242:
5231:
5190:
5153:
5116:
5029:
5018:
4963:
4919:
4882:
4710:
4699:
4688:
4677:
4666:
4652:
4534:
4523:
4482:
4445:
4387:
4376:
4365:
4354:
4343:
4329:
4325:
4230:
4219:
4208:
4156:
4024:
4013:
3972:
3935:
3924:
3913:
3868:
3857:
3846:
3787:
3776:
3765:
3754:
3743:
3729:
3538:
3527:
3468:
3457:
3446:
3335:
3298:
3261:
3250:
3151:
3092:
3081:
3070:
3059:
2988:
2977:
2966:
2955:
2815:
2804:
2749:
2738:
2727:
2716:
2638:
2627:
2616:
2605:
2594:
2580:
2576:
2550:
2536:
2528:
2164:
2150:
2142:
2066:Microbiology and Molecular Biology Reviews
1487:) CS1 maint: numeric names: authors list (
340:Centers for Disease Control and Prevention
31:
20:
2085:
1974:
1897:
1887:
1782:
1733:
1684:
1630:
1527:
1418:
1408:
1326:
1285:
1192:
1151:
1094:
626:. There are ten recognised genera in the
2406:Laboratory diagnosis of viral infections
1661:"The evolutionary biology of poxviruses"
382:(external enveloped virion), though the
378:viral particles (virions) are generally
1359:"Orthopoxvirus replication ~ ViralZone"
989:. University of Chicago Press. p.
896:
1476:
1079:10.1146/annurev-virology-031413-085442
924:
922:
415:Phylogenetic analysis of 26 different
1551:Prangishvili, D; Garrett, RA (2004).
1226:"ICTVdb Descriptions: 58. Poxviridae"
569:diverged 111,000 ± 29,000 years ago.
7:
2498:
1443:Racaniello, Vincent (4 March 2014).
2510:
2045:virus, have been collected by the
561:. The last common ancestor of the
14:
2509:
2497:
2486:
2485:
2473:
2237:
1613:Babkin, IV; Babkina, IN (2011).
44:
403:are 1/10 as large as a typical
159:is a family of double-stranded
1264:Gubser, C; Hué, S; Kellam, P;
930:"Virus Taxonomy: 2019 Release"
815:Diachasmimorpha entomopoxvirus
1:
1797:(for systematic see figure 2)
863:The prototypical poxvirus is
606:The species in the subfamily
145:
1955:Emerging Infectious Diseases
1763:Emerging Infectious Diseases
1677:10.1016/j.meegid.2009.10.001
981:Hopkins, Donald R. (2002) .
2078:10.1128/mr.55.1.80-122.1991
616:and those in the subfamily
384:intracellular mature virion
259:molluscum contagiosum virus
6533:
856:
395:in length and carries its
6230:
6216:
6101:
6087:
5591:
5577:
4665:
4651:
4342:
4328:
3742:
3728:
2593:
2579:
2469:
2235:
2198:Social history of viruses
2183:
1520:10.1016/j.str.2011.03.023
1328:10.1007/s11262-010-0568-2
1067:Annual Review of Virology
960:Antiviral Agents Bulletin
333:World Health Organization
239:bovine papular stomatitis
143:
138:
39:
30:
23:
1815:(Submitted manuscript).
1562:(Submitted manuscript).
1254:. Retrieved 2005-02-26.
1194:10.4103/jcrt.JCRT_157_18
1040:10.1001/jama.281.22.2127
280:The name of the family,
2062:"Poxvirus Pathogenesis"
1889:10.1073/pnas.0609268104
1833:10.1126/science.1125134
1410:10.1073/pnas.0912737107
1246:How Big is a ... ?
641:denotes subfamily and -
547: million years ago
532: million years ago
387:its size is around 200
251:yaba monkey tumor virus
2399:Helper dependent virus
2114:5 January 2012 at the
1967:10.3201/eid2704.203818
1868:Proc Natl Acad Sci USA
1775:10.3201/eid2310.171008
1063:"Oncolytic Poxviruses"
510:), Chlorella viruses (
488:
452:
372:
305:
1287:10.1099/vir.0.19565-0
483:Phylogenetic tree of
482:
447:
367:
303:
2563:virus classification
2451:Virus quantification
2446:Virus classification
2028:Electron micrographs
1726:10.1128/JVI.01174-15
1363:viralzone.expasy.org
962:: 196–7. August 1996
771:vespertilionpoxvirus
391:in diameter and 300
348:September 11 attacks
286:viral classification
41:Virus classification
5545:Permutotetraviridae
3304:Orthopolintovirales
2779:Malacoherpesviridae
2441:Virus-like particle
2012:talk.ictvonline.org
1936:talk.ictvonline.org
1880:2007PNAS..10415787L
1874:(40): 15787–15792.
1825:2006Sci...313..807E
1401:2010PNAS..107.5978M
934:talk.ictvonline.org
822:Gammaentomopoxvirus
808:Deltaentomopoxvirus
794:Alphaentomopoxvirus
3863:Malgrandaviricetes
3498:Sphaerolipoviridae
3484:Matshushitaviridae
3076:Nucleocytoviricota
2103:NCBI Taxonomy Page
1572:10.1042/bst0320204
1136:10.1038/mt.2010.14
801:Betaentomopoxvirus
687:Crocodylidpoxvirus
516:) and poxviruses (
489:
457:glycosaminoglycans
453:
373:
306:
96:Nucleocytoviricota
6499:
6498:
6359:Taxon identifiers
6350:
6349:
6346:
6345:
6342:
6341:
6338:
6337:
6334:
6333:
6330:
6329:
6326:
6325:
6322:
6321:
6212:
6211:
6208:
6207:
6204:
6203:
6200:
6199:
6196:
6195:
6192:
6191:
6074:
6073:
6070:
6069:
6066:
6065:
6062:
6061:
6058:
6057:
6054:
6053:
6050:
6049:
6020:Yunchangviricetes
6013:
6012:
5856:
5855:
5819:
5818:
5775:
5774:
5661:
5660:
5573:
5572:
5569:
5568:
5565:
5564:
5561:
5560:
5528:
5527:
5524:
5523:
5465:
5464:
5219:
5218:
5215:
5214:
5178:
5177:
5141:
5140:
5104:
5103:
5006:
5005:
5002:
5001:
4979:Carmotetraviridae
4951:
4950:
4907:
4906:
4870:
4869:
4854:Gammaflexiviridae
4847:Deltaflexiviridae
4833:Alphaflexiviridae
4726:Alphatetraviridae
4647:
4646:
4643:
4642:
4639:
4638:
4635:
4634:
4591:
4590:
4587:
4586:
4511:
4510:
4507:
4506:
4470:
4469:
4433:
4432:
4382:Chrymotiviricetes
4315:
4314:
4311:
4310:
4307:
4306:
4267:
4266:
4263:
4262:
4259:
4258:
4255:
4254:
4196:
4195:
4192:
4191:
4188:
4187:
4144:
4143:
4040:Bacilladnaviridae
4001:
4000:
3997:
3996:
3960:
3959:
3901:
3900:
3897:
3896:
3893:
3892:
3834:
3833:
3830:
3829:
3826:
3825:
3724:
3723:
3720:
3719:
3716:
3715:
3682:Portogloboviridae
3595:
3594:
3519:
3518:
3515:
3514:
3511:
3510:
3507:
3506:
3434:
3433:
3430:
3429:
3426:
3425:
3323:
3322:
3286:
3285:
3245:Preplasmiviricota
3238:
3237:
3234:
3233:
3139:
3138:
3047:
3046:
3043:
3042:
3039:
3038:
3035:
3034:
2943:
2942:
2939:
2938:
2935:
2934:
2931:
2930:
2838:Autographiviridae
2792:
2791:
2788:
2787:
2765:Alloherpesviridae
2704:
2703:
2700:
2699:
2696:
2695:
2692:
2691:
2525:
2524:
2416:Neurotropic virus
2261:Viral replication
1819:(5788): 807–812.
1769:(10): 1726–1729.
1720:(18): 9348–9867.
1665:Infect Genet Evol
1632:10.1159/000320964
1560:Biochem Soc Trans
1124:Molecular Therapy
630:and three in the
451:replication cycle
213:virus (variola),
152:
151:
16:Family of viruses
6524:
6492:
6491:
6479:
6478:
6466:
6465:
6453:
6452:
6440:
6439:
6427:
6426:
6414:
6413:
6401:
6400:
6399:
6386:
6385:
6384:
6354:
6276:
6265:
6254:
6243:
6232:
6224:dsDNA-RT viruses
6218:
6147:
6136:
6125:
6114:
6103:
6095:ssRNA-RT viruses
6089:
6085:
6026:
5869:
5837:Serpentovirales
5832:
5810:Orthomyxoviridae
5788:
5738:Peribunyaviridae
5674:
5637:
5631:Chunqiuviricetes
5626:
5615:
5604:
5593:
5585:(–)ssRNA viruses
5579:
5478:
5321:Nanghoshaviridae
5243:
5232:
5206:Botourmiaviridae
5191:
5154:
5148:Howeltoviricetes
5117:
5111:Amabiliviricetes
5030:
5019:
4964:
4920:
4883:
4840:Betaflexiviridae
4711:
4700:
4689:
4678:
4667:
4659:(+)ssRNA viruses
4653:
4571:Picobirnaviridae
4535:
4529:Duplopiviricetes
4524:
4483:
4477:Vidaverviricetes
4446:
4440:Resentoviricetes
4410:Megabirnaviridae
4388:
4377:
4371:Duplornaviricota
4366:
4355:
4344:
4330:
4326:
4231:
4225:Huolimaviricetes
4220:
4209:
4157:
4151:Repensiviricetes
4025:
4014:
4008:Cressdnaviricota
3973:
3936:
3925:
3914:
3869:
3858:
3847:
3788:
3777:
3766:
3755:
3744:
3730:
3539:
3528:
3469:
3458:
3447:
3336:
3330:Tectiliviricetes
3299:
3293:Polintoviricetes
3267:Priklausovirales
3262:
3251:
3225:Marseilleviridae
3152:
3093:
3082:
3071:
3060:
3026:Papillomaviridae
3016:Zurhausenvirales
2989:
2978:
2967:
2956:
2831:Ackermannviridae
2816:
2805:
2750:
2739:
2728:
2717:
2683:Tristromaviridae
2654:Lipothrixviridae
2639:
2628:
2617:
2606:
2595:
2581:
2577:
2552:
2545:
2538:
2529:
2513:
2512:
2501:
2500:
2489:
2488:
2477:
2312:Phenotype mixing
2248:Viral life cycle
2241:
2166:
2159:
2152:
2143:
2099:
2089:
2041:, including the
2016:
2015:
2004:
1998:
1995:
1989:
1988:
1978:
1961:(4): 1177–1180.
1946:
1940:
1939:
1928:
1922:
1918:
1912:
1911:
1901:
1891:
1859:
1853:
1852:
1804:
1798:
1796:
1786:
1754:
1748:
1747:
1737:
1705:
1699:
1698:
1688:
1656:
1645:
1644:
1634:
1610:
1601:
1598:
1592:
1591:
1557:
1548:
1542:
1541:
1531:
1514:(7): 1011–1020.
1499:
1493:
1492:
1482:
1474:
1466:
1460:
1459:
1457:
1455:
1440:
1434:
1432:
1422:
1412:
1380:
1374:
1373:
1371:
1369:
1355:
1349:
1348:
1330:
1306:
1300:
1299:
1289:
1261:
1255:
1243:
1237:
1236:
1234:
1232:
1221:
1215:
1214:
1196:
1172:
1166:
1165:
1155:
1115:
1109:
1108:
1098:
1058:
1052:
1051:
1022:
1016:
1005:
999:
998:
988:
978:
972:
971:
969:
967:
952:
946:
945:
943:
941:
926:
917:
916:
914:
912:
901:
786:Entomopoxvirinae
708:Molluscipoxvirus
651:Chordopoxvirinae
632:Entomopoxvirinae
628:Chordopoxvirinae
619:Entomopoxvirinae
609:Chordopoxvirinae
574:Chordopoxvirinae
548:
533:
342:(the C.D.C.) in
326:A century after
270:varicella zoster
255:Molluscipoxvirus
202:Molluscipoxvirus
49:
48:
35:
21:
6532:
6531:
6527:
6526:
6525:
6523:
6522:
6521:
6502:
6501:
6500:
6495:
6487:
6482:
6474:
6469:
6461:
6456:
6448:
6443:
6435:
6430:
6422:
6417:
6409:
6404:
6395:
6394:
6389:
6380:
6379:
6374:
6361:
6351:
6318:
6296:
6270:Revtraviricetes
6226:
6188:
6141:Revtraviricetes
6097:
6046:
6009:
5976:Paramyxoviridae
5924:Mononegavirales
5917:
5898:Crepuscuviridae
5852:
5815:
5793:Articulavirales
5782:Insthoviricetes
5771:
5657:
5620:Negarnaviricota
5587:
5557:
5520:
5498:
5472:Stelpaviricetes
5461:
5425:
5420:Polycipiviridae
5413:Dicistroviridae
5354:
5237:Pisoniviricetes
5211:
5174:
5137:
5100:
5071:
4998:
4958:Tolucaviricetes
4947:
4903:
4888:Amarillovirales
4866:
4816:
4776:Closteroviridae
4759:Martellivirales
4752:
4694:Kitrinoviricota
4661:
4631:
4615:Polymycoviridae
4583:
4503:
4466:
4429:
4338:
4303:
4291:Finnlakeviridae
4251:
4246:Pleolipoviridae
4184:
4140:
4118:
4089:
4067:
4045:
3993:
3967:Quintoviricetes
3956:
3889:
3822:
3738:
3712:
3591:
3573:
3503:
3474:Halopanivirales
3422:
3400:
3378:
3356:
3319:
3282:
3256:Maveriviricetes
3230:
3194:
3172:
3167:Phycodnaviridae
3135:
3113:
3087:Pokkesviricetes
3031:
3009:
2983:Papovaviricetes
2927:
2894:Rountreeviridae
2784:
2688:
2666:
2589:
2566:
2556:
2526:
2521:
2465:
2367:
2326:
2322:Viral evolution
2307:Antigenic shift
2302:Antigenic drift
2290:
2286:Lysogenic cycle
2242:
2233:
2207:
2179:
2170:
2116:Wayback Machine
2059:
2054:picture gallery
2024:
2019:
2006:
2005:
2001:
1996:
1992:
1948:
1947:
1943:
1930:
1929:
1925:
1919:
1915:
1861:
1860:
1856:
1806:
1805:
1801:
1756:
1755:
1751:
1707:
1706:
1702:
1658:
1657:
1648:
1612:
1611:
1604:
1599:
1595:
1555:
1550:
1549:
1545:
1501:
1500:
1496:
1475:
1468:
1467:
1463:
1453:
1451:
1442:
1441:
1437:
1395:(13): 5978–82.
1382:
1381:
1377:
1367:
1365:
1357:
1356:
1352:
1308:
1307:
1303:
1263:
1262:
1258:
1244:
1240:
1230:
1228:
1223:
1222:
1218:
1174:
1173:
1169:
1117:
1116:
1112:
1060:
1059:
1055:
1034:(22): 2127–37.
1024:
1023:
1019:
1006:
1002:
980:
979:
975:
965:
963:
954:
953:
949:
939:
937:
928:
927:
920:
910:
908:
903:
902:
898:
894:
881:Quokkapox virus
877:
861:
855:
848:, created 2016.
827:
743:Pteropopoxvirus
701:Macropopoxvirus
645:denotes genus):
604:
582:
543:
528:
513:Phycodnaviridae
477:
442:
413:
362:
357:
298:
278:
108:Pokkesviricetes
43:
17:
12:
11:
5:
6530:
6528:
6520:
6519:
6517:Virus families
6514:
6504:
6503:
6497:
6496:
6494:
6493:
6480:
6467:
6454:
6441:
6428:
6415:
6402:
6387:
6371:
6369:
6363:
6362:
6357:
6348:
6347:
6344:
6343:
6340:
6339:
6336:
6335:
6332:
6331:
6328:
6327:
6324:
6323:
6320:
6319:
6317:
6316:
6313:Caulimoviridae
6308:
6306:
6298:
6297:
6295:
6294:
6291:Hepadnaviridae
6286:
6284:
6273:
6262:
6259:Artverviricota
6251:
6240:
6228:
6227:
6221:
6214:
6213:
6210:
6209:
6206:
6205:
6202:
6201:
6198:
6197:
6194:
6193:
6190:
6189:
6187:
6186:
6179:
6172:
6165:
6157:
6155:
6144:
6133:
6130:Artverviricota
6122:
6111:
6099:
6098:
6092:
6082:
6076:
6075:
6072:
6071:
6068:
6067:
6064:
6063:
6060:
6059:
6056:
6055:
6052:
6051:
6048:
6047:
6045:
6044:
6036:
6034:
6031:Goujianvirales
6023:
6015:
6014:
6011:
6010:
6008:
6007:
6000:
5993:
5986:
5979:
5972:
5965:
5958:
5951:
5944:
5937:
5929:
5927:
5919:
5918:
5916:
5915:
5908:
5901:
5894:
5887:
5879:
5877:
5874:Jingchuvirales
5866:
5863:Monjiviricetes
5858:
5857:
5854:
5853:
5851:
5850:
5842:
5840:
5829:
5826:Milneviricetes
5821:
5820:
5817:
5816:
5814:
5813:
5806:
5798:
5796:
5785:
5777:
5776:
5773:
5772:
5770:
5769:
5762:
5755:
5748:
5741:
5734:
5727:
5720:
5717:Leishbuviridae
5713:
5706:
5699:
5692:
5684:
5682:
5671:
5668:Ellioviricetes
5663:
5662:
5659:
5658:
5656:
5655:
5647:
5645:
5634:
5623:
5612:
5601:
5589:
5588:
5582:
5575:
5574:
5571:
5570:
5567:
5566:
5563:
5562:
5559:
5558:
5556:
5555:
5552:Sarthroviridae
5548:
5536:
5534:
5530:
5529:
5526:
5525:
5522:
5521:
5519:
5518:
5510:
5508:
5500:
5499:
5497:
5496:
5488:
5486:
5475:
5467:
5466:
5463:
5462:
5460:
5459:
5452:
5445:
5442:Alvernaviridae
5437:
5435:
5427:
5426:
5424:
5423:
5416:
5409:
5402:
5395:
5388:
5385:Solinviviridae
5381:
5374:
5371:Picornaviridae
5366:
5364:
5361:Picornavirales
5356:
5355:
5353:
5352:
5345:
5338:
5331:
5328:Nanhypoviridae
5324:
5317:
5310:
5303:
5300:Medioniviridae
5296:
5289:
5282:
5275:
5272:Cremegaviridae
5268:
5261:
5253:
5251:
5240:
5229:
5221:
5220:
5217:
5216:
5213:
5212:
5210:
5209:
5201:
5199:
5188:
5180:
5179:
5176:
5175:
5173:
5172:
5164:
5162:
5151:
5143:
5142:
5139:
5138:
5136:
5135:
5127:
5125:
5122:Wolframvirales
5114:
5106:
5105:
5102:
5101:
5099:
5098:
5091:
5083:
5081:
5073:
5072:
5070:
5069:
5062:
5055:
5048:
5040:
5038:
5027:
5016:
5008:
5007:
5004:
5003:
5000:
4999:
4997:
4996:
4989:
4982:
4974:
4972:
4961:
4953:
4952:
4949:
4948:
4946:
4945:
4942:Sinhaliviridae
4938:
4930:
4928:
4917:
4914:Magsaviricetes
4909:
4908:
4905:
4904:
4902:
4901:
4893:
4891:
4880:
4877:Flasuviricetes
4872:
4871:
4868:
4867:
4865:
4864:
4857:
4850:
4843:
4836:
4828:
4826:
4818:
4817:
4815:
4814:
4807:
4800:
4793:
4786:
4783:Endornaviridae
4779:
4772:
4764:
4762:
4754:
4753:
4751:
4750:
4743:
4736:
4729:
4721:
4719:
4708:
4697:
4686:
4675:
4663:
4662:
4656:
4649:
4648:
4645:
4644:
4641:
4640:
4637:
4636:
4633:
4632:
4630:
4629:
4626:Botybirnavirus
4618:
4611:
4599:
4597:
4593:
4592:
4589:
4588:
4585:
4584:
4582:
4581:
4578:Partitiviridae
4574:
4567:
4560:
4557:Curvulaviridae
4553:
4545:
4543:
4532:
4521:
4513:
4512:
4509:
4508:
4505:
4504:
4502:
4501:
4493:
4491:
4480:
4472:
4471:
4468:
4467:
4465:
4464:
4456:
4454:
4443:
4435:
4434:
4431:
4430:
4428:
4427:
4420:
4413:
4406:
4398:
4396:
4385:
4374:
4363:
4352:
4340:
4339:
4333:
4323:
4317:
4316:
4313:
4312:
4309:
4308:
4305:
4304:
4302:
4301:
4294:
4287:
4275:
4273:
4269:
4268:
4265:
4264:
4261:
4260:
4257:
4256:
4253:
4252:
4250:
4249:
4241:
4239:
4228:
4217:
4206:
4198:
4197:
4194:
4193:
4190:
4189:
4186:
4185:
4183:
4182:
4175:
4167:
4165:
4162:Geplafuvirales
4154:
4146:
4145:
4142:
4141:
4139:
4138:
4135:Redondoviridae
4130:
4128:
4120:
4119:
4117:
4116:
4109:
4101:
4099:
4091:
4090:
4088:
4087:
4079:
4077:
4069:
4068:
4066:
4065:
4057:
4055:
4047:
4046:
4044:
4043:
4035:
4033:
4022:
4011:
4003:
4002:
3999:
3998:
3995:
3994:
3992:
3991:
3983:
3981:
3970:
3962:
3961:
3958:
3957:
3955:
3954:
3946:
3944:
3933:
3922:
3911:
3903:
3902:
3899:
3898:
3895:
3894:
3891:
3890:
3888:
3887:
3879:
3877:
3866:
3855:
3844:
3836:
3835:
3832:
3831:
3828:
3827:
3824:
3823:
3821:
3820:
3817:Plectroviridae
3813:
3810:Paulinoviridae
3806:
3798:
3796:
3785:
3782:Faserviricetes
3774:
3771:Hofneiviricota
3763:
3752:
3740:
3739:
3733:
3726:
3725:
3722:
3721:
3718:
3717:
3714:
3713:
3711:
3710:
3703:
3692:
3685:
3678:
3675:Polydnaviridae
3671:
3664:
3657:
3650:
3643:
3640:Globuloviridae
3636:
3633:Fuselloviridae
3629:
3622:
3619:Bicaudaviridae
3615:
3612:Ampullaviridae
3603:
3601:
3597:
3596:
3593:
3592:
3590:
3589:
3581:
3579:
3575:
3574:
3572:
3571:
3564:
3561:Hytrosaviridae
3557:
3549:
3547:
3536:
3533:Naldaviricetes
3525:
3521:
3520:
3517:
3516:
3513:
3512:
3509:
3508:
3505:
3504:
3502:
3501:
3494:
3487:
3479:
3477:
3466:
3463:Laserviricetes
3455:
3452:Dividoviricota
3444:
3436:
3435:
3432:
3431:
3428:
3427:
3424:
3423:
3421:
3420:
3417:Corticoviridae
3412:
3410:
3402:
3401:
3399:
3398:
3390:
3388:
3380:
3379:
3377:
3376:
3368:
3366:
3358:
3357:
3355:
3354:
3346:
3344:
3333:
3325:
3324:
3321:
3320:
3318:
3317:
3309:
3307:
3296:
3288:
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3284:
3283:
3281:
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3272:
3270:
3259:
3248:
3240:
3239:
3236:
3235:
3232:
3231:
3229:
3228:
3221:
3214:
3206:
3204:
3201:Pimascovirales
3196:
3195:
3193:
3192:
3184:
3182:
3174:
3173:
3171:
3170:
3162:
3160:
3149:
3141:
3140:
3137:
3136:
3134:
3133:
3125:
3123:
3115:
3114:
3112:
3111:
3103:
3101:
3090:
3079:
3068:
3057:
3049:
3048:
3045:
3044:
3041:
3040:
3037:
3036:
3033:
3032:
3030:
3029:
3021:
3019:
3011:
3010:
3008:
3007:
3004:Polyomaviridae
2999:
2997:
2986:
2975:
2964:
2953:
2945:
2944:
2941:
2940:
2937:
2936:
2933:
2932:
2929:
2928:
2926:
2925:
2918:
2911:
2904:
2901:Salasmaviridae
2897:
2890:
2883:
2876:
2873:Herelleviridae
2869:
2862:
2859:Drexlerviridae
2855:
2852:Demerecviridae
2848:
2841:
2834:
2826:
2824:
2813:
2810:Caudoviricetes
2802:
2794:
2793:
2790:
2789:
2786:
2785:
2783:
2782:
2775:
2768:
2760:
2758:
2747:
2744:Herviviricetes
2736:
2725:
2722:Heunggongvirae
2714:
2706:
2705:
2702:
2701:
2698:
2697:
2694:
2693:
2690:
2689:
2687:
2686:
2678:
2676:
2668:
2667:
2665:
2664:
2657:
2649:
2647:
2644:Ligamenvirales
2636:
2625:
2614:
2603:
2591:
2590:
2584:
2574:
2568:
2567:
2557:
2555:
2554:
2547:
2540:
2532:
2523:
2522:
2520:
2519:
2507:
2495:
2483:
2470:
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2463:
2458:
2453:
2448:
2443:
2438:
2433:
2428:
2423:
2418:
2413:
2411:Marine viruses
2408:
2403:
2402:
2401:
2391:
2386:
2381:
2379:Antiviral drug
2375:
2373:
2369:
2368:
2366:
2365:
2360:
2355:
2350:
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2334:
2332:
2328:
2327:
2325:
2324:
2319:
2314:
2309:
2304:
2298:
2296:
2292:
2291:
2289:
2288:
2283:
2278:
2273:
2268:
2266:Viral shedding
2263:
2258:
2252:
2250:
2244:
2243:
2236:
2234:
2232:
2231:
2226:
2224:Viral envelope
2221:
2215:
2213:
2209:
2208:
2206:
2205:
2200:
2195:
2190:
2184:
2181:
2180:
2171:
2169:
2168:
2161:
2154:
2146:
2140:
2139:
2134:
2127:
2119:
2106:
2100:
2057:
2023:
2022:External links
2020:
2018:
2017:
1999:
1990:
1941:
1923:
1913:
1854:
1799:
1749:
1700:
1646:
1625:(5): 253–260.
1602:
1593:
1566:(2): 204–208.
1543:
1494:
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1435:
1375:
1350:
1321:(2): 189–199.
1301:
1280:(1): 105–117.
1256:
1238:
1216:
1187:(4): 708–712.
1167:
1130:(5): 896–902.
1110:
1073:(1): 119–141.
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865:vaccinia virus
859:Vaccinia virus
857:Main article:
854:
853:Vaccinia virus
851:
850:
849:
840:
826:
825:
818:
811:
804:
797:
782:
781:
774:
767:
760:
757:Sciuripoxvirus
753:
750:Salmonpoxvirus
746:
739:
732:
725:
718:
715:Mustelpoxvirus
711:
704:
697:
694:Leporipoxvirus
690:
683:
680:Cervidpoxvirus
676:
669:
662:
647:
603:
600:
581:
578:
555:Leporipoxvirus
476:
473:
441:
438:
426:Camelpox virus
417:chordopoxvirus
412:
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6307:
6305:
6304:
6299:
6293:
6292:
6288:
6287:
6285:
6283:
6282:
6281:Blubervirales
6277:
6274:
6272:
6271:
6266:
6263:
6261:
6260:
6255:
6252:
6250:
6249:
6244:
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6238:
6233:
6229:
6225:
6219:
6215:
6185:
6184:
6180:
6178:
6177:
6176:Pseudoviridae
6173:
6171:
6170:
6166:
6164:
6163:
6162:Belpaoviridae
6159:
6158:
6156:
6154:
6153:
6148:
6145:
6143:
6142:
6137:
6134:
6132:
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6022:
6021:
6016:
6006:
6005:
6001:
5999:
5998:
5994:
5992:
5991:
5990:Rhabdoviridae
5987:
5985:
5984:
5983:Pneumoviridae
5980:
5978:
5977:
5973:
5971:
5970:
5966:
5964:
5963:
5962:Mymonaviridae
5959:
5957:
5956:
5952:
5950:
5949:
5945:
5943:
5942:
5938:
5936:
5935:
5931:
5930:
5928:
5926:
5925:
5920:
5914:
5913:
5912:Natareviridae
5909:
5907:
5906:
5902:
5900:
5899:
5895:
5893:
5892:
5888:
5886:
5885:
5881:
5880:
5878:
5876:
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5841:
5839:
5838:
5833:
5830:
5828:
5827:
5822:
5812:
5811:
5807:
5805:
5804:
5803:Amnoonviridae
5800:
5799:
5797:
5795:
5794:
5789:
5786:
5784:
5783:
5778:
5768:
5767:
5766:Wupedeviridae
5763:
5761:
5760:
5756:
5754:
5753:
5752:Phenuiviridae
5749:
5747:
5746:
5745:Phasmaviridae
5742:
5740:
5739:
5735:
5733:
5732:
5728:
5726:
5725:
5721:
5719:
5718:
5714:
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5711:
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5704:
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5698:
5697:
5693:
5691:
5690:
5686:
5685:
5683:
5681:
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5675:
5672:
5670:
5669:
5664:
5654:
5653:
5649:
5648:
5646:
5644:
5643:
5638:
5635:
5633:
5632:
5627:
5624:
5622:
5621:
5616:
5613:
5611:
5610:
5609:Orthornavirae
5605:
5602:
5600:
5599:
5594:
5590:
5586:
5580:
5576:
5554:
5553:
5549:
5547:
5546:
5541:
5538:
5537:
5535:
5531:
5517:
5516:
5512:
5511:
5509:
5507:
5506:
5505:Stellavirales
5501:
5495:
5494:
5490:
5489:
5487:
5485:
5484:
5483:Patatavirales
5479:
5476:
5474:
5473:
5468:
5458:
5457:
5456:Solemoviridae
5453:
5451:
5450:
5446:
5444:
5443:
5439:
5438:
5436:
5434:
5433:
5432:Sobelivirales
5428:
5422:
5421:
5417:
5415:
5414:
5410:
5408:
5407:
5403:
5401:
5400:
5396:
5394:
5393:
5392:Caliciviridae
5389:
5387:
5386:
5382:
5380:
5379:
5375:
5373:
5372:
5368:
5367:
5365:
5363:
5362:
5357:
5351:
5350:
5349:Tobaniviridae
5346:
5344:
5343:
5339:
5337:
5336:
5332:
5330:
5329:
5325:
5323:
5322:
5318:
5316:
5315:
5314:Mononiviridae
5311:
5309:
5308:
5307:Mesoniviridae
5304:
5302:
5301:
5297:
5295:
5294:
5293:Gresnaviridae
5290:
5288:
5287:
5286:Euroniviridae
5283:
5281:
5280:
5279:Coronaviridae
5276:
5274:
5273:
5269:
5267:
5266:
5265:Arteriviridae
5262:
5260:
5259:
5258:Abyssoviridae
5255:
5254:
5252:
5250:
5249:
5244:
5241:
5239:
5238:
5233:
5230:
5228:
5227:
5222:
5208:
5207:
5203:
5202:
5200:
5198:
5197:
5192:
5189:
5187:
5186:
5181:
5171:
5170:
5166:
5165:
5163:
5161:
5160:
5159:Cryppavirales
5155:
5152:
5150:
5149:
5144:
5134:
5133:
5129:
5128:
5126:
5124:
5123:
5118:
5115:
5113:
5112:
5107:
5097:
5096:
5095:Steitzviridae
5092:
5090:
5089:
5085:
5084:
5082:
5080:
5079:
5074:
5068:
5067:
5066:Solspiviridae
5063:
5061:
5060:
5056:
5054:
5053:
5049:
5047:
5046:
5045:Atkinsviridae
5042:
5041:
5039:
5037:
5036:
5031:
5028:
5026:
5025:
5024:Leviviricetes
5020:
5017:
5015:
5014:
5013:Lenarviricota
5009:
4995:
4994:
4993:Tombusviridae
4990:
4988:
4987:
4983:
4981:
4980:
4976:
4975:
4973:
4971:
4970:
4965:
4962:
4960:
4959:
4954:
4944:
4943:
4939:
4937:
4936:
4932:
4931:
4929:
4927:
4926:
4925:Nodamuvirales
4921:
4918:
4916:
4915:
4910:
4900:
4899:
4895:
4894:
4892:
4890:
4889:
4884:
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4856:
4855:
4851:
4849:
4848:
4844:
4842:
4841:
4837:
4835:
4834:
4830:
4829:
4827:
4825:
4824:
4819:
4813:
4812:
4808:
4806:
4805:
4801:
4799:
4798:
4794:
4792:
4791:
4787:
4785:
4784:
4780:
4778:
4777:
4773:
4771:
4770:
4766:
4765:
4763:
4761:
4760:
4755:
4749:
4748:
4747:Matonaviridae
4744:
4742:
4741:
4737:
4735:
4734:
4730:
4728:
4727:
4723:
4722:
4720:
4718:
4717:
4716:Hepelivirales
4712:
4709:
4707:
4706:
4705:Alsuviricetes
4701:
4698:
4696:
4695:
4690:
4687:
4685:
4684:
4683:Orthornavirae
4679:
4676:
4674:
4673:
4668:
4664:
4660:
4654:
4650:
4628:
4627:
4622:
4619:
4617:
4616:
4612:
4610:
4609:
4604:
4601:
4600:
4598:
4594:
4580:
4579:
4575:
4573:
4572:
4568:
4566:
4565:
4561:
4559:
4558:
4554:
4552:
4551:
4550:Amalgaviridae
4547:
4546:
4544:
4542:
4541:
4536:
4533:
4531:
4530:
4525:
4522:
4520:
4519:
4514:
4500:
4499:
4495:
4494:
4492:
4490:
4489:
4484:
4481:
4479:
4478:
4473:
4463:
4462:
4458:
4457:
4455:
4453:
4452:
4447:
4444:
4442:
4441:
4436:
4426:
4425:
4421:
4419:
4418:
4417:Quadriviridae
4414:
4412:
4411:
4407:
4405:
4404:
4403:Chrysoviridae
4400:
4399:
4397:
4395:
4394:
4393:Ghabrivirales
4389:
4386:
4384:
4383:
4378:
4375:
4373:
4372:
4367:
4364:
4362:
4361:
4360:Orthornavirae
4356:
4353:
4351:
4350:
4345:
4341:
4337:
4336:dsRNA viruses
4331:
4327:
4324:
4322:
4318:
4300:
4299:
4295:
4293:
4292:
4288:
4286:
4285:
4284:Anelloviridae
4280:
4277:
4276:
4274:
4270:
4248:
4247:
4243:
4242:
4240:
4238:
4237:
4236:Haloruvirales
4232:
4229:
4227:
4226:
4221:
4218:
4216:
4215:
4210:
4207:
4205:
4204:
4199:
4181:
4180:
4179:Genomoviridae
4176:
4174:
4173:
4172:Geminiviridae
4169:
4168:
4166:
4164:
4163:
4158:
4155:
4153:
4152:
4147:
4137:
4136:
4132:
4131:
4129:
4127:
4126:
4121:
4115:
4114:
4110:
4108:
4107:
4106:Metaxyviridae
4103:
4102:
4100:
4098:
4097:
4092:
4086:
4085:
4081:
4080:
4078:
4076:
4075:
4070:
4064:
4063:
4059:
4058:
4056:
4054:
4053:
4048:
4042:
4041:
4037:
4036:
4034:
4032:
4031:
4026:
4023:
4021:
4020:
4019:Arfiviricetes
4015:
4012:
4010:
4009:
4004:
3990:
3989:
3985:
3984:
3982:
3980:
3979:
3974:
3971:
3969:
3968:
3963:
3953:
3952:
3948:
3947:
3945:
3943:
3942:
3937:
3934:
3932:
3931:
3926:
3923:
3921:
3920:
3919:Cossaviricota
3915:
3912:
3910:
3909:
3904:
3886:
3885:
3881:
3880:
3878:
3876:
3875:
3870:
3867:
3865:
3864:
3859:
3856:
3854:
3853:
3848:
3845:
3843:
3842:
3837:
3819:
3818:
3814:
3812:
3811:
3807:
3805:
3804:
3800:
3799:
3797:
3795:
3794:
3793:Tubulavirales
3789:
3786:
3784:
3783:
3778:
3775:
3773:
3772:
3767:
3764:
3762:
3761:
3756:
3753:
3751:
3750:
3745:
3741:
3737:
3736:ssDNA viruses
3731:
3727:
3709:
3708:
3707:Rhizidiovirus
3704:
3702:
3701:
3696:
3693:
3691:
3690:
3689:Thaspiviridae
3686:
3684:
3683:
3679:
3677:
3676:
3672:
3670:
3669:
3668:Plasmaviridae
3665:
3663:
3662:
3658:
3656:
3655:
3654:Halspiviridae
3651:
3649:
3648:
3644:
3642:
3641:
3637:
3635:
3634:
3630:
3628:
3627:
3623:
3621:
3620:
3616:
3614:
3613:
3608:
3605:
3604:
3602:
3598:
3588:
3587:
3583:
3582:
3580:
3576:
3570:
3569:
3565:
3563:
3562:
3558:
3556:
3555:
3554:Baculoviridae
3551:
3550:
3548:
3546:
3545:
3540:
3537:
3535:
3534:
3529:
3526:
3522:
3500:
3499:
3495:
3493:
3492:
3491:Simuloviridae
3488:
3486:
3485:
3481:
3480:
3478:
3476:
3475:
3470:
3467:
3465:
3464:
3459:
3456:
3454:
3453:
3448:
3445:
3443:
3442:
3441:Helvetiavirae
3437:
3419:
3418:
3414:
3413:
3411:
3409:
3408:
3403:
3397:
3396:
3392:
3391:
3389:
3387:
3386:
3381:
3375:
3374:
3370:
3369:
3367:
3365:
3364:
3363:Kalamavirales
3359:
3353:
3352:
3348:
3347:
3345:
3343:
3342:
3341:Belfryvirales
3337:
3334:
3332:
3331:
3326:
3316:
3315:
3314:Adintoviridae
3311:
3310:
3308:
3306:
3305:
3300:
3297:
3295:
3294:
3289:
3279:
3278:
3277:Lavidaviridae
3274:
3273:
3271:
3269:
3268:
3263:
3260:
3258:
3257:
3252:
3249:
3247:
3246:
3241:
3227:
3226:
3222:
3220:
3219:
3215:
3213:
3212:
3208:
3207:
3205:
3203:
3202:
3197:
3191:
3190:
3186:
3185:
3183:
3181:
3180:
3179:Imitervirales
3175:
3169:
3168:
3164:
3163:
3161:
3159:
3158:
3153:
3150:
3148:
3147:
3146:Megaviricetes
3142:
3132:
3131:
3127:
3126:
3124:
3122:
3121:
3116:
3110:
3109:
3105:
3104:
3102:
3100:
3099:
3094:
3091:
3089:
3088:
3083:
3080:
3078:
3077:
3072:
3069:
3067:
3066:
3061:
3058:
3056:
3055:
3050:
3028:
3027:
3023:
3022:
3020:
3018:
3017:
3012:
3006:
3005:
3001:
3000:
2998:
2996:
2995:
2994:Sepolyvirales
2990:
2987:
2985:
2984:
2979:
2976:
2974:
2973:
2972:Cossaviricota
2968:
2965:
2963:
2962:
2957:
2954:
2952:
2951:
2946:
2924:
2923:
2922:Zobellviridae
2919:
2917:
2916:
2912:
2910:
2909:
2908:Schitoviridae
2905:
2903:
2902:
2898:
2896:
2895:
2891:
2889:
2888:
2884:
2882:
2881:
2877:
2875:
2874:
2870:
2868:
2867:
2866:Guelinviridae
2863:
2861:
2860:
2856:
2854:
2853:
2849:
2847:
2846:
2842:
2840:
2839:
2835:
2833:
2832:
2828:
2827:
2825:
2823:
2822:
2817:
2814:
2812:
2811:
2806:
2803:
2801:
2800:
2795:
2781:
2780:
2776:
2774:
2773:
2772:Herpesviridae
2769:
2767:
2766:
2762:
2761:
2759:
2757:
2756:
2755:Herpesvirales
2751:
2748:
2746:
2745:
2740:
2737:
2735:
2734:
2733:Peploviricota
2729:
2726:
2724:
2723:
2718:
2715:
2713:
2712:
2711:Duplodnaviria
2707:
2685:
2684:
2680:
2679:
2677:
2675:
2674:
2669:
2663:
2662:
2658:
2656:
2655:
2651:
2650:
2648:
2646:
2645:
2640:
2637:
2635:
2634:
2633:Tokiviricetes
2629:
2626:
2624:
2623:
2622:Taleaviricota
2618:
2615:
2613:
2612:
2607:
2604:
2602:
2601:
2596:
2592:
2588:
2587:dsDNA viruses
2582:
2578:
2575:
2573:
2569:
2564:
2560:
2553:
2548:
2546:
2541:
2539:
2534:
2533:
2530:
2518:
2517:
2508:
2506:
2505:
2496:
2494:
2493:
2484:
2482:
2481:
2476:
2472:
2471:
2468:
2462:
2459:
2457:
2454:
2452:
2449:
2447:
2444:
2442:
2439:
2437:
2434:
2432:
2431:Viral disease
2429:
2427:
2424:
2422:
2419:
2417:
2414:
2412:
2409:
2407:
2404:
2400:
2397:
2396:
2395:
2392:
2390:
2387:
2385:
2382:
2380:
2377:
2376:
2374:
2370:
2364:
2361:
2359:
2356:
2354:
2351:
2349:
2346:
2344:
2343:Bacteriophage
2341:
2339:
2336:
2335:
2333:
2329:
2323:
2320:
2318:
2315:
2313:
2310:
2308:
2305:
2303:
2300:
2299:
2297:
2293:
2287:
2284:
2282:
2279:
2277:
2276:Virus latency
2274:
2272:
2269:
2267:
2264:
2262:
2259:
2257:
2254:
2253:
2251:
2249:
2245:
2240:
2230:
2229:Viral protein
2227:
2225:
2222:
2220:
2217:
2216:
2214:
2210:
2204:
2201:
2199:
2196:
2194:
2191:
2189:
2186:
2185:
2182:
2178:
2174:
2167:
2162:
2160:
2155:
2153:
2148:
2147:
2144:
2138:
2135:
2133:
2132:
2128:
2126:
2124:
2120:
2117:
2113:
2110:
2107:
2104:
2101:
2097:
2093:
2088:
2083:
2079:
2075:
2072:(1): 80–122.
2071:
2067:
2063:
2058:
2055:
2053:
2048:
2044:
2040:
2037:
2033:
2032:Orthopoxvirus
2029:
2026:
2025:
2021:
2013:
2009:
2003:
2000:
1994:
1991:
1986:
1982:
1977:
1972:
1968:
1964:
1960:
1956:
1952:
1945:
1942:
1937:
1933:
1927:
1924:
1917:
1914:
1909:
1905:
1900:
1895:
1890:
1885:
1881:
1877:
1873:
1869:
1865:
1858:
1855:
1850:
1846:
1842:
1838:
1834:
1830:
1826:
1822:
1818:
1814:
1810:
1803:
1800:
1794:
1790:
1785:
1780:
1776:
1772:
1768:
1764:
1760:
1753:
1750:
1745:
1741:
1736:
1731:
1727:
1723:
1719:
1715:
1711:
1704:
1701:
1696:
1692:
1687:
1682:
1678:
1674:
1670:
1666:
1662:
1655:
1653:
1651:
1647:
1642:
1638:
1633:
1628:
1624:
1620:
1619:Intervirology
1616:
1609:
1607:
1603:
1597:
1594:
1589:
1585:
1581:
1577:
1573:
1569:
1565:
1561:
1554:
1547:
1544:
1539:
1535:
1530:
1525:
1521:
1517:
1513:
1509:
1505:
1498:
1495:
1490:
1486:
1480:
1472:
1465:
1462:
1450:
1449:Virology Blog
1446:
1439:
1436:
1430:
1426:
1421:
1416:
1411:
1406:
1402:
1398:
1394:
1390:
1386:
1379:
1376:
1364:
1360:
1354:
1351:
1346:
1342:
1338:
1334:
1329:
1324:
1320:
1316:
1312:
1305:
1302:
1297:
1293:
1288:
1283:
1279:
1275:
1271:
1267:
1260:
1257:
1253:
1252:
1247:
1242:
1239:
1227:
1220:
1217:
1212:
1208:
1204:
1200:
1195:
1190:
1186:
1182:
1178:
1171:
1168:
1163:
1159:
1154:
1149:
1145:
1141:
1137:
1133:
1129:
1125:
1121:
1114:
1111:
1106:
1102:
1097:
1092:
1088:
1084:
1080:
1076:
1072:
1068:
1064:
1057:
1054:
1049:
1045:
1041:
1037:
1033:
1029:
1021:
1018:
1014:
1010:
1004:
1001:
997:
992:
987:
986:
977:
974:
961:
957:
951:
948:
935:
931:
925:
923:
919:
906:
900:
897:
891:
887:
884:
882:
879:
878:
874:
872:
868:
866:
860:
852:
847:
846:
845:Centapoxvirus
841:
838:
837:
836:Oryzopoxvirus
832:
831:
830:
824:
823:
819:
817:
816:
812:
810:
809:
805:
803:
802:
798:
796:
795:
791:
790:
789:
788:
787:
780:
779:
775:
773:
772:
768:
766:
765:
761:
759:
758:
754:
752:
751:
747:
745:
744:
740:
738:
737:
733:
731:
730:
729:Oryzopoxvirus
726:
724:
723:
722:Orthopoxvirus
719:
717:
716:
712:
710:
709:
705:
703:
702:
698:
696:
695:
691:
689:
688:
684:
682:
681:
677:
675:
674:
673:Centapoxvirus
670:
668:
667:
666:Capripoxvirus
663:
661:
660:
656:
655:
654:
653:
652:
646:
644:
640:
635:
633:
629:
625:
621:
620:
615:
611:
610:
601:
599:
596:
595:herpesviruses
591:
586:
579:
577:
575:
570:
568:
564:
563:Capripoxvirus
560:
556:
552:
551:Orthopoxvirus
546:
541:
540:Orthopoxvirus
537:
531:
525:
521:
519:
515:
514:
509:
508:
503:
502:
496:
494:
486:
481:
474:
472:
470:
465:
460:
458:
450:
446:
439:
437:
434:
429:
427:
423:
422:Variola virus
418:
410:
408:
406:
402:
398:
394:
390:
385:
381:
377:
370:
366:
359:
354:
352:
349:
345:
341:
336:
334:
329:
328:Edward Jenner
324:
322:
318:
314:
311:
302:
295:
293:
291:
287:
283:
275:
273:
271:
268:
264:
260:
256:
252:
248:
244:
240:
236:
232:
228:
224:
220:
216:
212:
208:
207:Orthopoxvirus
204:
203:
198:
197:
192:
191:
186:
185:
184:Orthopoxvirus
179:
177:
173:
169:
165:
162:
158:
157:
148:
147:
142:
137:
134:
133:
129:
126:
125:
122:
121:
117:
114:
113:
110:
109:
105:
102:
101:
98:
97:
93:
90:
89:
86:
85:
81:
78:
77:
74:
73:
69:
66:
63:
62:
59:
56:
53:
52:
47:
42:
38:
34:
29:
26:
22:
19:
6366:
6311:
6303:Ortervirales
6301:
6289:
6279:
6268:
6257:
6248:Pararnavirae
6246:
6235:
6183:Retroviridae
6181:
6174:
6167:
6160:
6152:Ortervirales
6150:
6139:
6128:
6119:Pararnavirae
6117:
6106:
6039:
6029:
6018:
6004:Xinmoviridae
6002:
5995:
5988:
5981:
5974:
5969:Nyamiviridae
5967:
5960:
5955:Lispiviridae
5953:
5946:
5941:Bornaviridae
5939:
5932:
5922:
5910:
5905:Myriaviridae
5903:
5896:
5889:
5884:Aliusviridae
5882:
5872:
5861:
5845:
5835:
5824:
5808:
5801:
5791:
5780:
5764:
5759:Tospoviridae
5757:
5750:
5743:
5736:
5731:Nairoviridae
5729:
5722:
5715:
5710:Hantaviridae
5708:
5701:
5696:Arenaviridae
5694:
5689:Cruliviridae
5687:
5679:Bunyavirales
5677:
5666:
5650:
5640:
5629:
5618:
5607:
5596:
5550:
5543:
5539:
5515:Astroviridae
5513:
5503:
5491:
5481:
5470:
5454:
5449:Barnaviridae
5447:
5440:
5430:
5418:
5411:
5404:
5397:
5390:
5383:
5378:Marnaviridae
5376:
5369:
5359:
5347:
5340:
5335:Olifoviridae
5333:
5326:
5319:
5312:
5305:
5298:
5291:
5284:
5277:
5270:
5263:
5256:
5246:
5235:
5226:Pisuviricota
5224:
5204:
5196:Ourlivirales
5194:
5185:Miaviricetes
5183:
5167:
5157:
5146:
5132:Narnaviridae
5130:
5120:
5109:
5093:
5088:Blumeviridae
5086:
5078:Timlovirales
5076:
5064:
5059:Fiersviridae
5057:
5050:
5043:
5035:Norzivirales
5033:
5022:
5011:
4991:
4986:Luteoviridae
4984:
4977:
4967:
4956:
4940:
4933:
4923:
4912:
4898:Flaviviridae
4896:
4886:
4875:
4859:
4852:
4845:
4838:
4831:
4821:
4811:Virgaviridae
4809:
4802:
4795:
4788:
4781:
4774:
4769:Bromoviridae
4767:
4757:
4745:
4738:
4731:
4724:
4714:
4703:
4692:
4681:
4670:
4624:
4620:
4613:
4608:Birnaviridae
4606:
4602:
4576:
4569:
4562:
4555:
4548:
4540:Durnavirales
4538:
4527:
4518:Pisuviricota
4516:
4498:Cystoviridae
4496:
4488:Mindivirales
4486:
4475:
4459:
4449:
4438:
4422:
4415:
4408:
4401:
4391:
4380:
4369:
4358:
4347:
4298:Spiraviridae
4296:
4289:
4282:
4278:
4244:
4234:
4223:
4214:Saleviricota
4212:
4201:
4177:
4170:
4160:
4149:
4133:
4125:Recrevirales
4123:
4111:
4104:
4096:Mulpavirales
4094:
4084:Smacoviridae
4082:
4074:Cremevirales
4072:
4062:Circoviridae
4060:
4052:Cirlivirales
4050:
4038:
4030:Baphyvirales
4028:
4017:
4006:
3988:Parvoviridae
3986:
3978:Piccovirales
3976:
3965:
3951:Bidnaviridae
3949:
3939:
3930:Mouviricetes
3928:
3917:
3908:Shotokuvirae
3906:
3884:Microviridae
3882:
3874:Petitvirales
3872:
3861:
3852:Phixviricota
3850:
3839:
3815:
3808:
3801:
3791:
3780:
3769:
3758:
3749:Monodnaviria
3747:
3705:
3700:Dinodnavirus
3698:
3694:
3687:
3680:
3673:
3666:
3661:Ovaliviridae
3659:
3652:
3647:Guttaviridae
3645:
3638:
3631:
3626:Clavaviridae
3624:
3617:
3610:
3606:
3584:
3566:
3559:
3552:
3542:
3531:
3496:
3489:
3482:
3472:
3461:
3450:
3439:
3415:
3405:
3395:Adenoviridae
3393:
3383:
3373:Tectiviridae
3371:
3361:
3351:Turriviridae
3349:
3339:
3328:
3312:
3302:
3291:
3275:
3265:
3254:
3243:
3223:
3218:Iridoviridae
3216:
3209:
3199:
3187:
3177:
3165:
3155:
3144:
3129:
3128:
3120:Chitovirales
3118:
3108:Asfarviridae
3106:
3096:
3085:
3074:
3065:Bamfordvirae
3063:
3054:Varidnaviria
3052:
3024:
3014:
3002:
2992:
2981:
2970:
2961:Shotokuvirae
2959:
2950:Monodnaviria
2948:
2920:
2915:Siphoviridae
2913:
2906:
2899:
2892:
2885:
2878:
2871:
2864:
2857:
2850:
2845:Chaseviridae
2843:
2836:
2829:
2821:Caudovirales
2819:
2808:
2797:
2777:
2770:
2763:
2753:
2742:
2731:
2720:
2709:
2681:
2673:Primavirales
2671:
2659:
2652:
2642:
2631:
2620:
2609:
2598:
2514:
2502:
2490:
2478:
2394:Viral vector
2389:Helper virus
2353:Human virome
2338:Animal virus
2317:Reassortment
2193:Introduction
2173:Microbiology
2130:
2125:: Poxviridae
2122:
2069:
2065:
2051:
2036:Parapoxvirus
2035:
2031:
2011:
2002:
1993:
1958:
1954:
1944:
1935:
1926:
1916:
1871:
1867:
1857:
1816:
1812:
1802:
1766:
1762:
1752:
1717:
1713:
1703:
1671:(1): 50–59.
1668:
1664:
1622:
1618:
1596:
1563:
1559:
1546:
1511:
1507:
1497:
1464:
1452:. Retrieved
1448:
1438:
1392:
1388:
1378:
1366:. Retrieved
1362:
1353:
1318:
1314:
1304:
1277:
1273:
1259:
1251:Cells Alive!
1249:
1241:
1229:. Retrieved
1219:
1184:
1180:
1170:
1127:
1123:
1113:
1070:
1066:
1056:
1031:
1027:
1020:
1003:
994:
984:
976:
964:. Retrieved
959:
950:
938:. Retrieved
933:
909:. Retrieved
905:"Viral Zone"
899:
869:
862:
843:
834:
828:
820:
813:
806:
799:
792:
784:
783:
778:Yatapoxvirus
776:
769:
762:
755:
748:
741:
736:Parapoxvirus
734:
727:
720:
713:
706:
699:
692:
685:
678:
671:
664:
657:
649:
648:
642:
638:
636:
631:
627:
617:
607:
605:
587:
583:
573:
571:
566:
562:
559:Yatapoxvirus
558:
554:
550:
539:
535:
534:. The genus
526:
522:
517:
511:
507:Asfarviridae
505:
499:
497:
490:
484:
461:
454:
448:
430:
414:
404:
401:rhinoviruses
375:
374:
368:
355:Microbiology
337:
325:
307:
281:
279:
254:
243:Yatapoxvirus
242:
235:pseudocowpox
227:Parapoxvirus
226:
206:
200:
196:Yatapoxvirus
194:
190:Parapoxvirus
188:
182:
180:
155:
154:
153:
144:
139:Subfamilies
131:
130:
120:Chitovirales
119:
118:
107:
95:
84:Bamfordvirae
83:
72:Varidnaviria
71:
64:
54:(unranked):
24:
18:
6445:iNaturalist
6391:Wikispecies
6169:Metaviridae
5948:Filoviridae
5934:Artoviridae
5847:Aspiviridae
5724:Mypoviridae
5703:Fimoviridae
5493:Potyviridae
5406:Secoviridae
5399:Iflaviridae
5342:Roniviridae
5248:Nidovirales
5169:Mitoviridae
5052:Duinviridae
4969:Tolivirales
4935:Nodaviridae
4861:Tymoviridae
4823:Tymovirales
4804:Togaviridae
4797:Mayoviridae
4790:Kitaviridae
4740:Hepeviridae
4733:Benyviridae
4564:Hypoviridae
4424:Totiviridae
4113:Nanoviridae
3941:Polivirales
3841:Sangervirae
3586:Nimaviridae
3568:Nudiviridae
3544:Lefavirales
3407:Vinavirales
3385:Rowavirales
3211:Ascoviridae
3189:Mimiviridae
3157:Algavirales
3098:Asfuvirales
2887:Podoviridae
2799:Uroviricota
2661:Rudiviridae
2611:Zilligvirae
2516:WikiProject
2384:Giant virus
2363:Plant virus
2281:Lytic cycle
2256:Viral entry
1315:Virus Genes
1274:J Gen Virol
1231:26 February
886:Water warts
764:Suipoxvirus
659:Avipoxvirus
614:vertebrates
567:Suipoxvirus
536:Avipoxvirus
501:Rudiviridae
440:Replication
267:herpesvirus
168:Vertebrates
6512:Poxviruses
6506:Categories
6397:Poxviridae
6367:Poxviridae
6041:Yueviridae
5997:Sunviridae
5891:Chuviridae
5652:Qinviridae
5533:Unassigned
4596:Unassigned
4461:Reoviridae
4451:Reovirales
4272:Unassigned
4203:Trapavirae
3803:Inoviridae
3600:Unassigned
3578:Unassigned
3524:Unassigned
3130:Poxviridae
2880:Myoviridae
2461:Virosphere
2436:Viral load
2426:Satellites
2212:Components
2052:Poxviridae
1013:0816045631
892:References
518:Poxviridae
493:adenovirus
485:Poxviridae
449:Poxviridae
433:GC-content
405:Poxviridae
376:Poxviridae
369:Poxviridae
282:Poxviridae
263:chickenpox
172:arthropods
156:Poxviridae
132:Poxviridae
25:Poxviridae
6237:Riboviria
6108:Riboviria
5642:Muvirales
5598:Riboviria
4672:Riboviria
4349:Riboviria
3760:Loebvirae
2600:Adnaviria
2559:Baltimore
2421:Oncovirus
2358:Mycovirus
2348:Virophage
2271:Viroplasm
2123:Viralzone
2049:in their
1508:Structure
1266:Smith, GL
1203:1998-4138
1144:1525-0024
1087:2327-056X
590:Taterapox
475:Evolution
380:enveloped
360:Structure
276:Etymology
79:Kingdom:
6376:Wikidata
5540:Families
4603:Families
4279:Families
3607:Families
2492:Category
2295:Genetics
2203:Virology
2112:Archived
2043:smallpox
1985:33754985
1908:17901212
1849:39823899
1841:16873609
1793:28930029
1744:26136578
1695:19833230
1641:21228539
1588:20018642
1580:15046572
1538:21742267
1479:cite web
1429:20231474
1345:21779605
1337:21369827
1296:14718625
1268:(2004).
1211:32930107
1162:20160706
1105:25839047
1048:10367824
907:. ExPASy
875:See also
602:Taxonomy
580:Smallpox
407:virion.
317:Americas
313:Ramses V
290:smallpox
215:vaccinia
211:smallpox
176:smallpox
146:See text
127:Family:
91:Phylum:
6382:Q948421
2504:Commons
2331:By host
2188:History
2096:1851533
1976:8007330
1899:2000395
1876:Bibcode
1821:Bibcode
1813:Science
1784:5621524
1735:4542343
1714:J Virol
1686:2818276
1529:3136756
1454:4 March
1420:2851855
1397:Bibcode
1368:26 June
1153:2890119
1096:4380149
966:16 July
911:15 June
639:virinae
624:insects
622:infect
612:infect
469:nucleus
344:Atlanta
310:pharaoh
296:History
249:virus,
247:tanapox
241:virus;
233:virus,
225:virus;
221:virus,
217:virus,
164:viruses
115:Order:
103:Class:
6489:600031
6463:113827
6450:484646
4621:Genera
3695:Genera
2480:Portal
2456:Virome
2219:Capsid
2094:
2087:372802
2084:
2039:Genera
1983:
1973:
1906:
1896:
1847:
1839:
1791:
1781:
1742:
1732:
1693:
1683:
1639:
1586:
1578:
1536:
1526:
1427:
1417:
1343:
1335:
1294:
1209:
1201:
1160:
1150:
1142:
1103:
1093:
1085:
1046:
1015:p.337.
1011:
411:Genome
397:genome
371:virion
321:Aztecs
219:cowpox
6484:WoRMS
6476:10240
6458:IRMNG
6411:627B2
6222:VII:
4334:III:
2372:Other
2177:Virus
1921:Virol
1845:S2CID
1584:S2CID
1556:(PDF)
1341:S2CID
940:9 May
643:virus
65:Realm
58:Virus
6471:NCBI
6437:1937
6432:GBIF
6424:8050
6093:VI:
4657:IV:
3734:II:
2131:ICTV
2092:PMID
2034:and
1981:PMID
1904:PMID
1837:PMID
1789:PMID
1740:PMID
1691:PMID
1637:PMID
1576:PMID
1534:PMID
1489:link
1485:link
1456:2014
1425:PMID
1370:2022
1333:PMID
1292:PMID
1233:2005
1207:PMID
1199:ISSN
1158:PMID
1140:ISSN
1101:PMID
1083:ISSN
1044:PMID
1009:ISBN
968:2006
942:2020
913:2015
565:and
431:The
424:and
223:Mpox
170:and
6419:EoL
6406:CoL
5583:V:
4321:RNA
2585:I:
2572:DNA
2082:PMC
2074:doi
2030:of
1971:PMC
1963:doi
1894:PMC
1884:doi
1872:104
1829:doi
1817:313
1779:PMC
1771:doi
1730:PMC
1722:doi
1681:PMC
1673:doi
1627:doi
1568:doi
1524:PMC
1516:doi
1415:PMC
1405:doi
1393:107
1323:doi
1282:doi
1248:at
1189:doi
1148:PMC
1132:doi
1091:PMC
1075:doi
1036:doi
1032:281
545:0.3
530:0.5
520:).
464:DNA
257::
245::
231:orf
209::
161:DNA
6508::
6486::
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6378::
6080:RT
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4281::
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3609::
2175::
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2080:.
2070:55
2068:.
2064:.
2010:.
1979:.
1969:.
1959:27
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1870:.
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