81:
37:
56:
376:, with a wingspan of 1.24–1.41 m (4 ft 1 in – 4 ft 8 in). It weighs 400–500 g (0.88–1.10 lb). Its forearm is approximately 140 mm (5.5 in) long. The body of the bat is covered in long hairs, making the body seem almost woolly. The bat is reddish brown and has a yellowish white
459:
The Ryukyu flying fox is native to Taiwan, Japan and the
Philippines. In Japan it is found on the ĹŚsumi Islands, Tokara Islands, Okinawa Islands, Miyako Islands, Yaeyama Islands and DaitĹŤ Islands. In the Philippines it is present in Batan, Dalupiri and Fuga. Its habitat is forests where it roosts
468:
The largest population of these bats is probably on the
Philippines and is thought to be stable. In Japan there are estimated to be well over five thousand individuals but in Taiwan, there has been a large reduction in bat numbers. This species faces a number of threats. Some populations in the
473:. In Japan, habitat loss is the main threat but some individuals get entangled in nets placed to protect citrus crops and others are electrocuted by power-lines. Overall, most populations have been in decline though this seemed to have levelled off to some extent by 2008 when the
445:
202:
751:
Nakamoto, Atsushi; Kinjo, Kazumitsu; Izawa, Masako (2007). "Food habits of Orii's flying-fox, Pteropus dasymallus inopinatus, in relation to food availability in an urban area of
Okinawa-jima Island, the Ryukyu Archipelago, Japan".
396:, consuming the fruits of at least 53 plant species; the flowers of 20 plant species; the leaves of 18 plant species; and the bark of one plant species. It has also been observed consuming eight different species of
451:
450:
447:
446:
452:
449:
3764:
3941:
497:
to launch an investigation in 2012. In two villages surveyed in 2013, it was estimated that flying foxes cause 19 million yen ($ 175 thousand USD) in damages to
3829:
779:
Funakoshi, K; Watanabe, H; Kunisaki, T (1993). "Feeding ecology of the northern Ryukyu fruit bat, Pteropus dasymallus dasymallus, in a warm-temperate region".
636:"The role of Orii's flying-fox (Pteropus dasymallus inopinatus) as a pollinator and a seed disperser on Okinawa-jima Island, the Ryukyu Archipelago, Japan"
3896:
590:
Monographies de
Mammalogie, ou description de quelques genres de Mammifères, dont les espèces ont été observées dans les différens musées de l'Europe
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681:"Public awareness and perceptual factors in the conservation of elusive species: The case of the endangered Ryukyu flying fox"
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1769:
1760:
1072:
723:
Andrew T. Smith; Yan Xie; Robert S. Hoffmann; Darrin Lunde; John MacKinnon; Don E. Wilson; W. Chris
Wozencraft, eds. (2010).
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1911:
1336:
1039:
940:
817:
302:
3916:
3503:
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1970:
428:
species, usually solitary roosting in trees during the day and foraging at night. The Ryukyu flying fox enhances
80:
2840:
2590:
2279:
2270:
2180:
1811:
1453:
1435:
1187:
1163:
2795:
2500:
2464:
1402:
1259:
493:
In
Temminck's initial description, he wrote that it "devastates" orchards. Its depredation on orchards caused
3690:
765:
3577:
2687:
2419:
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2228:
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2020:
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1920:
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2011:
1961:
1600:
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45:
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2002:
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75:
60:
20:
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36:
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2154:
1820:
1802:
1590:
1501:
1477:
1354:
1139:
502:
250:
802:
699:
329: 'woolly'; Temminck described its fur as long and woolly. The five subspecies are:
3558:
2509:
2365:
870:
792:
429:
201:
679:
Vincenot, Christian Ernest; Collazo, Anja Maria; Wallmo, Kristy; Koyama, Lina (2015).
380:. Its ears are small and pointed, and are difficult to see beneath its thick fur. Its
3890:
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3333:
3133:
3091:
1369:
1178:
1054:
541:
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65:
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2873:
2048:
1985:
1902:
1861:
1154:
1100:
892:
325:
270:
234:
662:
297:. Temminck acquired the specimens used for his description from Dutch businessman
315:
3873:
3808:
3751:
3684:
3468:
3155:
2919:
1516:
1468:
1319:
708:
469:
Philippines are hunted for consumption and this bat is considered a delicacy on
254:
2932:
2243:
2121:
1944:
1581:
1233:
876:
654:
501:
crops annually. Many
Japanese farmers believe that the Ryukyu flying fox is a
405:
3675:
592:. Vol. 1. Paris: G. Dufour et E. d'Ocagne. pp. 180–181, pl. XX–XVI.
3521:
2982:
2219:
1638:
1557:
1492:
1417:
1113:
425:
393:
92:
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3669:
3531:
2303:
2195:
2061:
1662:
1393:
1209:
864:
858:
852:
846:
381:
364:
The subspecies are based on populations that occur on different islands.
152:
112:
3743:
3259:
834:
506:
258:
246:
230:
142:
3756:
3001:
498:
415:
397:
262:
242:
122:
102:
3646:
586:"Cinquième Monographie. Vues générales sur l'ordre des cheiroptères"
404:
tree is an important source of food year-round. It is an important
3868:
443:
433:
266:
238:
727:(illustrated ed.). Princeton University Press. p. 332.
474:
377:
274:
3730:
3650:
3130:
2916:
2045:
1845:
1622:
1097:
889:
806:
526:
524:
522:
132:
634:
Nakamoto, Atsushi; Kinjo, Kazumitsu; Izawa, Masako (2008).
436:
up to 1,833 m (1.139 mi) from the parent trees.
766:
10.3161/1733-5329(2007)9[237:FHOOFP]2.0.CO;2
414:, an evergreen tree. It also pollinates a species of
460:
during the day in trees, singly or in small groups.
3659:
3608:
3566:
3557:
3529:
3520:
3466:
3433:
3409:
3400:
3372:
3331:
3307:
3257:
3153:
3144:
2980:
2930:
2871:
2301:
2241:
2217:
2193:
2152:
2119:
2059:
1983:
1942:
1900:
1859:
1800:
1660:
1636:
1579:
1555:
1514:
1490:
1466:
1415:
1391:
1367:
1317:
1231:
1207:
1176:
1152:
1111:
1052:
903:
372:The Ryukyu flying fox is slightly smaller than the
432:, as seeds from digested fruits are deposited as
674:
672:
579:
577:
575:
573:
629:
627:
625:
556:10.2305/IUCN.UK.2017-2.RLTS.T18722A22080614.en
818:
8:
293:as a new species in 1825 by Dutch zoologist
3647:
3563:
3526:
3406:
3150:
3141:
3127:
2927:
2913:
2056:
2042:
1856:
1842:
1633:
1619:
1108:
1094:
900:
886:
825:
811:
803:
746:
744:
200:
54:
35:
26:
707:
610:. Integrated Taxonomic Information System
554:
3495:East African little collared fruit bat (
3384:Veldkamp's dwarf epauletted fruit bat (
518:
3942:Taxa named by Coenraad Jacob Temminck
3351:Hayman's dwarf epauletted fruit bat (
3209:Lesser Angolan epauletted fruit bat (
7:
3360:Peters' dwarf epauletted fruit bat (
3277:Buettikofer's epauletted fruit bat (
1779:New Britain naked-backed fruit bat (
1287:Nusatenggara short-nosed fruit bat (
923:Broad-striped tube-nosed fruit bat (
604:"Pteropus dasymallus Temminck, 1825"
3227:East African epauletted fruit bat (
1698:Philippine naked-backed fruit bat (
1689:Beaufort's naked-backed fruit bat (
1680:Andersen's naked-backed fruit bat (
542:IUCN Red List of Threatened Species
2715:Little golden-mantled flying fox (
1734:Solomon's naked-backed fruit bat (
1707:Halmahera naked-backed fruit bat (
1305:Indonesian short-nosed fruit bat (
959:Round-eared tube-nosed fruit bat (
793:10.1111/j.1469-7998.1993.tb02684.x
14:
3295:Franquet's epauletted fruit bat (
3245:Wahlberg's epauletted fruit bat (
2892:Sulawesi stripe-faced fruit bat (
2089:Giant golden-crowned flying fox (
1788:Greenish naked-backed fruit bat (
1752:Moluccan naked-backed fruit bat (
1725:Sulawesi naked-backed fruit bat (
1022:Philippine tube-nosed fruit bat (
481:" category and placed it in the "
3897:IUCN Red List vulnerable species
3218:Ethiopian epauletted fruit bat (
2883:Mindoro stripe-faced fruit bat (
1770:Western naked-backed fruit bat (
1761:Panniet naked-backed fruit bat (
1073:Steadfast tube-nosed fruit bat (
273:and by hunting for food and the
261:are subtropical or tropical dry
79:
3191:Peters's epauletted fruit bat (
3182:Ansell's epauletted fruit bat (
2438:New Ireland masked flying fox (
2289:New Georgian monkey-faced Bat (
2253:Bougainville monkey-faced Bat (
2140:Small white-winged flying fox (
1743:Lesser naked-backed fruit bat (
1296:Greater short-nosed fruit bat (
950:Mountain tube-nosed fruit bat (
725:A Guide to the Mammals of China
688:Global Ecology and Conservation
3200:Gambian epauletted fruit bat (
3173:Angolan epauletted fruit bat (
2262:Guadalcanal monkey-faced Bat (
1251:Lesser short-nosed fruit bat (
1004:Demonic tube-nosed fruit bat (
995:Malaita tube-nosed fruit bat (
977:Keast's tube-nosed fruit bat (
1:
3932:Near threatened biota of Asia
3486:São Tomé collared fruit bat (
1716:Biak naked-backed fruit bat (
1064:Lesser tube-nosed fruit bat (
986:Island tube-nosed fruit bat (
968:Dragon tube-nosed fruit bat (
932:Common tube-nosed fruit bat (
3476:(Little collared fruit bats)
3236:Minor epauletted fruit bat (
2787:Philippine gray flying fox (
2384:Bismarck masked flying fox (
1445:Ratanaworabhan's fruit bat (
1040:Umboi tube-nosed fruit bat (
265:and subtropical or tropical
3504:Little collared fruit bat (
2162:(Straw-coloured fruit bats)
1971:New Caledonia blossom bat (
1379:Fischer's pygmy fruit bat (
709:10.1016/j.gecco.2015.02.005
477:removed this bat from the "
3958:
3922:Mammals of the Philippines
2841:New Caledonia flying fox (
2591:Black-bearded flying fox (
2280:Montane monkey-faced Bat (
2271:Greater monkey-faced Bat (
2181:Straw-coloured fruit bat (
1812:Sulawesi harpy fruit bat (
1454:White-collared fruit bat (
1436:Javan tailless fruit bat (
1346:Rickart's dyak fruit bat (
1188:Spotted-winged fruit bat (
1164:Mindanao pygmy fruit bat (
505:that should be managed by
319: 'hairy' and
18:
3937:Mammals described in 1825
3578:Short-palated fruit bat (
3140:
3126:
2926:
2912:
2688:Geelvink Bay flying fox (
2519:Livingstone's fruit bat (
2420:Large Samoan flying fox (
2348:Small Samoan flying fox (
2229:Small-toothed fruit bat (
2205:Fijian monkey-faced Bat (
2131:White-winged flying fox (
2055:
2041:
2021:Moss-forest blossom bat (
1921:Black-bellied fruit bat (
1912:Fardoulis's blossom bat (
1879:Long-tongued nectar bat (
1869:(Long-tongued fruit bats)
1855:
1841:
1632:
1618:
1534:Greater musky fruit bat (
1337:Brooks's dyak fruit bat (
1107:
1093:
941:Pallas's tube-nosed bat (
899:
885:
841:
833:Extant species of family
655:10.1007/s11284-008-0516-y
384:are dark brown in color.
208:
199:
181:
174:
76:Scientific classification
74:
52:
43:
34:
29:
3541:D'Anchieta's fruit bat (
3052:Leschenault's rousette (
2697:Grey-headed flying fox (
2600:Black-eared flying fox (
2483:Ontong Java flying fox (
2375:Ashy-headed flying fox (
1952:(Long-tailed fruit bats)
1888:Long-tongued fruit bat (
1670:(Bare-backed fruit bats)
1543:Lesser musky fruit bat (
1219:Black-capped fruit bat (
1031:Eastern tube-nosed Bat (
440:Distribution and habitat
209:Ryukyu flying fox range
3927:Near threatened animals
3587:Campo-Ma’an fruit bat (
2796:Temminck's flying fox (
2769:Little red flying fox (
2751:Madagascan flying fox (
2501:Andersen's flying fox (
2465:Gilliard's flying fox (
2411:Spectacled flying fox (
2012:Halmahera blossom bat (
1962:Long-tailed fruit bat (
1403:Salim Ali's fruit bat (
1260:Horsfield's fruit bat (
1013:Lesser tube-nosed bat (
913:(tube-nosed fruit bats)
584:Temminck, C.J. (1825).
531:Vincenot, C.E. (2017).
295:Coenraad Jacob Temminck
3912:Bats of Southeast Asia
3341:(Dwarf epauleted bats)
3163:(Epauleted fruit bats)
3097:Long-haired rousette (
3079:Bare-backed rousette (
2778:Seychelles fruit bat (
2742:Rodrigues flying fox (
2627:Mauritian flying fox (
2555:Big-eared flying fox (
2321:Admiralty flying fox (
2172:Madagascan fruit bat (
1930:Woodford's fruit bat (
1567:Blanford's fruit bat (
1241:(Dog-faced fruit bats)
1198:Balionycteris seimundi
489:Relationship to humans
456:
345:Taiwanese fruit bat -
285:Taxonomy and etymology
269:. It is threatened by
3106:Madagascan rousette (
3025:Geoffroy's rousette (
2990:(rousette fruit bats)
2968:Philippine dawn bat (
2814:Vanikoro flying fox (
2724:Solomons flying fox (
2609:Caroline flying fox (
2393:Moluccan flying fox (
2071:Sulawesi flying fox (
754:Acta Chiropterologica
549:: e.T18722A22080614.
455:
309:" is likely from
3620:Zenker's fruit bat (
3286:Dobson's fruit bat (
2950:Greater nectar bat (
2805:Insular flying fox (
2733:Rennell flying fox (
2537:Okinawa flying fox (
2447:Nicobar flying fox (
2357:Vanuatu flying fox (
2330:Aldabra flying fox (
2003:Common blossom bat (
1648:Bulmer's fruit bat (
1427:Tailless fruit bat (
1269:Peters's fruit bat (
357:Yaeyama fruit bat -
3691:Pteropus dasymallus
3661:Pteropus dasymallus
3596:Pohle's fruit bat (
3445:Azagnyi fruit bat (
3319:Hammer-headed Bat (
3108:R. madagascariensis
3043:Egyptian rousette (
3034:Sulawesi rousette (
2823:Kosrae flying fox (
2679:Masked flying fox (
2654:Ornate flying fox (
2636:Temotu flying fox (
2582:Indian flying fox (
2573:Mariana fruit bat (
2564:Lesser flying fox (
2546:Lyle's flying fox (
2528:Lombok flying fox (
2429:Ryukyu flying fox (
2402:Makira flying fox (
2142:D. microleucopterus
2098:Palawan fruit bat (
2080:Talaud flying fox (
700:2015GEcoC...3..526V
643:Ecological Research
535:Pteropus dasymallus
408:of a subspecies of
388:Biology and ecology
351:Orii's fruit bat -
226:Pteropus dasymallus
185:Pteropus dasymallus
46:Conservation status
3421:Angolan rousette (
3061:Linduan Rousette (
3027:R. amplexicaudatus
3007:Manado fruit bat (
2859:Dwarf flying fox (
2850:Pemba flying fox (
2832:Large flying fox (
2760:Samoa flying fox (
2706:Bonin flying fox (
2670:Pelew flying fox (
2618:Great flying fox (
2492:Small flying fox (
2456:Banks flying fox (
2339:Black flying fox (
2107:Sunda flying fox (
1524:(Musky fruit bats)
1327:(Dayak fruit bats)
1278:Minute fruit bat (
1131:Borneo fruit bat (
1121:(Pygmy fruit bats)
781:Journal of Zoology
495:Okinawa Prefecture
457:
339:Erabu fruit bat -
333:Daito fruit bat -
229:) is a species of
167:P. dasymallus
30:Ryukyu flying fox
21:Okinawa flying fox
3884:
3883:
3856:Open Tree of Life
3653:Taxon identifiers
3644:
3643:
3640:
3639:
3636:
3635:
3632:
3631:
3589:C. campomaanensis
3553:
3552:
3516:
3515:
3479:
3396:
3395:
3344:
3270:
3166:
3122:
3121:
3118:
3117:
3070:Comoro rousette (
2993:
2959:Cave nectar bat (
2943:
2940:(Dawn fruit bats)
2908:
2907:
2904:
2903:
2645:Ceram fruit bat (
2474:Gray flying fox (
2413:P. conspicillatus
2314:
2165:
2037:
2036:
2033:
2032:
1996:
1955:
1872:
1837:
1836:
1833:
1832:
1821:Harpy fruit bat (
1673:
1626:Harpyionycterinae
1614:
1613:
1610:
1609:
1591:Swift fruit bat (
1527:
1502:Dusky fruit bat (
1480:O. cartilagonodus
1478:Luzon fruit bat (
1355:Dayak fruit bat (
1330:
1307:C. titthaecheilus
1244:
1221:C. melanocephalus
1140:Pygmy fruit bat (
1124:
1089:
1088:
1085:
1084:
916:
453:
421:Mucuna macrocarpa
374:Indian flying fox
299:Jan Cock Blomhoff
237:. It is found in
217:Ryukyu flying fox
213:
212:
69:
3949:
3917:Mammals of Japan
3877:
3876:
3864:
3863:
3851:
3850:
3838:
3837:
3825:
3824:
3812:
3811:
3799:
3798:
3786:
3785:
3773:
3772:
3760:
3759:
3747:
3746:
3734:
3733:
3721:
3720:
3708:
3707:
3695:
3694:
3693:
3680:
3679:
3678:
3648:
3564:
3527:
3488:M. brachycephala
3478:
3477:
3473:
3454:Woermann's bat (
3407:
3343:
3342:
3338:
3269:
3268:
3267:(Epauleted bats)
3264:
3165:
3164:
3160:
3151:
3142:
3128:
2992:
2991:
2987:
2942:
2941:
2937:
2928:
2914:
2780:P. seychellensis
2699:P. poliocephalus
2620:P. neohibernicus
2510:Kei flying fox (
2395:P. chrysoproctus
2366:Aru flying fox (
2323:P. admiralitatum
2313:
2312:
2308:
2164:
2163:
2159:
2057:
2043:
1995:
1994:
1990:
1973:N. neocaledonica
1954:
1953:
1949:
1871:
1870:
1866:
1857:
1843:
1672:
1671:
1667:
1634:
1620:
1526:
1525:
1521:
1329:
1328:
1324:
1243:
1242:
1238:
1123:
1122:
1118:
1109:
1095:
915:
914:
910:
901:
887:
827:
820:
813:
804:
797:
796:
776:
770:
769:
748:
739:
738:
720:
714:
713:
711:
685:
676:
667:
666:
640:
631:
620:
619:
617:
615:
600:
594:
593:
581:
568:
567:
565:
563:
558:
528:
454:
411:Schima wallichii
382:flight membranes
353:P. d. inopinatus
341:P. d. dasymallus
335:P. d. daitoensis
277:classify it as "
221:Ryukyu fruit bat
204:
187:
84:
83:
63:
58:
57:
39:
27:
3957:
3956:
3952:
3951:
3950:
3948:
3947:
3946:
3887:
3886:
3885:
3880:
3872:
3867:
3859:
3854:
3846:
3843:Observation.org
3841:
3833:
3828:
3820:
3815:
3807:
3802:
3794:
3789:
3781:
3776:
3768:
3763:
3755:
3750:
3742:
3737:
3729:
3724:
3716:
3711:
3703:
3698:
3689:
3688:
3683:
3674:
3673:
3668:
3655:
3645:
3628:
3604:
3549:
3512:
3475:
3474:
3472:
3462:
3429:
3392:
3368:
3340:
3339:
3337:
3327:
3303:
3279:E. buettikoferi
3266:
3265:
3263:
3253:
3162:
3161:
3159:
3136:
3114:
3054:R. leschenaulti
2989:
2988:
2986:
2976:
2939:
2938:
2936:
2922:
2900:
2867:
2816:P. tuberculatus
2638:P. nitendiensis
2521:P. livingstonii
2467:P. gilliardorum
2310:
2309:
2307:
2297:
2237:
2213:
2189:
2161:
2160:
2158:
2148:
2115:
2051:
2029:
1992:
1991:
1989:
1979:
1951:
1950:
1948:
1938:
1896:
1868:
1867:
1865:
1851:
1829:
1803:Harpyionycteris
1796:
1763:D. pannietensis
1669:
1668:
1666:
1656:
1628:
1606:
1575:
1551:
1523:
1522:
1520:
1510:
1486:
1462:
1411:
1387:
1363:
1326:
1325:
1323:
1313:
1289:C. nusatenggara
1240:
1239:
1237:
1227:
1203:
1172:
1166:A. paucidentata
1148:
1120:
1119:
1117:
1103:
1081:
1048:
912:
911:
909:
895:
881:
837:
831:
801:
800:
778:
777:
773:
750:
749:
742:
735:
722:
721:
717:
683:
678:
677:
670:
638:
633:
632:
623:
613:
611:
602:
601:
597:
583:
582:
571:
561:
559:
530:
529:
520:
515:
491:
466:
444:
442:
390:
370:
359:P. d. yayeyamae
287:
251:Babuyan Islands
195:
189:
183:
170:
78:
70:
59:
55:
48:
23:
17:
12:
11:
5:
3955:
3953:
3945:
3944:
3939:
3934:
3929:
3924:
3919:
3914:
3909:
3904:
3899:
3889:
3888:
3882:
3881:
3879:
3878:
3865:
3852:
3839:
3826:
3813:
3800:
3787:
3774:
3761:
3748:
3735:
3722:
3709:
3696:
3681:
3665:
3663:
3657:
3656:
3651:
3642:
3641:
3638:
3637:
3634:
3633:
3630:
3629:
3627:
3626:
3616:
3614:
3606:
3605:
3603:
3602:
3593:
3584:
3574:
3572:
3561:
3559:Scotonycterini
3555:
3554:
3551:
3550:
3548:
3547:
3537:
3535:
3524:
3518:
3517:
3514:
3513:
3511:
3510:
3501:
3492:
3482:
3480:
3464:
3463:
3461:
3460:
3451:
3441:
3439:
3431:
3430:
3428:
3427:
3417:
3415:
3404:
3398:
3397:
3394:
3393:
3391:
3390:
3380:
3378:
3370:
3369:
3367:
3366:
3357:
3353:M. intermedius
3347:
3345:
3329:
3328:
3326:
3325:
3315:
3313:
3305:
3304:
3302:
3301:
3292:
3283:
3273:
3271:
3255:
3254:
3252:
3251:
3242:
3233:
3224:
3215:
3206:
3197:
3188:
3179:
3169:
3167:
3148:
3138:
3137:
3131:
3124:
3123:
3120:
3119:
3116:
3115:
3113:
3112:
3103:
3086:
3085:
3076:
3067:
3058:
3049:
3045:R. aegyptiacus
3040:
3031:
3014:
3013:
2996:
2994:
2978:
2977:
2975:
2974:
2965:
2956:
2946:
2944:
2924:
2923:
2917:
2910:
2909:
2906:
2905:
2902:
2901:
2899:
2898:
2889:
2885:S. mindorensis
2879:
2877:
2869:
2868:
2866:
2865:
2856:
2847:
2838:
2829:
2820:
2811:
2802:
2793:
2784:
2775:
2766:
2757:
2748:
2744:P. rodricensis
2739:
2730:
2721:
2712:
2703:
2694:
2685:
2676:
2667:
2660:
2651:
2642:
2633:
2624:
2615:
2606:
2597:
2593:P. melanopogon
2588:
2579:
2570:
2561:
2552:
2543:
2539:P. loochoensis
2534:
2530:P. lombocensis
2525:
2516:
2507:
2503:P. intermedius
2498:
2494:P. hypomelanus
2489:
2480:
2471:
2462:
2453:
2444:
2435:
2426:
2417:
2408:
2399:
2390:
2386:P. capistratus
2381:
2372:
2363:
2354:
2345:
2336:
2332:P. aldabrensis
2327:
2317:
2315:
2311:(flying foxes)
2299:
2298:
2296:
2295:
2286:
2277:
2268:
2259:
2249:
2247:
2239:
2238:
2236:
2235:
2225:
2223:
2215:
2214:
2212:
2211:
2201:
2199:
2191:
2190:
2188:
2187:
2178:
2168:
2166:
2150:
2149:
2147:
2146:
2137:
2133:D. leucopterus
2127:
2125:
2117:
2116:
2114:
2113:
2104:
2095:
2086:
2077:
2067:
2065:
2053:
2052:
2046:
2039:
2038:
2035:
2034:
2031:
2030:
2028:
2027:
2018:
2009:
1999:
1997:
1993:(blossom bats)
1981:
1980:
1978:
1977:
1968:
1958:
1956:
1940:
1939:
1937:
1936:
1927:
1918:
1908:
1906:
1898:
1897:
1895:
1894:
1885:
1875:
1873:
1853:
1852:
1849:Macroglossinae
1846:
1839:
1838:
1835:
1834:
1831:
1830:
1828:
1827:
1818:
1808:
1806:
1798:
1797:
1795:
1794:
1785:
1776:
1767:
1758:
1754:D. moluccensis
1749:
1740:
1731:
1722:
1713:
1704:
1695:
1686:
1676:
1674:
1658:
1657:
1655:
1654:
1644:
1642:
1630:
1629:
1623:
1616:
1615:
1612:
1611:
1608:
1607:
1605:
1604:
1597:
1587:
1585:
1577:
1576:
1574:
1573:
1563:
1561:
1553:
1552:
1550:
1549:
1540:
1530:
1528:
1512:
1511:
1509:
1508:
1498:
1496:
1488:
1487:
1485:
1484:
1474:
1472:
1464:
1463:
1461:
1460:
1451:
1442:
1433:
1423:
1421:
1413:
1412:
1410:
1409:
1399:
1397:
1389:
1388:
1386:
1385:
1375:
1373:
1365:
1364:
1362:
1361:
1352:
1343:
1333:
1331:
1315:
1314:
1312:
1311:
1302:
1293:
1284:
1275:
1271:C. luzoniensis
1266:
1262:C. horsfieldii
1257:
1247:
1245:
1229:
1228:
1226:
1225:
1215:
1213:
1205:
1204:
1202:
1201:
1194:
1184:
1182:
1174:
1173:
1171:
1170:
1160:
1158:
1150:
1149:
1147:
1146:
1137:
1127:
1125:
1105:
1104:
1098:
1091:
1090:
1087:
1086:
1083:
1082:
1080:
1079:
1070:
1060:
1058:
1050:
1049:
1047:
1046:
1037:
1028:
1019:
1010:
1001:
997:N. malaitensis
992:
983:
974:
965:
956:
947:
938:
929:
919:
917:
897:
896:
890:
883:
882:
880:
879:
873:
871:Laurasiatheria
867:
861:
855:
849:
842:
839:
838:
832:
830:
829:
822:
815:
807:
799:
798:
771:
740:
733:
715:
668:
649:(2): 405–414.
621:
595:
569:
517:
516:
514:
511:
490:
487:
465:
462:
441:
438:
430:seed dispersal
402:Chinese banyan
389:
386:
369:
366:
362:
361:
355:
349:
347:P. d. formosus
343:
337:
286:
283:
257:. Its natural
233:in the family
211:
210:
206:
205:
197:
196:
190:
179:
178:
172:
171:
164:
162:
158:
157:
150:
146:
145:
140:
136:
135:
130:
126:
125:
120:
116:
115:
110:
106:
105:
100:
96:
95:
90:
86:
85:
72:
71:
53:
50:
49:
44:
41:
40:
32:
31:
16:Species of bat
15:
13:
10:
9:
6:
4:
3:
2:
3954:
3943:
3940:
3938:
3935:
3933:
3930:
3928:
3925:
3923:
3920:
3918:
3915:
3913:
3910:
3908:
3905:
3903:
3900:
3898:
3895:
3894:
3892:
3875:
3870:
3866:
3862:
3857:
3853:
3849:
3844:
3840:
3836:
3831:
3827:
3823:
3818:
3814:
3810:
3805:
3801:
3797:
3792:
3788:
3784:
3779:
3775:
3771:
3766:
3762:
3758:
3753:
3749:
3745:
3740:
3736:
3732:
3727:
3723:
3719:
3714:
3710:
3706:
3701:
3697:
3692:
3686:
3682:
3677:
3671:
3667:
3666:
3664:
3662:
3658:
3654:
3649:
3625:
3623:
3618:
3617:
3615:
3613:
3612:
3611:Scotonycteris
3607:
3601:
3599:
3594:
3592:
3590:
3585:
3583:
3581:
3576:
3575:
3573:
3571:
3570:
3565:
3562:
3560:
3556:
3546:
3544:
3539:
3538:
3536:
3534:
3533:
3528:
3525:
3523:
3519:
3509:
3507:
3502:
3500:
3498:
3493:
3491:
3489:
3484:
3483:
3481:
3471:
3470:
3465:
3459:
3457:
3452:
3450:
3448:
3443:
3442:
3440:
3438:
3437:
3436:Megaloglossus
3432:
3426:
3424:
3423:L. angolensis
3419:
3418:
3416:
3414:
3413:
3412:Lissonycteris
3408:
3405:
3403:
3399:
3389:
3387:
3386:N. veldkampii
3382:
3381:
3379:
3377:
3376:
3371:
3365:
3363:
3358:
3356:
3354:
3349:
3348:
3346:
3336:
3335:
3334:Micropteropus
3330:
3324:
3322:
3321:H. monstrosus
3317:
3316:
3314:
3312:
3311:
3306:
3300:
3298:
3293:
3291:
3289:
3284:
3282:
3280:
3275:
3274:
3272:
3262:
3261:
3256:
3250:
3248:
3243:
3241:
3239:
3234:
3232:
3230:
3225:
3223:
3221:
3216:
3214:
3212:
3207:
3205:
3203:
3198:
3196:
3194:
3189:
3187:
3185:
3180:
3178:
3176:
3175:E. angolensis
3171:
3170:
3168:
3158:
3157:
3152:
3149:
3147:
3143:
3139:
3135:
3134:Epomophorinae
3129:
3125:
3111:
3109:
3104:
3102:
3100:
3094:
3093:
3092:Stenonycteris
3088:
3087:
3084:
3082:
3081:R. spinalatus
3077:
3075:
3073:
3072:R. obliviosus
3068:
3066:
3064:
3063:R. linduensis
3059:
3057:
3055:
3050:
3048:
3046:
3041:
3039:
3037:
3036:R. celebensis
3032:
3030:
3028:
3022:
3021:
3016:
3015:
3012:
3010:
3004:
3003:
2998:
2997:
2995:
2985:
2984:
2979:
2973:
2971:
2966:
2964:
2962:
2957:
2955:
2953:
2948:
2947:
2945:
2935:
2934:
2929:
2925:
2921:
2915:
2911:
2897:
2895:
2890:
2888:
2886:
2881:
2880:
2878:
2876:
2875:
2870:
2864:
2862:
2857:
2855:
2853:
2852:P. voeltzkowi
2848:
2846:
2844:
2839:
2837:
2835:
2830:
2828:
2826:
2821:
2819:
2817:
2812:
2810:
2808:
2803:
2801:
2799:
2798:P. temminckii
2794:
2792:
2790:
2785:
2783:
2781:
2776:
2774:
2772:
2771:P. scapulatus
2767:
2765:
2763:
2758:
2756:
2754:
2749:
2747:
2745:
2740:
2738:
2736:
2731:
2729:
2727:
2722:
2720:
2718:
2713:
2711:
2709:
2704:
2702:
2700:
2695:
2693:
2691:
2686:
2684:
2682:
2681:P. personatus
2677:
2675:
2673:
2672:P. pelewensis
2668:
2666:
2665:
2661:
2659:
2657:
2652:
2650:
2648:
2643:
2641:
2639:
2634:
2632:
2630:
2625:
2623:
2621:
2616:
2614:
2612:
2611:P. molossinus
2607:
2605:
2603:
2598:
2596:
2594:
2589:
2587:
2585:
2580:
2578:
2576:
2571:
2569:
2567:
2562:
2560:
2558:
2553:
2551:
2549:
2544:
2542:
2540:
2535:
2533:
2531:
2526:
2524:
2522:
2517:
2515:
2513:
2508:
2506:
2504:
2499:
2497:
2495:
2490:
2488:
2486:
2481:
2479:
2477:
2472:
2470:
2468:
2463:
2461:
2459:
2454:
2452:
2450:
2445:
2443:
2441:
2436:
2434:
2432:
2431:P. dasymallus
2427:
2425:
2423:
2418:
2416:
2414:
2409:
2407:
2405:
2400:
2398:
2396:
2391:
2389:
2387:
2382:
2380:
2378:
2373:
2371:
2369:
2364:
2362:
2360:
2355:
2353:
2351:
2346:
2344:
2342:
2337:
2335:
2333:
2328:
2326:
2324:
2319:
2318:
2316:
2306:
2305:
2300:
2294:
2292:
2287:
2285:
2283:
2278:
2276:
2274:
2269:
2267:
2265:
2260:
2258:
2256:
2251:
2250:
2248:
2246:
2245:
2240:
2234:
2232:
2227:
2226:
2224:
2222:
2221:
2216:
2210:
2208:
2203:
2202:
2200:
2198:
2197:
2192:
2186:
2184:
2179:
2177:
2175:
2170:
2169:
2167:
2157:
2156:
2151:
2145:
2143:
2138:
2136:
2134:
2129:
2128:
2126:
2124:
2123:
2118:
2112:
2110:
2105:
2103:
2101:
2096:
2094:
2092:
2087:
2085:
2083:
2078:
2076:
2074:
2073:A. celebensis
2069:
2068:
2066:
2064:
2063:
2058:
2054:
2050:
2044:
2040:
2026:
2024:
2019:
2017:
2015:
2010:
2008:
2006:
2001:
2000:
1998:
1988:
1987:
1982:
1976:
1974:
1969:
1967:
1965:
1964:N. macdonaldi
1960:
1959:
1957:
1947:
1946:
1941:
1935:
1933:
1928:
1926:
1924:
1919:
1917:
1915:
1914:M. fardoulisi
1910:
1909:
1907:
1905:
1904:
1899:
1893:
1891:
1886:
1884:
1882:
1877:
1876:
1874:
1864:
1863:
1858:
1854:
1850:
1844:
1840:
1826:
1824:
1823:H. whiteheadi
1819:
1817:
1815:
1814:H. celebensis
1810:
1809:
1807:
1805:
1804:
1799:
1793:
1791:
1786:
1784:
1782:
1781:D. praedatrix
1777:
1775:
1773:
1768:
1766:
1764:
1759:
1757:
1755:
1750:
1748:
1746:
1741:
1739:
1737:
1732:
1730:
1728:
1723:
1721:
1719:
1714:
1712:
1710:
1705:
1703:
1701:
1696:
1694:
1692:
1687:
1685:
1683:
1678:
1677:
1675:
1665:
1664:
1659:
1653:
1651:
1646:
1645:
1643:
1641:
1640:
1635:
1631:
1627:
1621:
1617:
1603:
1602:
1601:T. suhaniahae
1598:
1596:
1594:
1593:T. nigrescens
1589:
1588:
1586:
1584:
1583:
1578:
1572:
1570:
1565:
1564:
1562:
1560:
1559:
1554:
1548:
1546:
1541:
1539:
1537:
1532:
1531:
1529:
1519:
1518:
1513:
1507:
1505:
1500:
1499:
1497:
1495:
1494:
1489:
1483:
1481:
1476:
1475:
1473:
1471:
1470:
1465:
1459:
1457:
1452:
1450:
1448:
1443:
1441:
1439:
1434:
1432:
1430:
1425:
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1422:
1420:
1419:
1414:
1408:
1406:
1401:
1400:
1398:
1396:
1395:
1390:
1384:
1382:
1377:
1376:
1374:
1372:
1371:
1370:Haplonycteris
1366:
1360:
1358:
1353:
1351:
1349:
1344:
1342:
1340:
1335:
1334:
1332:
1322:
1321:
1316:
1310:
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1285:
1283:
1281:
1276:
1274:
1272:
1267:
1265:
1263:
1258:
1256:
1254:
1253:C. brachyotis
1249:
1248:
1246:
1236:
1235:
1230:
1224:
1222:
1217:
1216:
1214:
1212:
1211:
1206:
1200:
1199:
1195:
1193:
1191:
1186:
1185:
1183:
1181:
1180:
1179:Balionycteris
1175:
1169:
1167:
1162:
1161:
1159:
1157:
1156:
1151:
1145:
1143:
1138:
1136:
1134:
1129:
1128:
1126:
1116:
1115:
1110:
1106:
1102:
1096:
1092:
1078:
1076:
1071:
1069:
1067:
1062:
1061:
1059:
1057:
1056:
1055:Paranyctimene
1051:
1045:
1043:
1038:
1036:
1034:
1029:
1027:
1025:
1020:
1018:
1016:
1011:
1009:
1007:
1002:
1000:
998:
993:
991:
989:
984:
982:
980:
975:
973:
971:
970:N. draconilla
966:
964:
962:
957:
955:
953:
948:
946:
944:
943:N. cephalotes
939:
937:
935:
934:N. albiventer
930:
928:
926:
921:
920:
918:
908:
907:
902:
898:
894:
888:
884:
878:
874:
872:
868:
866:
862:
860:
856:
854:
850:
848:
844:
843:
840:
836:
828:
823:
821:
816:
814:
809:
808:
805:
794:
790:
786:
782:
775:
772:
767:
763:
759:
755:
747:
745:
741:
736:
734:9781400834112
730:
726:
719:
716:
710:
705:
701:
697:
693:
689:
682:
675:
673:
669:
664:
660:
656:
652:
648:
644:
637:
630:
628:
626:
622:
609:
605:
599:
596:
591:
587:
580:
578:
576:
574:
570:
557:
552:
548:
544:
543:
538:
536:
527:
525:
523:
519:
512:
510:
508:
504:
500:
496:
488:
486:
484:
480:
476:
472:
471:Babuyan Claro
463:
461:
439:
437:
435:
431:
427:
423:
422:
417:
416:climbing vine
413:
412:
407:
403:
399:
395:
392:It is mostly
387:
385:
383:
379:
375:
367:
365:
360:
356:
354:
350:
348:
344:
342:
338:
336:
332:
331:
330:
328:
327:
322:
321:Ancient Greek
318:
317:
312:
311:Ancient Greek
308:
304:
300:
296:
292:
284:
282:
280:
276:
272:
268:
264:
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240:
236:
232:
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227:
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207:
203:
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186:
180:
177:
176:Binomial name
173:
169:
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160:
159:
156:
155:
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148:
147:
144:
141:
138:
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108:
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91:
88:
87:
82:
77:
73:
67:
62:
51:
47:
42:
38:
33:
28:
25:
22:
3907:Bats of Asia
3660:
3621:
3609:
3597:
3588:
3579:
3569:Casinycteris
3567:
3543:P. anchietae
3542:
3530:
3505:
3496:
3487:
3467:
3456:M. woermanni
3455:
3446:
3434:
3422:
3410:
3402:Myonycterini
3385:
3375:Nanonycteris
3373:
3361:
3352:
3332:
3320:
3310:Hypsignathus
3308:
3297:E. franqueti
3296:
3287:
3278:
3258:
3247:E. wahlbergi
3246:
3237:
3228:
3219:
3210:
3202:E. gambianus
3201:
3193:E. crypturus
3192:
3183:
3174:
3154:
3146:Epomophorini
3107:
3098:
3089:
3080:
3071:
3062:
3053:
3044:
3035:
3026:
3017:
3008:
2999:
2981:
2969:
2960:
2951:
2931:
2893:
2884:
2874:Styloctenium
2872:
2861:P. woodfordi
2860:
2851:
2842:
2833:
2824:
2815:
2806:
2797:
2789:P. speciosus
2788:
2779:
2770:
2762:P. samoensis
2761:
2752:
2743:
2734:
2725:
2716:
2708:P. pselaphon
2707:
2698:
2689:
2680:
2671:
2664:P. pelagicus
2662:
2655:
2646:
2637:
2628:
2619:
2610:
2602:P. melanotus
2601:
2592:
2583:
2575:P. mariannus
2574:
2566:P. mahaganus
2565:
2556:
2547:
2538:
2529:
2520:
2511:
2502:
2493:
2484:
2475:
2466:
2457:
2448:
2439:
2430:
2428:
2421:
2412:
2403:
2394:
2385:
2376:
2367:
2359:P. anetianus
2358:
2350:P. allenorum
2349:
2340:
2331:
2322:
2302:
2290:
2281:
2273:P. flanneryi
2272:
2263:
2254:
2242:
2230:
2218:
2207:M. acrodonta
2206:
2194:
2182:
2174:E. dupreanum
2173:
2153:
2141:
2132:
2120:
2108:
2099:
2090:
2081:
2072:
2060:
2049:Pteropodinae
2022:
2014:S. carolinae
2013:
2005:S. australis
2004:
1986:Syconycteris
1984:
1972:
1963:
1943:
1932:M. woodfordi
1931:
1922:
1913:
1903:Melonycteris
1901:
1889:
1880:
1862:Macroglossus
1860:
1822:
1813:
1801:
1789:
1780:
1771:
1762:
1753:
1744:
1735:
1726:
1717:
1709:D. crenulata
1708:
1699:
1691:D. beauforti
1690:
1682:D. anderseni
1681:
1661:
1649:
1637:
1599:
1592:
1580:
1569:S. blanfordi
1568:
1556:
1544:
1535:
1515:
1503:
1491:
1479:
1467:
1455:
1446:
1437:
1429:M. ecaudatus
1428:
1416:
1405:L. salimalii
1404:
1392:
1380:
1368:
1357:D. spadiceus
1356:
1347:
1338:
1318:
1306:
1297:
1288:
1279:
1270:
1261:
1252:
1232:
1220:
1208:
1196:
1189:
1177:
1165:
1155:Alionycteris
1153:
1141:
1132:
1112:
1101:Cynopterinae
1074:
1065:
1053:
1042:N. vizcaccia
1041:
1033:N. robinsoni
1032:
1023:
1014:
1005:
996:
987:
978:
969:
960:
951:
942:
933:
924:
904:
893:Nyctimeninae
869:Superorder:
863:Infraclass:
835:Pteropodidae
784:
780:
774:
757:
753:
724:
718:
691:
687:
646:
642:
612:. Retrieved
607:
598:
589:
560:. Retrieved
546:
540:
534:
492:
485:" category.
467:
458:
419:
409:
391:
371:
363:
358:
352:
346:
340:
334:
324:
314:
306:
303:species name
288:
271:habitat loss
235:Pteropodidae
225:
224:
220:
216:
214:
184:
182:
166:
165:
153:
143:Pteropodidae
24:
3752:iNaturalist
3685:Wikispecies
3598:C. ophiodon
3580:C. argynnis
3506:M. torquata
3469:Myonycteris
3362:M. pusillus
3220:E. labiatus
3156:Epomophorus
2920:Rousettinae
2894:S. wallacei
2834:P. vampyrus
2807:P. tonganus
2735:P. rennelli
2647:P. ocularis
2557:P. macrotis
2512:P. keyensis
2485:P. howensis
2458:P. fundatus
2449:P. faunulus
2404:P. cognatus
2377:P. caniceps
2368:P. aruensis
2122:Desmalopex
2109:A. mackloti
2100:A. leucotis
1923:M. melanops
1890:M. sobrinus
1700:D. chapmani
1650:A. bulmerae
1517:Ptenochirus
1469:Otopteropus
1456:M. wetmorei
1447:M. niphanae
1438:M. kusnotoi
1381:H. fischeri
1348:D. rickarti
1320:Dyacopterus
1190:B. maculata
1133:A. aequalis
961:N. cyclotis
760:: 237–249.
694:: 526–540.
614:22 February
608:ITIS Report
562:16 November
394:frugivorous
368:Description
255:Philippines
3891:Categories
3622:S. zenkeri
3497:M. relicta
3447:M. azagnyi
3288:E. dobsoni
3229:E. minimus
3211:E. grandis
3184:E. anselli
3132:Subfamily
3099:R. lanosus
2970:E. robusta
2961:E. spelaea
2933:Eonycteris
2918:Subfamily
2843:P. vetulus
2825:P. ualanus
2726:P. rayneri
2717:P. pumilus
2656:P. ornatus
2476:P. griseus
2440:P. ennisae
2282:P. pulchra
2244:Pteralopex
2091:A. jubatus
2082:A. humilis
2047:Subfamily
1945:Notopteris
1881:M. minimus
1847:Subfamily
1790:D. viridis
1772:D. peronii
1736:D. inermis
1727:D. exoleta
1624:Subfamily
1582:Thoopterus
1339:D. brooksi
1280:C. minutus
1234:Cynopterus
1099:Subfamily
1015:N. minutus
1006:N. masalai
952:N. certans
891:Subfamily
877:Chiroptera
787:(2): 221.
513:References
483:Vulnerable
479:Endangered
424:. It is a
406:pollinator
307:dasymallus
279:Vulnerable
245:, and the
133:Chiroptera
61:Vulnerable
19:See also:
3522:Plerotini
3090:Subgenus
3020:Rousettus
3018:Subgenus
3009:R. bidens
3000:Subgenus
2983:Rousettus
2690:P. pohlei
2584:P. medius
2341:P. alecto
2264:P. atrata
2255:P. anceps
2231:N. frosti
2220:Neopteryx
2183:E. helvum
2023:S. hobbit
1718:D. emersa
1639:Aproteles
1558:Sphaerias
1536:P. jagori
1504:P. lucasi
1493:Penthetor
1418:Megaerops
1298:C. sphinx
1142:A. alecto
1114:Aethalops
1066:P. raptor
1024:N. rabori
979:N. keasti
906:Nyctimene
845:Kingdom:
426:nocturnal
291:described
161:Species:
99:Kingdom:
93:Eukaryota
3902:Pteropus
3869:Species+
3822:13800273
3770:10464713
3670:Wikidata
3532:Plerotes
3238:E. minor
2952:E. major
2753:P. rufus
2629:P. niger
2548:P. lylei
2304:Pteropus
2196:Mirimiri
2062:Acerodon
1745:D. minor
1663:Dobsonia
1545:P. minor
1394:Latidens
1210:Chironax
1075:P. tenax
988:N. major
925:N. aello
865:Eutheria
859:Mammalia
853:Chordata
851:Phylum:
847:Animalia
259:habitats
192:Temminck
154:Pteropus
139:Family:
123:Mammalia
113:Chordata
109:Phylum:
103:Animalia
89:Domain:
66:IUCN 3.1
3809:1004461
3744:5218670
3676:Q575969
3260:Epomops
2422:P. coxi
2291:P. taki
2155:Eidolon
875:Order:
857:Class:
696:Bibcode
507:culling
289:It was
263:forests
253:of the
247:Batanes
231:megabat
149:Genus:
129:Order:
119:Class:
64: (
3861:608040
3835:126282
3783:631626
3731:327263
3002:Boneia
731:
663:901864
661:
499:citrus
464:Status
400:. The
398:insect
326:mallĂłs
267:swamps
243:Taiwan
194:, 1825
3848:86991
3796:18722
3765:IRMNG
3757:40887
3718:4Q2Z8
3705:30429
684:(PDF)
659:S2CID
639:(PDF)
434:guano
323:
316:dasĂşs
313:
239:Japan
3874:3837
3830:NCBI
3791:IUCN
3778:ITIS
3739:GBIF
3700:BOLD
729:ISBN
616:2018
564:2021
547:2017
503:pest
475:IUCN
378:nape
301:Its
275:IUCN
249:and
215:The
3817:MSW
3804:MDD
3726:EoL
3713:CoL
789:doi
785:230
762:doi
704:doi
651:doi
551:doi
281:".
219:or
3893::
3871::
3858::
3845::
3832::
3819::
3806::
3793::
3780::
3767::
3754::
3741::
3728::
3715::
3702::
3687::
3672::
3095::
3023::
3005::
783:.
756:.
743:^
702:.
690:.
686:.
671:^
657:.
647:24
645:.
641:.
624:^
606:.
588:.
572:^
545:.
539:.
521:^
509:.
418:,
241:,
3624:)
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3499:)
3490:)
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