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1248:, probably serving instead for the recognition of individual birds. However, the intensity of the red on the bills is regarded an indicator of the animal's quality and social dominance. Red-billed quelea males mate with one female only within one breeding cycle. There are usually three eggs in each clutch (though the full range is one to five) of approximately 18 mm (0.71 in) long and 13 mm (0.51 in) in diameter. The eggs are light bluish or greenish in colour, sometimes with some dark spots. Some clutches contain six eggs, but large clutches may be the result of other females dumping an egg in a stranger's nest.
48:
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31:
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The temporarily wet areas do not form a single zone that periodically moves back and forth across the entirety of Sub-Saharan Africa, but rather consist of five or six regions, within which the wet areas "move" or "jump". Red-billed quelea populations thus migrate between the temporarily wet areas within each of these five to six geographical regions. Each of the subspecies, as distinguished by different male breeding plumage, is confined to one or more of these geographical regions.
97:
1459:
was calculated to consume roughly 13 t (29,000 lb) of insects and 1,000 t (2,200,000 lb) of grass seeds during its breeding cycle. At sunrise they form flocks that co-operate to find food. After a successful search, they settle to feed. In the heat of the day, they rest in the shade, preferably near water, and preen. Birds seem to prefer drinking at least twice a day. In the evening, they once again fly off in search of food.
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72:
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efficiently. It is regarded as the most numerous undomesticated bird on earth, with the total post-breeding population sometimes peaking at an estimated 1.5 billion individuals. It feeds in huge flocks of millions of individuals, with birds that run out of food at the rear flying over the entire group to a fresh feeding zone at the front, creating an image of a rolling cloud. The
557:. It ranges across central and southern Africa, where it has been recorded from southwestern Gabon, southern Congo, Angola (except the northeast and arid coastal southwest), southern Democratic Republic of Congo and the mouth of the Congo River, Zambia, Malawi and western Mozambique across to Namibia (except the coastal desert) and central, southern and eastern South Africa.
1233:
of a hanging forked twig, and from there bridging the gaps in the circle his beak can reach, having one foot on each of the branchlets, using the same footholds and the same orientation throughout the building process. Two parallel stems of reeds or sugar cane can also be used to attach the nest from. They use both their bills and feet in adding the initial knots needed.
1237:
looks like a small oval or globular ball of grass, around 18 cm (7 in) high and 16 cm (6 in) wide, with a 2.5 cm (1 in) wide entrance high up one side, sheltered by a shallow awning. About six to seven hundred fresh, green grass strips are used for each nest. This species may nest several times per year when conditions are favourable.
1708:
962:. The legs are orangey in colour. The bill is bright raspberry red. Outside the breeding season, the male lacks bright colours; it has a grey-brown head with dark streaks, whitish chin and throat, and a faint light stripe above the eyes. At this time, the bill becomes pink or dull red and the legs turn flesh-coloured.
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from the air on breeding colonies and roosts. In
Botswana and Zimbabwe, spraying was also executed from ground vehicles and manually. Kenya and South Africa regularly used fire-bombs. Attempts during the 1950s and '60s to eradicate populations, at least regionally, failed. Consequently, management is
1455:, but entire maize kernels are too big for them to swallow. A single bird may eat about 15 g (0.53 oz) in seeds each day. As much as half of the diet of nestlings consists of insects, such as grasshoppers, ants, beetles, bugs, caterpillars, flies and termites, as well as snails and spiders.
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As soon as the ring is finished the male displays, trying to attract a female, after which the nest may be completed in two days. The nest chamber is created in front of the ring. The entrance may be constructed after the egg laying has started, while the male works from the outside. A finished nest
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Red-billed queleas migrate seasonally over long distances in anticipation of the availability of their main natural food source, seeds of annual grasses. The presence of these grass seeds is the result of the beginning of rains weeks earlier, and the rainfall varies in a seasonal geographic pattern.
1047:
Flying flocks make a distinct sound due to the many wing beats. After arriving at the roost or nest site, birds keep moving around and make a lot of noise for about half an hour before settling in. Both males and females call. The male sings in short bursts, starting with some chatter, followed by a
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of the eggs during the day, but the female alone does so during the cool night, and feeds during the day when air temperatures are high enough to sustain the development of the embryo. The breeding cycle of the red-billed quelea is one of the shortest known in any bird. Incubation takes nine or ten
1232:
in
Zimbabwe one colony was 20 km (12 mi) long and 1 km (0.6 mi) wide. In southern Africa, suitable branches are stripped of leaves a few days in advance of the onset of nest construction. The male starts the nest by creating a ring of grass by twining strips around both branches
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in 2000. Occasionally, it can be found as high as 3,000 m (9,800 ft) above sea-level, but mostly resides below 1,500 m (4,900 ft). It visits agricultural areas, where it feeds on cereal crops, although it is thought to prefer seeds of wild annual grasses. It needs to drink daily
996:
the mask does not extend far above the bill, and the underparts may have a pink wash. There is much variability within subspecies, and some birds cannot be ascribed to a subspecies based on outward appearance alone. Because of interbreeding, specimens intermediate between subspecies may occur where
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by hobbyists. It thrives if kept in large and high cages, with space to fly to minimise the risk of obesity. A sociable bird, the red-billed quelea tolerates mixed-species aviaries. Keeping many individuals mimics its natural occurrence in large flocks. This species withstands frosts, but requires
1458:
Insects are generally eaten during the breeding season, though winged termites are eaten at other times. Breeding females consume snail-shell fragments and calcareous grit, presumably to enable egg-shell formation. One colony in
Namibia, of an estimated five million adults and five million chicks,
380:
or detonating fire-bombs in the enormous colonies during the night. Extensive control measures have been largely unsuccessful in limiting the quelea population. When food runs out, the species migrates to locations of recent rainfall and plentiful grass seed; hence it exploits its food source very
1305:
Flocks of red-billed queleas usually feed on the ground, with birds in the rear constantly leap-frogging those in the front to exploit the next strip of fallen seeds. This behaviour creates the impression of a rolling cloud, and enables efficient exploitation of the available food. The birds also
1113:
is thought to consist of two sub-populations. One moves from
Central Tanzania to southern Somalia, to return to breed in Tanzania in February and March, followed by successive migrations to breed ever further north, the season's last usually occurring in central Kenya during May. The second group
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valley, for instance, the rainy season has already passed and the grass has produced new seeds. After about six weeks, the birds migrate northwards to find a suitable breeding area, nurture a generation, and then repeat this sequence moving further north. Some populations may also move northwards
953:
and higher parts of the throat. Occasionally males have a white mask. The mask is surrounded by a variable band of yellow, rusty, pink or purple. White masks are sometimes bordered by black. This colouring may only reach the lower throat or extend along the belly, with the rest of the underparts
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The red-billed quelea is a small sparrow-like bird, approximately 12 cm (4.7 in) long and weighing 15–26 g (0.53–0.92 oz), with a heavy, cone-shaped bill, which is red (in females outside the breeding season and males) or orange to yellow (females during the breeding season).
972:
on the top of the head and the shoulders. The eyes open during the fourth day, at the same time as the first feathers appear. Older nestlings have a horn-coloured bill with a hint of lavender, though it turns orange-purple before the post-juvenile moult. Young birds change feathers two to three
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each year, representing a market value of approximately US$ 37,500–75,000. Between 13 June and 21 August 1994 alone, 1.2 million queleas were caught. Birds were taken from roosts in the trees during the moonless period each night. The feathers were plucked and the carcasses fried the following
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and a reddish bill. Breeding females attain a yellowish bill. Breeding males have a black (or rarely white) facial mask, surrounded by a purplish, pinkish, rusty or yellowish wash on the head and breast. The species avoids forests, deserts and colder areas such as those at high altitude and in
1125:. In November, part of the population migrates to the northwest to northwestern Angola, while the remainder migrates to the southeast to southern Mozambique and eastern South-Africa, but no proof has been found that these migration cohorts are genetically or morphologically divergent.
1988:
Ornithologia, sive, Synopsis methodica sistens avium divisionem in ordines, sectiones, genera, species, ipsarumque varietates : cum brevi & accurata cujusque speciei descriptione, citationibus auctorum de iis tractantium, nominibus eis ab ipsis impositis, nominibusque
539:, is native to west and central Africa, where it has been recorded from Mauritania, western and northern Senegal, Gambia, central Mali, Burkina Faso, southwestern and southern Niger, northern Nigeria, Cameroon, south-central Chad and northern Central African Republic.
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species—below 1,000 m (3,300 ft) elevation. While foraging for food, they may fly 50–65 km (31–40 mi) each day and return to the roosting or nesting site in the evening. Small groups of red-billed queleas often mix with different weaver birds
1137:
on feeding on seeds of annual grass species, which may be ripe, or still green, but have not germinated yet. Since the availability of these seeds varies with time and space, occurring in particular weeks after the local off-set of rains, queleas migrate as a
575:. It is found in eastern Africa where it occurs in southern Sudan, eastern South Sudan, Ethiopia and Eritrea south to the northeastern parts of the Democratic Republic of Congo, Uganda, Kenya, central and eastern Tanzania and northwestern and southern Somalia.
1100:
In
Nigeria, the nominate subspecies generally travels 300–600 km (190–370 mi) southwards during the start of the rains in the north during June and July, when the grass seed germinates, and is no longer eaten by the queleas. When they reach the
1202:
The red-billed quelea needs 300–800 mm (12–31 in) of precipitation to breed, with nest building usually commencing four to nine weeks after the onset of the rains. Nests are usually built in stands of thorny trees such as umbrella thorn acacia
710:(IOC). Other names in English include black-faced dioch, cardinal, common dioch, Latham's weaver-bird, pink-billed weaver, quelea finch, quelea weaver, red-billed dioch, red-billed weaver, Russ' weaver, South-African dioch, Sudan dioch and Uganda dioch.
1742:
2467:
De Silva, Thilina N.; Peterson, A. Townsend; Bates, John M.; Fernandoa, Sumudu W.; Girard, Matthew G. (2017). "Phylogenetic relationships of weaverbirds (Aves: Ploceidae): A first robust phylogeny based on mitochondrial and nuclear markers".
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in 1960 on the basis of more greyish nonbreeding plumage of populations of wetter habitats of northeastern South Africa, Eswatini and southern
Mozambique. However, further analysis indicated no clear distinction in plumage between it and
1004:
could be mistaken for the red-billed quelea in non-breeding plumage, since both are sparrow-like birds with conical red-coloured bills, but the whydah has a whitish brow between a black stripe through the eye and a black stripe above.
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The governments of
Botswana, Ethiopia, Kenya, South Africa, Sudan, Tanzania, and Zimbabwe have regularly made attempts to lessen quelea populations. The most common method to kill members of problematic flocks was by spraying the
437:. He incorrectly mentioned that it originated in India, probably because ships from the East Indies picked up birds when visiting the African coast during their return voyage to Europe. It is likely that he had seen a draft of
1723:
1517:
sometimes attack drinking queleas, and an individual in
Ethiopia hit birds out of the vegetation on the bank into the water with its tail, subsequently eating them. Queleas drinking at a waterhole were grabbed from below by
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shelter from rain and wind. Affixing hanging branches, such as hawthorn, in the cage facilitates nesting. Adults are typically given a diet of tropical seeds enriched with grass seeds, augmented by living insects such as
1306:
take seeds from the grass ears directly. They prefer grains of 1–2 mm (0.04–0.08 in) in size. Red-billed queleas feed mainly on grass seeds, which includes a large number of annual species from the genera
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tribe use triangular hand-held nets, which are both selective and efficient. Each team of six trappers caught about twenty thousand birds each night. An estimated five to ten million queleas are trapped near
954:
light brown or whitish with some dark stripes. The upperparts have light and dark brown longitudinal stripes, particularly at midlength, and are paler on the rump. The tail and upper wing are dark brown. The
1587:
fishermen cast nets over groups of birds. The impact of hunting on the quelea population (about 200 million individuals in the Lake Chad Basin) is deemed insignificant. Woven traps made from star grass
1133:
The red-billed quelea is regarded as the most numerous undomesticated bird on earth, with the total post-breeding population sometimes peaking at an estimated 1.5 billion individuals. The species is
1114:
moves from northern and central Sudan and central
Ethiopia in May and June, to breed in southern Sudan, South Sudan, southern Ethiopia and northern Kenya, moving back north from August to October.
1182:. Red-billed queleas may also roost together with weavers, estrildids and barn swallows. Their life expectancy is two to three years in the wild, but one captive bird lived for eighteen years.
1776:
965:
The females resemble the males in non-breeding plumage, but have a yellow or orangey bill and eye-ring during the breeding season. At other times, the female bill is pink or dull red.
1688:
at present directed at removing those congregations that are likely to attack vulnerable fields. In eastern and southern Africa, the control of quelea is often coordinated by the
481:
illustrated the species in colour, based on a live male specimen owned by a Mrs
Clayton in Surrey. He called it the "Brazilian sparrow", despite being unsure whether it came from
1689:
1085:
and can only be found within about 30 km (19 mi) distance of the nearest body of water. It is found in wet habitats, congregating at the shores of waterbodies, such as
1761:
2989:
Shefte, N.; Bruggers, R. L.; Schafer Jr., E. W. (April 1982). "Repellency and Toxicity of Three Bird Control Chemicals to Four Species of African Grain-Eating Birds".
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southern South Africa. It constructs oval roofed nests woven from strips of grass hanging from thorny branches, sugar cane or reeds. It breeds in very large colonies.
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Workshop on Research Priorities for Migrant Pests of Agriculture in Southern Africa, Plant Protection Research Institute, Pretoria, South Africa, 24 to 26 March 1999
4804:
1902:
2946:
Mulliè, Wim C. (2000). "Traditional capture of Red-billed Quelea Quelea quelea in the Lake Chad Basin and its possible role in reducing damage levels in cereals".
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fields or reeds. Colonies can consist of millions of nests, in densities of 30,000 per ha (12,000 per acre). Over 6000 nests in a single tree have been counted.
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The quelea feeds primarily on seeds of annual grasses, but also causes extensive damage to cereal crops. Therefore, it is sometimes called "Africa's feathered
4936:
1604:
is collected from under large roosts in Nigeria and used as a fertiliser. Tourists like to watch the large flocks of queleas, such as during visits of the
402:
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1260:-rich insects. Later the nestlings mainly get seeds. The young birds fledge after about two weeks in the nest. They are sexually mature in one year.
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when the rains have started, to eat the remaining ungerminated seeds. In Senegal migration is probably between the southeast and the northwest.
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months after hatching, after which the plumage resembles that of non-breeding adults, although the head is grey, the cheeks whitish, and wing
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The Problems of Birds as Pests: Proceedings of a Symposium Held at the Royal Geographical Society, London, on 28 and 29 September 1967
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1963:
Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis
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2564:"Lack of genetic and plumage differentiation in the red-billed quelea Quelea quelea across a migratory divide in southern Africa"
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410:
317:, is a small—approximately 12 cm (4.7 in) long and weighing 15–26 g (0.53–0.92 oz)—migratory, sparrow-like
212:
47:
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3307:
984:
Different subspecies are distinguished by different colour patterns of the male breeding plumage. In the typical subspecies,
364:
from Gabon to Mozambique and South Africa. Non-breeding birds have light underparts, striped brown upper parts, yellow-edged
4835:
439:
Ornithologia, sive Synopsis methodica sistens avium divisionem in ordines, sectiones, genera, species, ipsarumque varietates
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1653:, spiders, or boiled shredded egg during the breeding season. Fine stone grit and calcium sources, such as shell grit and
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30:
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3227:
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748:). The genus belongs to the group of true weavers (subfamily Ploceinae), and is most closely related to the fodies (
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3803:
3693:
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2437:
676:, meaning "quail", linking the prodigious numbers of queleas to the hordes of quail that fed the Israelites during
2887:
3903:
3365:
3204:
1669:
Sometimes called "Africa's feathered locust", the red-billed quelea is considered a serious agricultural pest in
1511:
that probably lays eggs in nests of queleas. Some predators, such as snakes, raid nests and eat eggs and chicks.
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527:
Three subspecies are recognised. In the field, these are distinguished by differences in male breeding plumage.
4502:
3113:
1357:
546:
442:
96:
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1451:) are eaten on a large scale. Red-billed queleas have also been observed feeding on crushed corn from cattle
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3753:
3713:
3136:
2710:
2067:
1528:
1519:
988:, breeding males have a buff crown, nape and underparts and the black mask extends high up the forehead. In
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2636:. Invasive animal risk assessment. Department of Agriculture and Fisheries, State of Queensland, Australia.
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1961:
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4003:
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3533:
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1491:, foxes, jackals, hyaenas, cats, lions and leopards. Bird species that prey on queleas include the
1211:
382:
61:
3108:
2837:
Crook, C.H.; Ward, P. (1967). "The Quelea Problem in Africa". In R.K. Murton; E.N. Wright (eds.).
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After 8:00 in the morning, tightly packed flocks await their chance to drink at waterholes in the
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again and their plumage starts to look like that of breeding adults, with a pinkish-purple bill.
601:
490:
341:
326:
240:
91:
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1068:, and its distribution covers most of sub-Saharan Africa. It avoids forests, however, including
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that was to be published in 1760, and which contained a black and white drawing of the species.
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showed that two-thirds of the seeds eaten belonged to only three species: African wild rice (
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Ward, Peter (1966). "Distribution, systematics, and polymorphism of the African weaver-bird
1971:
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1562:, three traditional methods are used to catch red-billed queleas. Trappers belonging to the
934:
764:. The following tree represents current insight of the relationships between the species of
629:
274:
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Natural enemies of the red-billed quelea include other birds, snakes, warthogs, squirrels,
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4882:
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4013:
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1992:. Vol. 2. Lugduni Batavorum (Leiden, Netherlands): Apud Theodorum Haak. p. 337.
462:. Brisson mentions that the bird originates from Senegal, where it had been collected by
2959:
2918:
Pocock, 1892 (Scorpiones, Liochelidae) as a possible predator of the red-billed quelea,
2732:
Crook, J. H. (1960). "Nest form and construction in certain West African weaver-birds".
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The red-billed quelea is mostly found in tropical and subtropical areas with a seasonal
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4086:
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1526:. Among the invertebrates that kill and eat youngsters are the armoured bush cricket (
1146:
is needed for the queleas to gain enough fat to allow migration to new feeding areas.
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1957:
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406:
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208:
81:
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the mask also extends high up the forehead, but the underparts are mainly white. In
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and arid coastal regions of Namibia and Angola. It was introduced to the island of
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949:
Over 75% of males have a black facial "mask", comprising a black forehead, cheeks,
723:
4757:
4705:
3025:
2967:
2537:
1568:
1033:
758:. These two genera are in turn the sister clade to the Asian species of the genus
3076:. Desert Locust Control Organization for Eastern Africa (DLCO-EA). Archived from
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morning, dried in the sun, and transported to the city to be sold on the market.
592:, with no evidence of genetic isolation. Hence it is not recognised as distinct.
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to ensure year-round food availability. The consumption of a lot of food with a
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such as those in central Africa, and is generally absent from western parts of
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are edged greenish or yellow. The eye has a narrow naked red ring and a brown
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349:
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International Red Locust Control Organization for Central and Southern Africa
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1967:
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1657:, provide nutrients as well. If provided with material like fresh grass or
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In the breeding season, males are diversely coloured. These differences in
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and flight feathers have buff margins. At an age of about five months they
2761:"An Experimental Study of the Mechanisms of Nest Building in a Weaverbird"
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1897:
1558:
The red-billed quelea is caught and eaten in many parts of Africa. Around
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between red-billed and red-headed queleas has been observed in captivity.
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2711:"The weaving of the red-billed weaver bird, Quelea quelea in captivity"
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1696:(IRLCO-CSA), which make their aircraft available for this purpose.
726:
of a clade that contains both other remaining species of the genus
4726:
3167:
1601:
1488:
1466:
1420:
1189:
1065:
968:
Newborns have white bills and are almost naked with some wisps of
706:"Red-billed quelea" has been designated the official name by the
672:. M.W. Jeffreys suggested that the term came from medieval Latin
1658:
1412:
1380:
1150:
1090:
318:
138:
4605:
3125:
2562:
Dallimer, M.; Jones, P.J; Pemberton, J.M.; Cheke, R.A. (2003).
1624:. The animal's large distribution and population resulted in a
1089:, during flooding. It needs shrubs, reeds or trees to nest and
2855:
Ward, Peter (1965). "Feeding ecology of the black-faced dioch
2665:) Using Remote Sensing And Geographic Information System (GIS)
1428:
1256:
days. After the chicks hatch, they are fed for some days with
286:
2809:. Princeton, New Jersey: Princeton University Press. p.
2280:(in German). Vol. 3. Neudamm: J. Neumann. p. 109.
2220:
Jones, P.J.; Dallimer, M.; Cheke, R.A.; Mundy, P.J. (2002).
997:
the ranges of the subspecies overlap, such as at Lake Chad.
2627:
Markula, Anna; Hannan-Jones, Martin; Csurhes, Steve (200).
1117:
In southern Africa, the total population of the subspecies
295:
292:
2805:
Avian Architecture: How Birds Design, Engineer, and Build
1594:) are used to catch hundreds of these birds daily in the
16:
Small, migratory weaver bird native to Sub-Saharan Africa
2685:
Traditional Ecological Knowledge: A Collection of Essays
474:
in Latin, both meaning "red-billed Senegalese sparrow".
452:
of India was corrected to Africa in the 12th edition of
2167:
The Journal of the Royal Geographical Society of London
2163:"Report of the Expedition for Exploring Central Africa"
1579:
use standing fishing nets with a very fine mesh, while
1064:, resulting in dry thornbush grassland, including the
3031:. In R.A. Cheke; L.J. Rosenberg; M.E. Kieser (eds.).
3026:"5. Quelea Management in Southern and Eastern Africa"
1690:
Desert Locust Control Organization for Eastern Africa
1367:). When the supply of these seeds runs out, seeds of
754:), a genus of six or seven species that occur on the
283:
280:
2661:
Monitoring and Habitat Location of the Weaver bird (
1752:
in breeding plumage with a yellow wash on the head,
579:
Formerly, two other subspecies have been described.
466:
during his 1748-1752 expedition. He called the bird
289:
4614:
4576:
4391:
4306:
4261:
4238:
4215:
4100:
4077:
3412:
3389:
3354:
3331:
3276:
3241:
3218:
3183:
3165:
277:
2802:
1970:: (Laurentii Salvii). p. 176 – via The
497:, but rather a weaver, and created the genus name
1966:(in Latin). Vol. I (10th revised ed.).
1838:10.2305/IUCN.UK.2016-3.RLTS.T22719128A94613042.en
2888:"Turtles take Red-billed Quelea (Quelea quelea)"
2759:Collias, Nicholas E.; Collias, Elsie C. (1962).
2420:"NRM 568681 Ploceus aethiopicus Sundevall, 1850"
2222:"Are there two subspecies of Red-billed Quelea,
1608:. The birds themselves eat pest insects such as
1013:
687:is probably named in honor of the ornithologist
376:". The usual pest-control measures are spraying
2442:"Old World sparrows, snowfinches & weavers"
2206:Check-list of the birds of the world. Volume 15
1767:Females in breeding plumage with yellow bills,
2929:Bulletin of the British Arachnological Society
1538:. Internal parasites found in queleas include
3137:
8:
2368:(3rd ed.). Robina: Alexander Editions.
2329:The Helm Dictionary of Scientific Bird Names
2209:. Museum of Comparative Zoology. p. 62.
1903:Random House Webster's Unabridged Dictionary
2198:
2196:
1170:), and in western Africa they may join the
4602:
3144:
3130:
3122:
3052:. Springer Sciencefiction & Business.
2912:Vincent, Leonard S.; Breitman, Ty (2010).
2622:
2620:
2618:
2616:
1153:with thorny or spiny vegetation—typically
616:Linnaeus himself did not explain the name
401:The red-billed quelea was one of the many
221:
70:
46:
29:
20:
2850:
2848:
1952:
1950:
1836:
520:in 1877 and named after the aviculturist
3046:McCullough, Dale; Barrett, R.H. (2012).
2136:"Zwartmasker roodbekwever Quelea quelea"
2092:
2090:
2088:
1149:When breeding, it selects areas such as
600:in 1886 from east Africa, is regarded a
360:from Sudan to Somalia and Tanzania, and
356:occurring roughly from Senegal to Chad,
2948:Ostrich: Journal of African Ornithology
2051:
2049:
2047:
2045:
2043:
2041:
2039:
2037:
2035:
2033:
2031:
2029:
2027:
2025:
2023:
2021:
2019:
1799:
1703:
2653:
2651:
2649:
2647:
2645:
2643:
2130:
2128:
2126:
2124:
2017:
2015:
2013:
2011:
2009:
2007:
2005:
2003:
2001:
1999:
1852:
1850:
1848:
1805:
1803:
708:International Ornithological Committee
2658:Abdelwahid, Amel Abdelraheem (2008).
2470:Molecular Phylogenetics and Evolution
2448:. International Ornithologists' Union
512:was described as a separate species,
344:assigned it in 1850 to the new genus
7:
2332:. London: Christopher Helm. p.
1924:
1922:
1920:
1918:
1916:
1914:
1912:
1862:Handbook of Birds of the World Alive
1733:in breeding plumage with pink wash,
4937:IUCN Red List least concern species
1824:IUCN Red List of Threatened Species
1644:The red-billed quelea is sometimes
822:
798:
785:
778:
771:
756:islands of the western Indian Ocean
2991:The Journal of Wildlife Management
2873:10.1111/j.1474-919X.1965.tb07296.x
2746:10.1111/j.1474-919X.1960.tb05090.x
2115:10.1111/j.1474-919X.1966.tb07250.x
1985:Brisson, Mathurin Jacques (1763).
1109:In eastern Africa, the subspecies
699:was collected in the neighbouring
472:Passer senegalensis erythrorynchos
14:
2513:"Red-billed Quelea Quelea quelea"
3318:Donaldson Smith's sparrow-weaver
3093:
2591:10.1046/j.1365-294X.2003.01733.x
1775:
1760:
1741:
1722:
1706:
1283:
1272:
1018:
273:
95:
4784:red-billed-quelea-quelea-quelea
3298:Chestnut-crowned sparrow-weaver
2682:Johannes, Robert Earle (1989).
2440:; Donsker, David, eds. (2017).
768:, and their closest relatives.
722:, the red-billed quelea is the
567:in 1850, but later assigned to
549:in 1836, but later assigned to
3664:Northern brown-throated weaver
3654:Southern brown-throated weaver
3308:Chestnut-backed sparrow-weaver
612:Etymology and vernacular names
493:thought the species was not a
468:Moineau à bec rouge du Senegal
1:
2968:10.1080/00306525.2000.9639856
2841:(revised ed.). Elsevier.
2058:"Quelea quelea (weaver bird)"
691:. The name of the subspecies
336:in 1758, who considered it a
3024:Elliott, Clive C.H. (2000).
2302:"Roodbekwever Quelea quelea"
1121:in October converges on the
695:refers to Ethiopia, and its
4957:Birds of Sub-Saharan Africa
4952:Taxa named by Carl Linnaeus
3288:White-browed sparrow-weaver
3228:White-headed buffalo weaver
3195:White-billed buffalo weaver
2709:Friedmann, Herbert (1924).
2482:10.1016/j.ympev.2016.12.013
2446:World Bird List Version 7.1
2400:. World Agroforestry Center
2063:Invasive Species Compendium
908:Asian species of the genus
4980:
3376:Black-capped social weaver
2801:Goodfellow, Peter (2011).
2389:Orwa; et al. (2009).
2326:Jobling, James A. (2010).
1646:kept and bred in captivity
358:Q. q. aethiopica
3366:Grey-capped social weaver
3205:Red-billed buffalo weaver
3109:"Red-billed quelea media"
2671:. University of Khartoum.
2424:Naturhistorica Riksmuseet
2398:Agroforestry Database 4.0
2274:Reichenow, Anton (1900).
2253:10.2989/00306520209485349
2203:Peters, James L. (1962).
1786:in non-breeding plumage,
994:Q. quelea aethiopica
906:
879:
844:
827:
820:
803:
796:
783:
776:
594:Q. quelea intermedia
458:of 1766, and Brisson was
246:
239:
229:
220:
197:
190:
92:Scientific classification
90:
68:
59:
54:
45:
38:Male breeding plumage of
37:
28:
23:
4503:Yellow-mantled widowbird
3114:Internet Bird Collection
2663:Quelea quelea aethiopica
2140:Werkgroep voor Ploceidae
1831:: e.T22719128A94613042.
1554:Interactions with humans
1520:African helmeted turtles
1358:Sorghum purpureosericeum
1056:Distribution and habitat
581:Q. quelea spoliator
443:Mathurin Jacques Brisson
419:. Classifying it in the
385:of red-billed quelea is
362:Q. q. lathamii
40:Q. q. lathamii
4947:Birds described in 1758
4433:Black-winged red bishop
3804:Vieillot's black weaver
3754:Tanzanian masked weaver
3714:Vitelline masked weaver
3694:Heuglin's masked weaver
2825:Red-billed quelea nest.
2068:University of Greenwich
1612:, and the moth species
1529:Acanthoplus discoidalis
1471:Flocking at a waterhole
1463:Predators and parasites
1119:Q. quelea lathamii
590:Q. quelea lathamii
4523:Red-collared widowbird
4513:White-winged widowbird
3904:Maxwell's black weaver
3724:Southern masked weaver
3704:Northern masked weaver
3686:or Rueppell's weaver (
3263:Scaly-feathered weaver
3253:Speckle-fronted weaver
2886:Robel, Detlef (2008).
2511:Weaver Research Unit.
2226:, in southern Africa?"
2161:Smith, Andrew (1836).
2056:Cheke, Robert (2015).
1870:10.2173/bow.rebque1.01
1811:BirdLife International
1472:
1199:
1050:tweedle-toodle-tweedle
321:of the weaver family,
4553:Long-tailed widowbird
4403:Yellow-crowned bishop
3974:Black-breasted weaver
3894:Yellow-mantled weaver
3734:Katanga masked weaver
3594:Holub's golden weaver
3584:Eastern golden weaver
3464:Slender-billed weaver
1470:
1441:Phagopyrum esculentum
1251:Both sexes share the
1218:Dichrostachys cinerea
1193:
1129:Ecology and behaviour
986:Q. quelea quelea
409:in the landmark 1758
352:are recognised, with
309:), also known as the
55:Non-breeding plumage
4493:Fan-tailed widowbird
4473:Golden-backed bishop
4064:Yellow-legged weaver
4004:Yellow-capped weaver
3854:Golden-backed weaver
3744:Lufira masked weaver
3494:Lesser masked weaver
3434:Black-chinned weaver
3341:Rufous-tailed weaver
2517:Weavers of the World
2391:"Xylopia aethiopica"
1615:Helicoverpa armigera
1606:Kruger National Park
1221:), but sometimes in
1194:Two males of subsp.
1172:Sudan golden sparrow
565:Carl Jakob Sundevall
537:Quelea quelea quelea
441:, a book written by
354:Quelea quelea quelea
4586:Thick-billed weaver
4563:Jackson's widowbird
4463:Zanzibar red bishop
4453:Southern red bishop
4443:Northern red bishop
4413:Fire-fronted bishop
4322:F. madagascariensis
4182:Blue-billed malimbe
4172:Red-bellied malimbe
4112:Red-crowned malimbe
4034:Brown-capped weaver
4024:Olive-headed weaver
3954:Asian golden weaver
3884:Golden-naped weaver
3834:Black-headed weaver
3544:Black-billed weaver
3534:Black-necked weaver
2960:2000Ostri..71...15M
2916:Cheloctonus jonesii
2583:2003MolEc..12..345D
2245:2002Ostri..73...36J
1930:"Red-billed Quelea"
1858:"Red-billed quelea"
1626:conservation status
1591:Cynodon nlemfuensis
1535:Cheloctonus jonesii
1532:) and the scorpion
1212:Senegalia mellifera
1144:high energy content
1123:Zimbabwean Highveld
626:kwelea domo-jekundu
561:Ploceus aethiopicus
397:Taxonomy and naming
383:conservation status
253:Linnaeus, 1758
62:Conservation status
4942:Agricultural pests
4445:or orange bishop (
4202:Red-headed malimbe
4152:Red-vented malimbe
3984:Dark-backed weaver
3624:Golden palm weaver
3514:Bannerman's weaver
2366:Australian Parrots
2358:Forshaw, Joseph M.
2239:(1&2): 36–42.
1692:(DLCO-EA) and the
1671:Sub-Saharan Africa
1661:they can be bred.
1621:Spodoptera exempta
1473:
1364:Echinochloa colona
1215:) and sicklebush (
1206:Vachellia tortilis
1200:
1198:at their nest site
990:Q. quelea lathamii
831:Q. cardinalis
738:Q. cardinalis
703:in today's Sudan.
624:is locally called
571:as its subspecies
553:as its subspecies
508:. The white-faced
491:Ludwig Reichenbach
342:Ludwig Reichenbach
327:Sub-Saharan Africa
234:rough distribution
24:Red-billed quelea
4924:
4923:
4896:Open Tree of Life
4608:Taxon identifiers
4599:
4598:
4543:Montane widowbird
4293:Red-billed quelea
4283:Red-headed quelea
4248:Bob-tailed weaver
4225:Red-headed weaver
4044:Bar-winged weaver
4008:P. dorsomaculatus
3914:Nelicourvi weaver
3848:P. dichrocephalus
3838:P. melanocephalus
3504:Spectacled weaver
3444:Baglafecht weaver
3312:P. rufoscapulatus
3074:"The Quelea Bird"
2922:(Linnaeus, 1758)"
2630:Red-billed quelea
2571:Molecular Ecology
2375:978-0-9581212-0-0
2362:William T. Cooper
2343:978-1-4081-2501-4
2277:Die Vögel Afrikas
1717:published in 1760
1610:migratory locusts
1449:Helianthus annuus
1409:Panicum miliaceum
1361:and jungle rice (
1062:semi-arid climate
1044:
1043:
1002:pin-tailed whydah
931:
930:
922:
921:
895:
894:
868:
867:
859:
858:
848:Q. erythrops
746:Q. erythrops
742:red-headed quelea
583:was described by
563:was described by
545:was described by
501:, as well as the
429:, he gave it the
389:according to the
311:red-billed weaver
269:red-billed quelea
265:
264:
85:
4969:
4917:
4916:
4904:
4903:
4891:
4890:
4878:
4877:
4865:
4864:
4862:NHMSYS0001689630
4852:
4851:
4839:
4838:
4826:
4825:
4813:
4812:
4800:
4799:
4787:
4786:
4774:
4773:
4761:
4760:
4748:
4747:
4735:
4734:
4722:
4721:
4709:
4708:
4699:
4698:
4686:
4685:
4676:
4675:
4673:2B6DE1366B1D557C
4663:
4662:
4650:
4649:
4648:
4635:
4634:
4633:
4603:
4547:E. psammacromius
4332:F. eminentissima
4176:M. erythrogaster
4142:Rachel's malimbe
4122:Cassin's malimbe
4091:P. superciliosus
3684:Rüppell's weaver
3674:Kilombero weaver
3608:Ploceus princeps
3478:P. subpersonatus
3454:Bertram's weaver
3428:P. superciliosus
3302:P. superciliosus
3146:
3139:
3132:
3123:
3118:
3098:Data related to
3097:
3082:
3081:
3070:
3064:
3063:
3043:
3037:
3036:
3030:
3021:
3015:
3014:
2986:
2980:
2979:
2943:
2937:
2936:
2926:
2909:
2903:
2902:
2892:
2883:
2877:
2876:
2852:
2843:
2842:
2834:
2828:
2827:
2808:
2798:
2792:
2791:
2765:
2756:
2750:
2749:
2729:
2723:
2722:
2706:
2700:
2699:
2679:
2673:
2672:
2670:
2655:
2638:
2637:
2635:
2624:
2611:
2610:
2568:
2559:
2553:
2552:
2550:
2548:
2534:
2528:
2527:
2525:
2523:
2508:
2502:
2501:
2464:
2458:
2457:
2455:
2453:
2434:
2428:
2427:
2416:
2410:
2409:
2407:
2405:
2395:
2386:
2380:
2379:
2354:
2348:
2347:
2323:
2317:
2316:
2314:
2312:
2298:
2292:
2291:
2271:
2265:
2264:
2230:
2217:
2211:
2210:
2200:
2191:
2190:
2158:
2152:
2151:
2149:
2147:
2132:
2119:
2118:
2094:
2083:
2082:
2080:
2078:
2053:
1994:
1993:
1982:
1976:
1975:
1972:Internet Archive
1954:
1945:
1944:
1942:
1940:
1926:
1907:
1895:
1889:
1888:
1886:
1884:
1854:
1843:
1842:
1840:
1807:
1784:Q. q. aethiopica
1779:
1764:
1750:Q. q. aethiopica
1745:
1731:Q. q. aethiopica
1726:
1710:
1377:Hordeum disticum
1287:
1276:
1070:miombo woodlands
1022:
1014:
823:
799:
786:
779:
772:
718:Based on recent
332:It was named by
315:red-billed dioch
302:
301:
298:
297:
294:
291:
288:
285:
282:
279:
254:
233:
225:
203:
100:
99:
79:
74:
73:
50:
33:
21:
4979:
4978:
4972:
4971:
4970:
4968:
4967:
4966:
4927:
4926:
4925:
4920:
4912:
4907:
4899:
4894:
4886:
4883:Observation.org
4881:
4873:
4868:
4860:
4855:
4847:
4842:
4834:
4829:
4821:
4816:
4808:
4803:
4795:
4790:
4782:
4777:
4769:
4764:
4756:
4751:
4743:
4738:
4730:
4725:
4717:
4712:
4704:
4702:
4694:
4689:
4681:
4679:
4671:
4666:
4658:
4653:
4644:
4643:
4638:
4629:
4628:
4623:
4610:
4600:
4595:
4572:
4533:Marsh widowbird
4467:E. nigroventris
4455:or red bishop (
4447:E. franciscanus
4387:
4368:Seychelles fody
4302:
4273:Cardinal quelea
4257:
4234:
4211:
4192:Crested malimbe
4096:
4073:
4058:P. sanctithomae
4054:São Tomé weaver
4028:P. olivaceiceps
4014:Usambara weaver
3994:Preuss's weaver
3978:P. benghalensis
3968:P. megarhynchus
3934:Streaked weaver
3924:Sakalava weaver
3874:Chestnut weaver
3864:Cinnamon weaver
3824:Clarke's weaver
3708:P. taeniopterus
3658:P. xanthopterus
3638:P. castaneiceps
3604:Príncipe weaver
3564:Bocage's weaver
3548:P. melanogaster
3408:
3399:Sociable weaver
3385:
3350:
3327:
3272:
3237:
3214:
3179:
3161:
3150:
3107:
3090:
3085:
3080:on 28 May 2022.
3072:
3071:
3067:
3060:
3045:
3044:
3040:
3028:
3023:
3022:
3018:
3003:10.2307/3808656
2988:
2987:
2983:
2945:
2944:
2940:
2924:
2911:
2910:
2906:
2890:
2885:
2884:
2880:
2854:
2853:
2846:
2836:
2835:
2831:
2821:
2800:
2799:
2795:
2780:10.2307/4082640
2763:
2758:
2757:
2753:
2731:
2730:
2726:
2708:
2707:
2703:
2696:
2681:
2680:
2676:
2668:
2657:
2656:
2641:
2633:
2626:
2625:
2614:
2566:
2561:
2560:
2556:
2546:
2544:
2538:"Quelea quelea"
2536:
2535:
2531:
2521:
2519:
2510:
2509:
2505:
2466:
2465:
2461:
2451:
2449:
2436:
2435:
2431:
2418:
2417:
2413:
2403:
2401:
2393:
2388:
2387:
2383:
2376:
2356:
2355:
2351:
2344:
2325:
2324:
2320:
2310:
2308:
2300:
2299:
2295:
2288:
2273:
2272:
2268:
2228:
2219:
2218:
2214:
2202:
2201:
2194:
2179:10.2307/1797576
2160:
2159:
2155:
2145:
2143:
2134:
2133:
2122:
2096:
2095:
2086:
2076:
2074:
2055:
2054:
1997:
1984:
1983:
1979:
1956:
1955:
1948:
1938:
1936:
1928:
1927:
1910:
1896:
1892:
1882:
1880:
1856:
1855:
1846:
1809:
1808:
1801:
1797:
1790:
1780:
1771:
1765:
1756:
1746:
1737:
1727:
1718:
1711:
1702:
1679:organophosphate
1667:
1665:Pest management
1642:
1596:Kondoa District
1556:
1515:Nile crocodiles
1505:diederik cuckoo
1465:
1401:Setaria italica
1393:Sorghum bicolor
1303:
1302:
1301:
1300:
1299:
1288:
1279:
1278:
1277:
1266:
1209:), blackthorn (
1188:
1164:) and bishops (
1131:
1058:
1045:
1040:
1031:
1030:
1028:Songs and calls
1011:
956:flight feathers
943:
932:
923:
896:
869:
860:
734:cardinal quelea
716:
701:Sennar province
683:The subspecies
670:Tsonga language
614:
598:Anton Reichenow
596:, described by
585:Phillip Clancey
503:new combination
455:Systema Naturae
435:Emberiza quelea
416:Systema Naturae
403:birds described
399:
366:flight feathers
276:
272:
252:
250:Emberiza quelea
235:
231:
216:
205:
199:
186:
94:
86:
75:
71:
64:
17:
12:
11:
5:
4977:
4976:
4973:
4965:
4964:
4959:
4954:
4949:
4944:
4939:
4929:
4928:
4922:
4921:
4919:
4918:
4905:
4892:
4879:
4866:
4853:
4840:
4827:
4814:
4801:
4788:
4775:
4762:
4749:
4736:
4723:
4710:
4700:
4687:
4677:
4664:
4651:
4636:
4620:
4618:
4612:
4611:
4606:
4597:
4596:
4594:
4593:
4582:
4580:
4574:
4573:
4571:
4570:
4560:
4550:
4540:
4530:
4520:
4517:E. albonotatus
4510:
4500:
4490:
4480:
4470:
4460:
4450:
4440:
4430:
4420:
4410:
4399:
4397:
4389:
4388:
4386:
4385:
4378:Rodrigues fody
4375:
4372:F. sechellarum
4365:
4358:Mauritius fody
4355:
4345:
4335:
4325:
4314:
4312:
4304:
4303:
4301:
4300:
4290:
4280:
4269:
4267:
4259:
4258:
4256:
4255:
4244:
4242:
4236:
4235:
4233:
4232:
4221:
4219:
4213:
4212:
4210:
4209:
4206:M. rubricollis
4199:
4189:
4179:
4169:
4166:M. ibadanensis
4162:Ibadan malimbe
4159:
4149:
4139:
4129:
4119:
4108:
4106:
4098:
4097:
4095:
4094:
4087:Compact weaver
4083:
4081:
4075:
4074:
4072:
4071:
4061:
4051:
4041:
4031:
4021:
4011:
4001:
3991:
3981:
3971:
3961:
3958:P. hypoxanthus
3951:
3948:P. philippinus
3941:
3931:
3921:
3911:
3901:
3891:
3881:
3878:P. rubiginosus
3871:
3861:
3851:
3841:
3831:
3821:
3814:Weyns's weaver
3811:
3801:
3791:
3784:Speke's weaver
3781:
3771:
3764:Village weaver
3761:
3751:
3741:
3731:
3721:
3711:
3701:
3691:
3681:
3671:
3661:
3651:
3648:P. holoxanthus
3641:
3631:
3621:
3611:
3601:
3591:
3581:
3571:
3561:
3554:Strange weaver
3551:
3541:
3538:P. nigricollis
3531:
3524:Bates's weaver
3521:
3511:
3501:
3498:P. intermedius
3491:
3481:
3471:
3461:
3451:
3441:
3438:P. nigrimentus
3431:
3424:Compact weaver
3420:
3418:
3410:
3409:
3407:
3406:
3395:
3393:
3387:
3386:
3384:
3383:
3373:
3362:
3360:
3352:
3351:
3349:
3348:
3337:
3335:
3329:
3328:
3326:
3325:
3315:
3305:
3295:
3284:
3282:
3274:
3273:
3271:
3270:
3267:S. squamifrons
3260:
3249:
3247:
3239:
3238:
3236:
3235:
3224:
3222:
3216:
3215:
3213:
3212:
3202:
3199:B. albirostris
3191:
3189:
3181:
3180:
3173:
3171:
3163:
3162:
3151:
3149:
3148:
3141:
3134:
3126:
3120:
3119:
3105:
3104:at Wikispecies
3089:
3088:External links
3086:
3084:
3083:
3065:
3058:
3038:
3016:
2997:(2): 453–457.
2981:
2954:(1–2): 15–20.
2938:
2914:"The scorpion
2904:
2878:
2867:(2): 173–214.
2844:
2829:
2819:
2793:
2774:(4): 568–595.
2751:
2724:
2721:(16): 357–372.
2701:
2694:
2674:
2639:
2612:
2577:(2): 345–353.
2554:
2529:
2503:
2459:
2429:
2411:
2381:
2374:
2349:
2342:
2318:
2293:
2286:
2266:
2212:
2192:
2153:
2120:
2084:
2072:United Kingdom
1995:
1977:
1958:Linnaeus, Carl
1946:
1908:
1890:
1844:
1798:
1796:
1793:
1792:
1791:
1781:
1774:
1772:
1766:
1759:
1757:
1747:
1740:
1738:
1728:
1721:
1719:
1715:George Edwards
1712:
1705:
1701:
1698:
1666:
1663:
1641:
1638:
1555:
1552:
1509:brood parasite
1464:
1461:
1435:), as well as
1385:Eragrostis tef
1345:One survey at
1289:
1282:
1281:
1280:
1271:
1270:
1269:
1268:
1267:
1265:
1262:
1244:do not signal
1187:
1184:
1178:) and various
1130:
1127:
1057:
1054:
1042:
1041:
1032:
1026:
1024:
1012:
1010:
1007:
942:
939:
929:
928:
925:
924:
920:
919:
916:
915:
905:
902:
901:
898:
897:
893:
892:
889:
888:
878:
875:
874:
871:
870:
866:
865:
862:
861:
857:
856:
853:
852:
843:
840:
839:
836:
835:
826:
821:
819:
816:
815:
812:
811:
802:
797:
795:
784:
782:
777:
775:
770:
715:
712:
613:
610:
604:of subspecies
577:
576:
569:Q. quelea
558:
551:Q. quelea
543:Loxia lathamii
540:
514:Q. russii
506:Q. quelea
479:George Edwards
477:Also in 1766,
470:in French and
464:Michel Adanson
448:The erroneous
405:originally by
398:
395:
263:
262:
261:
260:
255:
244:
243:
237:
236:
230:
227:
226:
218:
217:
206:
195:
194:
188:
187:
183:Q. quelea
180:
178:
174:
173:
166:
162:
161:
156:
152:
151:
146:
142:
141:
136:
132:
131:
126:
122:
121:
116:
112:
111:
106:
102:
101:
88:
87:
69:
66:
65:
60:
57:
56:
52:
51:
43:
42:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
4975:
4974:
4963:
4960:
4958:
4955:
4953:
4950:
4948:
4945:
4943:
4940:
4938:
4935:
4934:
4932:
4915:
4914:Quelea-quelea
4910:
4906:
4902:
4897:
4893:
4889:
4884:
4880:
4876:
4871:
4867:
4863:
4858:
4854:
4850:
4845:
4841:
4837:
4832:
4828:
4824:
4819:
4815:
4811:
4806:
4802:
4798:
4793:
4789:
4785:
4780:
4776:
4772:
4767:
4763:
4759:
4754:
4750:
4746:
4741:
4737:
4733:
4728:
4724:
4720:
4715:
4711:
4707:
4701:
4697:
4692:
4688:
4684:
4678:
4674:
4669:
4665:
4661:
4660:Quelea_quelea
4656:
4652:
4647:
4646:Quelea quelea
4641:
4637:
4632:
4626:
4622:
4621:
4619:
4617:
4616:Quelea quelea
4613:
4609:
4604:
4591:
4587:
4584:
4583:
4581:
4579:
4575:
4568:
4564:
4561:
4558:
4554:
4551:
4548:
4544:
4541:
4538:
4534:
4531:
4528:
4524:
4521:
4518:
4514:
4511:
4508:
4504:
4501:
4498:
4494:
4491:
4488:
4484:
4483:Yellow bishop
4481:
4478:
4474:
4471:
4468:
4464:
4461:
4458:
4454:
4451:
4448:
4444:
4441:
4438:
4437:E. hordeaceus
4434:
4431:
4428:
4424:
4421:
4418:
4417:E. diadematus
4414:
4411:
4408:
4404:
4401:
4400:
4398:
4396:
4395:
4390:
4383:
4379:
4376:
4373:
4369:
4366:
4363:
4359:
4356:
4353:
4349:
4346:
4343:
4339:
4336:
4333:
4329:
4326:
4323:
4319:
4316:
4315:
4313:
4311:
4310:
4305:
4298:
4294:
4291:
4288:
4284:
4281:
4278:
4277:Q. cardinalis
4274:
4271:
4270:
4268:
4266:
4265:
4260:
4253:
4249:
4246:
4245:
4243:
4241:
4237:
4230:
4226:
4223:
4222:
4220:
4218:
4214:
4207:
4203:
4200:
4197:
4196:M. malimbicus
4193:
4190:
4187:
4183:
4180:
4177:
4173:
4170:
4167:
4163:
4160:
4157:
4153:
4150:
4147:
4143:
4140:
4137:
4133:
4130:
4127:
4123:
4120:
4117:
4113:
4110:
4109:
4107:
4105:
4104:
4099:
4092:
4088:
4085:
4084:
4082:
4080:
4076:
4069:
4065:
4062:
4059:
4055:
4052:
4049:
4048:P. angolensis
4045:
4042:
4039:
4035:
4032:
4029:
4025:
4022:
4019:
4015:
4012:
4009:
4005:
4002:
3999:
3995:
3992:
3989:
3985:
3982:
3979:
3975:
3972:
3969:
3965:
3964:Finn's weaver
3962:
3959:
3955:
3952:
3949:
3945:
3942:
3939:
3935:
3932:
3929:
3925:
3922:
3919:
3918:P. nelicourvi
3915:
3912:
3909:
3905:
3902:
3899:
3895:
3892:
3889:
3888:P. aureonucha
3885:
3882:
3879:
3875:
3872:
3869:
3865:
3862:
3859:
3855:
3852:
3849:
3845:
3842:
3839:
3835:
3832:
3829:
3825:
3822:
3819:
3815:
3812:
3809:
3808:P. nigerrimus
3805:
3802:
3799:
3798:P. spekeoides
3795:
3792:
3789:
3785:
3782:
3779:
3775:
3772:
3769:
3768:P. cucullatus
3765:
3762:
3759:
3755:
3752:
3749:
3745:
3742:
3739:
3735:
3732:
3729:
3725:
3722:
3719:
3718:P. vitellinus
3715:
3712:
3709:
3705:
3702:
3699:
3695:
3692:
3689:
3685:
3682:
3679:
3675:
3672:
3669:
3665:
3662:
3659:
3655:
3652:
3649:
3645:
3642:
3639:
3635:
3634:Taveta weaver
3632:
3629:
3625:
3622:
3619:
3615:
3614:Orange weaver
3612:
3609:
3605:
3602:
3599:
3595:
3592:
3589:
3585:
3582:
3579:
3575:
3572:
3569:
3568:P. temporalis
3565:
3562:
3559:
3555:
3552:
3549:
3545:
3542:
3539:
3535:
3532:
3529:
3525:
3522:
3519:
3518:P. bannermani
3515:
3512:
3509:
3505:
3502:
3499:
3495:
3492:
3489:
3485:
3484:Little weaver
3482:
3479:
3475:
3474:Loango weaver
3472:
3469:
3465:
3462:
3459:
3455:
3452:
3449:
3448:P. baglafecht
3445:
3442:
3439:
3435:
3432:
3429:
3425:
3422:
3421:
3419:
3417:
3416:
3411:
3404:
3400:
3397:
3396:
3394:
3392:
3388:
3381:
3377:
3374:
3371:
3367:
3364:
3363:
3361:
3359:
3358:
3357:Pseudonigrita
3353:
3346:
3342:
3339:
3338:
3336:
3334:
3330:
3323:
3322:P. donaldsoni
3319:
3316:
3313:
3309:
3306:
3303:
3299:
3296:
3293:
3289:
3286:
3285:
3283:
3281:
3280:
3275:
3268:
3264:
3261:
3258:
3254:
3251:
3250:
3248:
3246:
3245:
3240:
3233:
3229:
3226:
3225:
3223:
3221:
3217:
3210:
3206:
3203:
3200:
3196:
3193:
3192:
3190:
3188:
3187:
3182:
3178:
3177:
3172:
3170:
3169:
3164:
3160:
3159:
3154:
3147:
3142:
3140:
3135:
3133:
3128:
3127:
3124:
3116:
3115:
3110:
3106:
3103:
3102:
3101:Quelea quelea
3096:
3092:
3091:
3087:
3079:
3075:
3069:
3066:
3061:
3059:9789401128681
3055:
3051:
3050:
3049:Wildlife 2001
3042:
3039:
3034:
3027:
3020:
3017:
3012:
3008:
3004:
3000:
2996:
2992:
2985:
2982:
2977:
2973:
2969:
2965:
2961:
2957:
2953:
2949:
2942:
2939:
2934:
2930:
2923:
2921:
2920:Quelea quelea
2917:
2908:
2905:
2900:
2896:
2889:
2882:
2879:
2874:
2870:
2866:
2862:
2859:in Nigeria".
2858:
2857:Quelea quelea
2851:
2849:
2845:
2840:
2833:
2830:
2826:
2822:
2820:9781400838318
2816:
2812:
2807:
2806:
2797:
2794:
2789:
2785:
2781:
2777:
2773:
2769:
2762:
2755:
2752:
2747:
2743:
2739:
2735:
2728:
2725:
2720:
2716:
2712:
2705:
2702:
2697:
2695:9782880329983
2691:
2687:
2686:
2678:
2675:
2667:
2666:
2662:
2654:
2652:
2650:
2648:
2646:
2644:
2640:
2632:
2631:
2623:
2621:
2619:
2617:
2613:
2608:
2604:
2600:
2596:
2592:
2588:
2584:
2580:
2576:
2572:
2565:
2558:
2555:
2543:
2539:
2533:
2530:
2518:
2514:
2507:
2504:
2499:
2495:
2491:
2487:
2483:
2479:
2475:
2471:
2463:
2460:
2447:
2443:
2439:
2433:
2430:
2425:
2421:
2415:
2412:
2399:
2392:
2385:
2382:
2377:
2371:
2367:
2363:
2359:
2353:
2350:
2345:
2339:
2335:
2331:
2330:
2322:
2319:
2307:
2303:
2297:
2294:
2289:
2287:9785882381157
2283:
2279:
2278:
2270:
2267:
2262:
2258:
2254:
2250:
2246:
2242:
2238:
2234:
2227:
2225:
2224:Quelea quelea
2216:
2213:
2208:
2207:
2199:
2197:
2193:
2188:
2184:
2180:
2176:
2172:
2168:
2164:
2157:
2154:
2141:
2137:
2131:
2129:
2127:
2125:
2121:
2116:
2112:
2108:
2104:
2100:
2099:Quelea quelea
2093:
2091:
2089:
2085:
2073:
2069:
2065:
2064:
2059:
2052:
2050:
2048:
2046:
2044:
2042:
2040:
2038:
2036:
2034:
2032:
2030:
2028:
2026:
2024:
2022:
2020:
2018:
2016:
2014:
2012:
2010:
2008:
2006:
2004:
2002:
2000:
1996:
1991:
1990:
1981:
1978:
1973:
1969:
1965:
1964:
1959:
1953:
1951:
1947:
1935:
1931:
1925:
1923:
1921:
1919:
1917:
1915:
1913:
1909:
1905:
1904:
1899:
1894:
1891:
1879:
1875:
1871:
1867:
1863:
1859:
1853:
1851:
1849:
1845:
1839:
1834:
1830:
1826:
1825:
1820:
1818:
1817:Quelea quelea
1812:
1806:
1804:
1800:
1794:
1789:
1785:
1778:
1773:
1770:
1763:
1758:
1755:
1751:
1744:
1739:
1736:
1732:
1725:
1720:
1716:
1709:
1704:
1699:
1697:
1695:
1691:
1686:
1683:
1680:
1674:
1672:
1664:
1662:
1660:
1659:coconut fibre
1656:
1652:
1647:
1639:
1637:
1635:
1634:IUCN Red List
1631:
1630:least concern
1627:
1623:
1622:
1617:
1616:
1611:
1607:
1603:
1599:
1597:
1593:
1592:
1586:
1582:
1578:
1573:
1570:
1565:
1561:
1553:
1551:
1549:
1548:
1543:
1542:
1537:
1536:
1531:
1530:
1525:
1521:
1516:
1512:
1510:
1506:
1502:
1501:marabou stork
1498:
1494:
1493:lanner falcon
1490:
1486:
1482:
1478:
1469:
1462:
1460:
1456:
1454:
1450:
1446:
1442:
1438:
1434:
1433:Avena aestiva
1430:
1426:
1422:
1418:
1414:
1410:
1406:
1402:
1398:
1394:
1390:
1386:
1382:
1378:
1374:
1370:
1366:
1365:
1360:
1359:
1354:
1353:
1352:Oryza barthii
1348:
1343:
1341:
1340:
1335:
1334:
1329:
1328:
1323:
1322:
1317:
1316:
1311:
1310:
1298:respectively.
1297:
1293:
1286:
1275:
1263:
1261:
1259:
1254:
1249:
1247:
1243:
1238:
1234:
1231:
1226:
1224:
1220:
1219:
1214:
1213:
1208:
1207:
1197:
1192:
1185:
1183:
1181:
1177:
1176:Passer luteus
1173:
1169:
1168:
1163:
1162:
1156:
1152:
1147:
1145:
1141:
1136:
1128:
1126:
1124:
1120:
1115:
1112:
1107:
1104:
1098:
1094:
1092:
1088:
1083:
1079:
1075:
1071:
1067:
1063:
1055:
1053:
1051:
1039:
1038:on xeno-canto
1037:
1036:Quelea quelea
1029:
1025:
1023:
1021:
1016:
1015:
1008:
1006:
1003:
998:
995:
991:
987:
982:
980:
976:
971:
966:
963:
961:
957:
952:
947:
940:
938:
936:
935:Interbreeding
927:
926:
918:
917:
914:
913:
912:
904:
903:
900:
899:
891:
890:
887:
886:
885:
877:
876:
873:
872:
864:
863:
855:
854:
851:
850:
849:
842:
841:
838:
837:
834:
833:
832:
825:
824:
818:
817:
814:
813:
810:
809:
808:
801:
800:
794:
793:
788:
787:
781:
780:
774:
773:
769:
767:
763:
762:
757:
753:
752:
747:
743:
739:
735:
732:, namely the
731:
730:
725:
721:
713:
711:
709:
704:
702:
698:
694:
690:
686:
681:
679:
675:
671:
667:
663:
659:
655:
651:
647:
643:
639:
635:
631:
627:
623:
622:Quelea quelea
619:
611:
609:
607:
603:
599:
595:
591:
586:
582:
574:
570:
566:
562:
559:
556:
552:
548:
544:
541:
538:
534:
530:
529:
528:
525:
523:
519:
515:
511:
507:
504:
500:
496:
492:
488:
484:
480:
475:
473:
469:
465:
461:
457:
456:
451:
450:type locality
446:
444:
440:
436:
432:
431:binomial name
428:
427:
422:
418:
417:
412:
408:
404:
396:
394:
392:
391:IUCN Red List
388:
387:least concern
384:
379:
375:
370:
367:
363:
359:
355:
351:
347:
343:
339:
335:
330:
328:
324:
320:
316:
312:
308:
307:
306:Quelea quelea
300:
270:
259:
258:Quelea russii
256:
251:
248:
247:
245:
242:
238:
228:
224:
219:
214:
210:
204:
202:
201:Quelea quelea
196:
193:
192:Binomial name
189:
185:
184:
179:
176:
175:
172:
171:
167:
164:
163:
160:
157:
154:
153:
150:
149:Passeriformes
147:
144:
143:
140:
137:
134:
133:
130:
127:
124:
123:
120:
117:
114:
113:
110:
107:
104:
103:
98:
93:
89:
83:
78:
77:Least Concern
67:
63:
58:
53:
49:
44:
41:
36:
32:
27:
22:
19:
4615:
4590:A. albifrons
4589:
4577:
4566:
4556:
4546:
4537:E. hartlaubi
4536:
4526:
4516:
4506:
4497:E. axillaris
4496:
4486:
4476:
4466:
4456:
4446:
4436:
4426:
4423:Black bishop
4416:
4406:
4392:
4382:F. flavicans
4381:
4371:
4361:
4351:
4342:F. aldabrana
4341:
4338:Aldabra fody
4331:
4328:Comoros fody
4321:
4307:
4296:
4292:
4287:Q. erythrops
4286:
4276:
4262:
4251:
4239:
4229:A. rubriceps
4228:
4216:
4205:
4195:
4185:
4175:
4165:
4155:
4146:M. racheliae
4145:
4136:M. ballmanni
4135:
4132:Gola malimbe
4125:
4116:M. coronatus
4115:
4101:
4090:
4079:Pachyphantes
4078:
4067:
4057:
4047:
4037:
4027:
4017:
4007:
3997:
3987:
3977:
3967:
3957:
3947:
3937:
3927:
3917:
3908:P. albinucha
3907:
3897:
3887:
3877:
3867:
3857:
3847:
3837:
3827:
3817:
3807:
3797:
3794:Fox's weaver
3787:
3777:
3774:Giant weaver
3767:
3758:P. reichardi
3757:
3747:
3737:
3727:
3717:
3707:
3697:
3687:
3677:
3668:P. castanops
3667:
3657:
3647:
3637:
3627:
3618:P. aurantius
3617:
3607:
3597:
3588:P. subaureus
3587:
3577:
3567:
3557:
3547:
3537:
3527:
3517:
3507:
3497:
3487:
3477:
3467:
3458:P. bertrandi
3457:
3447:
3437:
3427:
3413:
3402:
3390:
3379:
3369:
3355:
3345:H. ruficauda
3344:
3332:
3321:
3311:
3301:
3291:
3277:
3266:
3257:S. frontalis
3256:
3242:
3232:D. dinemelli
3231:
3219:
3208:
3198:
3184:
3174:
3166:
3156:
3112:
3100:
3078:the original
3068:
3048:
3041:
3032:
3019:
2994:
2990:
2984:
2951:
2947:
2941:
2932:
2928:
2919:
2915:
2907:
2898:
2894:
2881:
2864:
2860:
2856:
2838:
2832:
2824:
2804:
2796:
2771:
2767:
2754:
2737:
2733:
2727:
2718:
2714:
2704:
2684:
2677:
2664:
2660:
2629:
2574:
2570:
2557:
2545:. Retrieved
2541:
2532:
2520:. Retrieved
2516:
2506:
2473:
2469:
2462:
2450:. Retrieved
2445:
2432:
2423:
2414:
2402:. Retrieved
2397:
2384:
2365:
2352:
2328:
2321:
2309:. Retrieved
2305:
2296:
2276:
2269:
2236:
2232:
2223:
2215:
2205:
2170:
2166:
2156:
2144:. Retrieved
2139:
2109:(1): 34–40.
2106:
2102:
2098:
2075:. Retrieved
2061:
1987:
1980:
1962:
1937:. Retrieved
1934:Weaver Watch
1933:
1901:
1893:
1881:. Retrieved
1861:
1828:
1822:
1816:
1783:
1769:South Africa
1749:
1730:
1675:
1668:
1643:
1619:
1613:
1600:
1598:, Tanzania.
1589:
1574:
1557:
1545:
1541:Haemoproteus
1539:
1533:
1527:
1513:
1474:
1457:
1448:
1440:
1432:
1424:
1417:Oryza sativa
1416:
1408:
1400:
1392:
1384:
1376:
1362:
1356:
1350:
1344:
1337:
1331:
1325:
1319:
1313:
1307:
1304:
1250:
1239:
1235:
1227:
1216:
1210:
1204:
1201:
1195:
1175:
1165:
1159:
1154:
1148:
1132:
1118:
1116:
1110:
1108:
1103:Benoue River
1099:
1095:
1078:South Africa
1059:
1049:
1046:
1035:
1027:
1017:
999:
993:
989:
985:
983:
967:
964:
948:
944:
933:
909:
907:
882:
880:
847:
845:
830:
828:
806:
805:
804:
790:
765:
759:
749:
745:
737:
727:
724:sister group
720:DNA analysis
717:
705:
692:
684:
682:
680:from Egypt.
673:
665:
657:
649:
641:
633:
625:
621:
617:
615:
605:
593:
589:
580:
578:
572:
568:
560:
554:
550:
547:Andrew Smith
542:
536:
535:subspecies,
526:
513:
505:
498:
495:true bunting
476:
471:
467:
453:
447:
438:
434:
424:
414:
411:10th edition
400:
371:
361:
357:
353:
345:
331:
325:, native to
314:
310:
305:
304:
268:
266:
257:
249:
200:
198:
182:
181:
169:
39:
18:
4792:iNaturalist
4640:Wikispecies
4578:Amblyospiza
4567:E. jacksoni
4507:E. macroura
4487:E. capensis
4427:E. gierowii
4348:Forest fody
4156:M. scutatus
4068:P. flavipes
4038:P. insignis
3944:Baya weaver
3928:P. sakalava
3898:P. tricolor
3858:P. jacksoni
3844:Juba weaver
3738:P. katangae
3698:P. heuglini
3678:P. burnieri
3644:Ruvu weaver
3598:P. xanthops
3578:P. capensis
3574:Cape weaver
3508:P. ocularis
3488:P. luteolus
3468:P. pelzelni
3391:Philetairus
3380:P. cabanisi
3279:Plocepasser
2935:(2): 59–60.
2895:Lanioturdus
2438:Gill, Frank
2173:: 394–413.
1713:Etching by
1577:Sara people
1497:tawny eagle
1479:, monkeys,
1309:Echinochloa
1135:specialised
1074:rainforests
1000:The female
941:Description
789:genus
689:John Latham
518:Otto Finsch
489:. In 1850,
4931:Categories
4909:Xeno-canto
4252:B. anomala
4240:Brachycope
4217:Anaplectes
4126:M. cassini
4018:P. nicolli
3998:P. preussi
3988:P. bicolor
3828:P. golandi
3778:P. grandis
3728:P. velatus
3688:P. galbula
3558:P. alienus
3370:P. arnaudi
3333:Histurgops
3244:Sporopipes
3220:Dinemellia
3186:Bubalornis
2542:Xeno-canto
2142:(in Dutch)
1989:vulgaribus
1795:References
1655:cuttlebone
1640:Aviculture
1628:listed as
1547:Plasmodium
1333:Tetrapogon
1253:incubation
1230:Malilangwe
1223:sugar cane
1180:estrildids
1111:aethiopica
1087:Lake Ngami
1034:Listen to
740:) and the
693:aethiopica
678:the Exodus
606:aethiopica
573:aethiopica
350:subspecies
4557:E. progne
4527:E. ardens
4477:E. aureus
4394:Euplectes
4352:F. omissa
4297:Q. quelea
4186:M. nitens
3938:P. manyar
3868:P. badius
3818:P. weynsi
3788:P. spekei
3748:P. ruweti
3628:P. bojeri
3528:P. batesi
3403:P. socius
3292:P. mahali
3158:Ploceidae
2715:Zoologica
2547:15 August
2498:205841906
2476:: 21–32.
2404:1 January
1878:216311213
1651:mealworms
1569:N'Djamena
1560:Lake Chad
1481:mongooses
1445:sunflower
1437:buckwheat
1347:Lake Chad
1292:Kgalagadi
1246:condition
1167:Euplectes
1048:warbling
807:Q. quelea
714:Phylogeny
666:ndzheyana
650:inyonyane
642:chimokoto
522:Karl Russ
323:Ploceidae
177:Species:
159:Ploceidae
115:Kingdom:
109:Eukaryota
4849:22719128
4810:11240909
4696:22719128
4691:BirdLife
4680:BioLib:
4625:Wikidata
4362:F. rubra
4318:Red fody
4103:Malimbus
3209:B. niger
2976:84053588
2740:: 1–25.
2688:. IUCN.
2599:12535086
2490:28012957
2452:16 April
2364:(2002).
2261:84764540
1960:(1758).
1898:"Quelea"
1813:(2016).
1788:Ethiopia
1685:fenthion
1564:Hadjerai
1453:feedlots
1425:Triticum
1371:such as
1339:Urochloa
1196:lathamii
1186:Breeding
1140:strategy
685:lathamii
638:Kwangali
555:lathamii
533:nominate
426:Emberiza
407:Linnaeus
378:avicides
348:. Three
334:Linnaeus
241:Synonyms
209:Linnaeus
155:Family:
129:Chordata
125:Phylum:
119:Animalia
105:Domain:
82:IUCN 3.1
4771:2493987
4732:rebque1
4706:rebque1
4668:Avibase
4631:Q367327
4457:E. orix
4407:E. afer
3415:Ploceus
3176:Species
3011:3808656
2956:Bibcode
2788:4082640
2768:The Auk
2607:1191425
2579:Bibcode
2306:Avibase
2241:Bibcode
2233:Ostrich
2187:1797576
1968:Holmiae
1782:female
1700:Gallery
1682:avicide
1632:on the
1477:galagos
1389:sorghum
1369:cereals
1327:Sorghum
1321:Setaria
1315:Panicum
1264:Feeding
1258:protein
1242:plumage
1161:Ploceus
1151:lowveld
1082:Réunion
975:coverts
911:Ploceus
761:Ploceus
668:in the
662:Sesotho
654:Siswati
634:enzunge
630:Swahili
602:synonym
421:bunting
413:of his
338:bunting
165:Genus:
145:Order:
135:Class:
80: (
4962:Quelea
4875:187445
4836:562669
4753:EURING
4745:QUELQU
4309:Foudia
4264:Quelea
3153:Family
3056:
3009:
2974:
2817:
2786:
2692:
2605:
2597:
2522:15 May
2496:
2488:
2372:
2340:
2284:
2259:
2185:
2146:11 May
1883:11 May
1876:
1754:Uganda
1735:Uganda
1585:Musgum
1524:Etosha
1503:. The
1489:civets
1485:genets
1443:) and
1405:millet
1373:barley
1296:Etosha
1155:Acacia
884:Foudia
881:genus
792:Quelea
766:Quelea
751:Foudia
729:Quelea
674:qualea
618:quelea
499:Quelea
487:Angola
483:Brazil
423:genus
374:locust
346:Quelea
340:, but
232:
170:Quelea
4901:95306
4888:78067
4823:66441
4805:IRMNG
4797:14005
4758:20240
4727:eBird
4719:4R42L
4703:BOW:
4683:30724
3168:Genus
3029:(PDF)
3007:JSTOR
2972:S2CID
2925:(PDF)
2891:(PDF)
2784:JSTOR
2764:(PDF)
2669:(PDF)
2634:(PDF)
2603:S2CID
2567:(PDF)
2494:S2CID
2394:(PDF)
2311:5 May
2257:S2CID
2229:(PDF)
2183:JSTOR
2077:4 May
1939:4 May
1874:S2CID
1748:male
1729:male
1602:Guano
1507:is a
1421:wheat
1397:manna
1091:roost
1066:Sahel
1009:Sound
979:moult
951:lores
658:thaha
646:Shona
510:morph
460:cited
4870:NCBI
4844:IUCN
4831:ITIS
4766:GBIF
4740:EPPO
3054:ISBN
2861:Ibis
2815:ISBN
2734:Ibis
2690:ISBN
2595:PMID
2549:2017
2524:2017
2486:PMID
2454:2017
2406:2013
2370:ISBN
2338:ISBN
2313:2017
2282:ISBN
2148:2017
2103:Ibis
2079:2017
1941:2017
1885:2017
1829:2016
1618:and
1583:and
1581:Masa
1575:The
1544:and
1499:and
1429:oats
1413:rice
1381:teff
1336:and
1294:and
1072:and
970:down
960:iris
697:type
664:and
531:The
319:bird
267:The
213:1758
139:Aves
4857:NBN
4818:ISC
4779:IBC
4714:CoL
4655:ADW
4340:, (
2999:doi
2964:doi
2869:doi
2865:107
2776:doi
2742:doi
2738:102
2587:doi
2478:doi
2474:109
2334:328
2249:doi
2175:doi
2111:doi
2107:108
2101:".
1866:doi
1833:doi
1522:in
1427:),
1419:),
1411:),
1403:),
1395:),
1387:),
1379:),
1355:),
1228:At
660:in
652:in
644:in
636:in
628:in
516:by
485:or
433:of
313:or
4933::
4911::
4898::
4885::
4872::
4859::
4846::
4833::
4820::
4807::
4794::
4781::
4768::
4755::
4742::
4729::
4716::
4693::
4670::
4657::
4642::
4627::
3155::
3111:.
3005:.
2995:46
2993:.
2970:.
2962:.
2952:71
2950:.
2933:15
2931:.
2927:.
2899:41
2897:.
2893:.
2863:.
2847:^
2823:.
2813:.
2811:96
2782:.
2772:79
2770:.
2766:.
2736:.
2717:.
2713:.
2642:^
2615:^
2601:.
2593:.
2585:.
2575:12
2573:.
2569:.
2540:.
2515:.
2492:.
2484:.
2472:.
2444:.
2422:.
2396:.
2360:;
2336:.
2304:.
2255:.
2247:.
2237:73
2235:.
2231:.
2195:^
2181:.
2169:.
2165:.
2138:.
2123:^
2105:.
2087:^
2070:,
2066:.
2060:.
1998:^
1949:^
1932:.
1911:^
1900:.
1872:.
1864:.
1860:.
1847:^
1827:.
1821:.
1802:^
1673:.
1636:.
1550:.
1495:,
1487:,
1483:,
1342:.
1330:,
1324:,
1318:,
1312:,
1093:.
1052:.
656:,
648:,
640:,
632:,
620:.
608:.
524:.
393:.
329:.
303:;
287:iː
211:,
4592:)
4588:(
4569:)
4565:(
4559:)
4555:(
4549:)
4545:(
4539:)
4535:(
4529:)
4525:(
4519:)
4515:(
4509:)
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