1478:
31:
53:
202:
and swivel their heads to cut off pieces of food. Adults do not make burrows, instead just squeezing between the cell layers of the decomposed wood, generally leaving no visible trace of their passage, while larvae live in short tunnels.
154:
in the past, and subsequent DNA analysis then placed it within
Carabidae, where it was sometimes treated as the tribe Rhysodini, but the most recent analyses place it as a subfamily in a clade along with subfamilies
183:
are deeply grooved lengthwise, thus giving these beetles their common name. The head is also grooved, and posteriorly constricted into a short but visible "neck". The 11-segment
1282:"Phylogenomic analyses clarify the pattern of evolution of Adephaga (Coleoptera) and highlight phylogenetic artefacts due to model misspecification and excessive data trimming"
1280:
Vasilikopoulos, A.; Balke, M.; Kukowka, S.; Pflug, J.M.; Martin, S.; Meusemann, K.; Hendrich, L.; Mayer, C.; Maddison, D.R.; Niehuis, O.; Beutel, R.G.; Misof, B. (2021).
1440:
129:
1519:
1192:"The beetle tree of life reveals that Coleoptera survived endâPermian mass extinction to diversify during the Cretaceous terrestrial revolution"
150:. There are 19 genera and at least 380 described species in Rhysodinae. The group of genera making up Rhysodinae had been treated as the family
1477:
30:
1548:
1512:
830:
175:
These beetles are elongate, in size ranging from 5â8 mm, and color ranging from a reddish brown to black. Both the
1362:
188:
1406:
52:
198:, which are believed to be their diet. Instead of using their mandibles to bite, they use the anterior edge of the
1445:
1553:
1419:
1505:
871:
1315:
1056:
39:
952:
1135:
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011).
1388:
1538:
47:
1543:
1453:
1375:
1262:
1190:
Mckenna, Duane D.; Wild, Alexander L.; Kanda, Kojun; Bellamy, Charles L.; et al. (2015).
1172:
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965:
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1458:
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lack cutting edges and are thus nonfunctional. The front legs are short and strongly built.
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They occur on all continents with forested areas, the richest fauna being found in
164:
1367:
1427:
1347:
960:
907:
879:
838:
215:
195:
1338:
233:
Position in relation to the
Carabidae according to Vasilikopoulos et al., 2021
207:
160:
1247:
1158:
1235:"Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico"
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925:
889:
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156:
147:
84:
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Adults and larvae live in moist rotten wood that is infested with
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1115:
1493:
1322:
187:are short, resembling a string of beads, while the
1513:
8:
1137:"Family-group names in Coleoptera (Insecta)"
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20:
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246:
239:
1098:"Clinidium. Version 01 March 2000"
14:
1143:(88). Pensoft Publishers: 1â972.
142:is a subfamily (sometimes called
1476:
51:
1:
1492:. You can help Knowledge by
167:to the remaining Carabidae.
1570:
1471:
1077:R.T. & J.R.Bell, 1991
1066:R.T. & J.R.Bell, 1991
1046:R.T. & J.R.Bell, 1982
1036:R.T. & J.R.Bell, 1978
1018:R.T. & J.R.Bell, 1979
1008:R.T. & J.R.Bell, 1978
998:R.T. & J.R.Bell, 1978
978:R.T. & J.R.Bell, 1978
937:R.T. & J.R.Bell, 1978
930:R.T. & J.R.Bell, 1987
919:R.T. & J.R.Bell, 1987
912:R.T. & J.R.Bell, 1978
901:R.T. & J.R.Bell, 1978
894:R.T. & J.R.Bell, 1982
884:R.T. & J.R.Bell, 1978
860:R.T. & J.R.Bell, 1978
843:R.T. & J.R.Bell, 1978
819:R.T. & J.R.Bell, 1978
730:
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439:
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244:
48:Scientific classification
46:
37:
28:
23:
1248:10.3897/zookeys.245.3416
1241:(245). Pensoft: 1â1722.
498:⊞other subfamilies
1233:Bousquet, Yves (2012).
1488:-related article is a
1150:10.3897/zookeys.88.807
1064:Tribe Sloanoglymmiini
40:Omoglymmius americanus
1549:Carabidae subfamilies
1196:Systematic Entomology
144:wrinkled bark beetles
1415:Fauna Europaea (new)
1102:Tree of Life Project
16:Subfamily of beetles
935:Tribe Omoglymmiini
899:Tribe Leoglymmiini
1299:10.1111/syen.12508
1209:10.1111/syen.12132
1501:
1500:
1469:
1468:
1454:Open Tree of Life
1316:Taxon identifiers
1078:
1067:
1060:
1047:
1037:
1026:
1019:
1009:
999:
989:
979:
969:
956:
938:
931:
920:
917:Tribe Medisorini
913:
902:
895:
885:
875:
861:
854:
844:
834:
820:
817:Tribe Clinidiini
815:
814:
810:
809:
801:
800:
792:
791:
784:
783:
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767:
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758:
750:
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469:
401:
400:
385:
384:
376:
375:
367:
366:
358:
357:
137:
136:
133:
1561:
1554:Rhysodinae stubs
1522:
1515:
1508:
1480:
1473:
1462:
1461:
1449:
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1436:
1435:
1423:
1422:
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1397:
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1187:
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1125:
1123:
1122:
1112:
1106:
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1093:
1076:
1065:
1055:
1045:
1035:
1024:
1023:Tribe Rhysodini
1017:
1007:
997:
987:
977:
964:
947:
936:
929:
918:
911:
900:
893:
883:
870:
859:
858:Tribe Dhysorini
852:
842:
829:
818:
778:
713:
693:
673:
613:
571:
564:
522:
515:
508:
492:
491:
442:
422:
415:
395:
330:
310:
290:
270:
254:
253:
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230:
229:
146:) in the family
128:
56:
55:
33:
21:
1569:
1568:
1564:
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1529:
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1470:
1465:
1457:
1452:
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1439:
1431:
1426:
1418:
1413:
1405:
1400:
1392:
1387:
1379:
1374:
1366:
1361:
1352:
1351:
1346:
1337:
1336:
1331:
1318:
1308:
1307:
1279:
1278:
1274:
1232:
1231:
1227:
1189:
1188:
1184:
1134:
1133:
1129:
1120:
1118:
1114:
1113:
1109:
1095:
1094:
1090:
1085:
988:Grouvelle, 1903
853:Fairmaire, 1895
811:
802:
793:
780:
779:
776:
771:
770:
769:
760:
751:
742:
660:
651:
642:
600:
551:
501:
500:
499:
480:
471:
397:
396:
393:
388:
387:
386:
377:
368:
359:
263:
262:
261:
228:
218:, and northern
173:
127:
50:
17:
12:
11:
5:
1567:
1565:
1557:
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1546:
1541:
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1525:
1524:
1517:
1510:
1502:
1499:
1498:
1481:
1467:
1466:
1464:
1463:
1450:
1437:
1424:
1411:
1402:Fauna Europaea
1398:
1385:
1372:
1359:
1344:
1328:
1326:
1320:
1319:
1314:
1306:
1305:
1272:
1225:
1202:(4): 835â880.
1182:
1127:
1107:
1096:Bell, Ross T.
1087:
1086:
1084:
1081:
1080:
1079:
1073:Sloanoglymmius
1062:
1061:
1048:
1038:
1021:
1020:
1010:
1000:
990:
980:
974:Plesioglymmius
970:
957:
933:
932:
915:
914:
897:
896:
886:
876:
856:
855:
845:
835:
813:
812:
808:
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804:
803:
799:
798:
795:
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786:
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772:
766:
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748:
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721:
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333:
328:
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296:Megacephalini
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279:
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273:
268:
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259:
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172:
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135:
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68:
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44:
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26:
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15:
13:
10:
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6:
4:
3:
2:
1566:
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1369:
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1360:
1355:
1349:
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1334:
1330:
1329:
1327:
1325:
1321:
1317:
1312:
1300:
1295:
1291:
1287:
1286:Syst. Entomol
1283:
1276:
1273:
1268:
1264:
1259:
1254:
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1240:
1236:
1229:
1226:
1220:
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1197:
1193:
1186:
1183:
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1169:
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1128:
1117:
1111:
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1103:
1099:
1092:
1089:
1082:
1075:
1074:
1070:
1069:
1068:
1058:
1054:
1053:
1049:
1044:
1043:
1039:
1034:
1033:
1029:
1028:
1027:
1025:Laporte, 1840
1016:
1015:
1011:
1006:
1005:
1001:
996:
995:
991:
986:
985:
981:
976:
975:
971:
967:
963:
962:
958:
954:
950:
946:
945:
941:
940:
939:
928:
927:
923:
922:
921:
910:
909:
905:
904:
903:
892:
891:
887:
882:
881:
877:
873:
869:
868:
864:
863:
862:
851:
850:
846:
841:
840:
836:
832:
828:
827:
823:
822:
821:
806:
805:
797:
796:
788:
787:
773:
764:
763:
755:
754:
746:
745:
737:
736:
733:
728:
727:
724:
723:
720:
715:
714:
708:
707:
704:
703:
700:
695:
694:
688:
687:
684:
683:
680:
675:
674:
668:
667:
664:
663:
655:
654:
646:
645:
637:
636:
633:
632:Omophroninae
628:
627:
624:
623:
620:
615:
614:
608:
607:
604:
603:
595:
594:
591:
586:
585:
582:
581:
578:
573:
572:
566:
565:
559:
558:
555:
554:
546:
545:
542:
537:
536:
533:
532:
529:
524:
523:
517:
516:
510:
509:
503:
494:
493:
488:
487:
484:
483:
475:
474:
466:
465:
462:
457:
456:
453:
452:
449:
444:
443:
437:
436:
433:
432:
429:
424:
423:
417:
416:
409:
408:
405:
404:
390:
381:
380:
372:
371:
363:
362:
354:
353:
350:
345:
344:
341:
340:
337:
332:
331:
325:
324:
321:
320:
317:
312:
311:
305:
304:
301:
300:
297:
292:
291:
285:
284:
281:
280:
277:
272:
271:
265:
256:
255:
250:Cicindelidae
249:
248:
242:
241:
237:
236:
232:
231:
225:
223:
221:
220:South America
217:
213:
209:
204:
201:
197:
192:
190:
186:
182:
178:
170:
168:
166:
162:
158:
153:
149:
145:
141:
131:
126:
123:
120:
119:
116:
113:
110:
109:
106:
103:
100:
99:
96:
93:
90:
89:
86:
83:
80:
79:
76:
73:
70:
69:
66:
63:
60:
59:
54:
49:
45:
42:
41:
36:
32:
27:
22:
19:
1494:expanding it
1485:
1483:
1323:
1292:: 991â1018.
1289:
1285:
1275:
1238:
1228:
1199:
1195:
1185:
1140:
1130:
1119:. Retrieved
1110:
1101:
1091:
1071:
1063:
1050:
1040:
1030:
1022:
1012:
1002:
992:
982:
972:
959:
942:
934:
924:
916:
906:
898:
888:
878:
865:
857:
849:Rhyzodiastes
847:
837:
824:
816:
731:
719:Brachininae
718:
698:
678:
631:
619:Loricerinae
618:
589:
576:
540:
527:
460:
447:
427:
349:Oxycheilini
348:
336:Cicindelini
335:
316:Collyridini
315:
295:
276:Manticorini
275:
205:
196:slime moulds
193:
174:
163:, forming a
151:
143:
139:
138:
124:
38:
18:
1428:iNaturalist
1348:Wikispecies
1219:10057/11540
961:Omoglymmius
908:Leoglymmius
880:Neodhysores
839:Grouvellina
732:Harpalinae
679:Scaritinae
590:Elaphrinae
461:Siagoninae
428:Rhysodinae
216:Philippines
171:Description
121:Subfamily:
24:Rhysodinae
1539:Rhysodinae
1533:Categories
1486:Rhysodinae
1368:Rhysodinae
1354:Rhysodinae
1324:Rhysodinae
1121:2018-11-24
1083:References
966:Ganglbauer
699:Trechinae
577:Broscinae
541:Nebriinae
528:Carabinae
448:Paussinae
410:Carabidae
208:New Guinea
161:Siagoninae
152:Rhysodidae
140:Rhysodinae
125:Rhysodinae
105:Coleoptera
85:Arthropoda
1159:1313-2989
953:J.R. Bell
949:R.T. Bell
926:Medisores
890:Tangarona
872:Grouvelle
826:Clinidium
212:Indonesia
189:mandibles
157:Paussinae
148:Carabidae
115:Carabidae
71:Kingdom:
65:Eukaryota
1544:Adephaga
1339:Q2784682
1333:Wikidata
1267:23431087
1177:21594053
1052:Rhysodes
1032:Kaveinga
1014:Yamatosa
1004:Xhosores
984:Shyrodes
944:Arrowina
867:Dhysores
185:antennae
179:and the
111:Family:
81:Phylum:
75:Animalia
61:Domain:
1258:3577090
1239:ZooKeys
1168:3088472
1141:ZooKeys
994:Srimara
777:⊟
394:⊟
260:⊞
130:Laporte
101:Order:
95:Insecta
91:Class:
1459:710477
1433:423043
1407:385498
1265:
1255:
1175:
1165:
1157:
1116:"GBIF"
1059:, 1822
1057:Germar
1042:Kupeus
968:, 1891
955:, 1978
951:&
874:, 1903
833:, 1830
226:Genera
214:, the
200:mentum
181:elytra
177:thorax
165:sister
132:, 1840
1484:This
1446:63897
1381:8LTM4
831:Kirby
1490:stub
1441:NCBI
1394:8610
1263:PMID
1173:PMID
1155:ISSN
159:and
1389:EoL
1376:CoL
1363:AFD
1294:doi
1253:PMC
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1214:hdl
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1163:PMC
1145:doi
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1290:46
1288:.
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1200:40
1198:.
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1521:e
1514:t
1507:v
1496:.
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1222:.
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1179:.
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