42:
786:
858:
886:
910:
898:
742:; their diet consists of a wide variety of plants including forbs, thistles, grasses, and succulents. They consume berries and other fruits when available. They sometimes gnaw on bones or even ash and hyena feces to obtain calcium, necessary for bone development and their eggshells. Seeds can pass undigested through the gut, so the leopard tortoise plays a significant role in seed dispersal. Normally active during the day, they are less active during hot weather or during the dry season.
746:
mating, the male makes grunting vocalizations. Nesting occurs between May and
October when the female digs a hole and lays a clutch of five to 30 eggs. As many as five to seven clutches may be laid in a single season. Incubation takes 8–15 months depending on temperature. The numerous predators of the eggs and hatchlings include monitor lizards, snakes, jackals, and crows. Adults have few natural predators, but lions and hyenas have occasionally been reported preying on them.
663:
810:
115:
822:
834:
874:
774:
798:
846:
73:
1530:
716:
678:
is high and domed with steep, almost vertical sides. Juveniles and young adults are attractively marked with black blotches, spots, or even dashes and stripes on a yellow background. In mature adults, the markings tend to fade to a nondescript brown or grey. The head and limbs are uniformly colored
754:
The leopard tortoise is a widespread species and remains common throughout most of its range. Human activities, including agricultural burning, consumption, and especially commercial exploitation in the pet trade, are potential threats, but have not yet caused significant population declines. They
745:
The leopard tortoise reaches sexual maturity between 12 and 15 years old, and may live as long as 80 to 100 years. During the mating season, males fight over females, ramming and butting their competitors. They trail after females for quite some distance, often ramming them into submission. When
670:
The leopard tortoise is the fourth-largest species of tortoise in the world, with typical adults reaching 40 cm (16 in) and weighing 13 kg (29 lb). Adults tend to be larger in the northern and southern ends of their range, where typical specimens weigh up to 20 kg
724:
707:. Over this range, the leopard tortoise occupies the most varied habitats of any African tortoise, including grasslands, thorn-scrub, mesic brushland, and savannas. They can be found at altitudes ranging from sea level to 2,900 m (9,500 ft).
729:
728:
725:
730:
755:
are increasingly being bred in captivity for the pet trade. For example, most tortoises exported from Kenya and
Tanzania originate in captive-breeding programs, alleviating collection from the wild.
585:
burrows. The leopard tortoise does not dig other than to make nests in which to lay eggs. Given its propensity for grassland habitats, it grazes extensively upon mixed grasses. It also favors
727:
4903:
581:. This tortoise is a grazing species that favors semiarid, thorny to grassland habitats. In both very hot and very cold weather, it may dwell in abandoned fox, jackal, or
4965:
4833:
5023:
4872:
4993:
726:
758:
The leopard tortoise has been listed in
Appendix II of CITES since 1975, and in 2000, the United States banned their import because of the risk posed by
4952:
4807:
885:
909:
809:
4846:
1775:
1964:
1958:
1431:
1035:
Conservation
Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group
857:
1176:
785:
5008:
2535:
1328:(1935). "Scientific Results of an Expedition to Rain Forest Regions in Eastern Africa. I. New Reptiles and Amphibians from East Africa".
1069:
Fritz, U.; Bininda-Emonds, O. R. P. (2007-07-03). "When genes meet nomenclature: Tortoise phylogeny and the shifting generic concepts of
5018:
1050:
1301:
1253:
897:
41:
1586:
797:
1817:
1028:"Turtles of the world, 7th edition: annotated checklist of taxonomy, synonymy, distribution with maps, and conservation status"
4568:
3651:
1946:
1940:
1812:
1571:
1027:
4851:
4695:
Turtles of the World 2017 Update: Annotated
Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status
2504:
turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status
2200:
2015:
833:
601:
The phylogenic placement of the leopard tortoise has been subject to several revisions. Different authors have placed it in
1351:
873:
5013:
2185:
2175:
1837:
1037:. Chelonian Research Monographs. Vol. 5. IUCN/SSC Tortoise and Freshwater Turtle Specialist Group. pp. 329–479.
4545:
4693:
2503:
671:(44 lb), and an exceptionally large tortoise may reach 70 cm (28 in) and weigh 40 kg (88 lb).
114:
4768:
4704:
3227:
2611:
2325:
2190:
1842:
773:
350:
1579:
389:
5003:
4877:
2960:
2180:
1786:
1424:
821:
4759:
2195:
1952:
1603:
3606:
2528:
2106:
1827:
1796:
1765:
1653:
687:
They are widely distributed across the arid and savanna regions of eastern and southern Africa, extending from
1862:
1735:
432:
4462:
2475:
2025:
1874:
1868:
1272:
845:
644:
436:
246:
2422:
2416:
2374:
4721:
4434:
3234:
3192:
2832:
2434:
2380:
2074:
2056:
1832:
1770:
1729:
1705:
1699:
1671:
1665:
1659:
1634:
1564:
376:
4773:
2410:
2080:
2062:
762:, an infectious disease carried by tortoise ticks that could seriously impact the US livestock industry.
4998:
3178:
2680:
2491:
2485:
2446:
2428:
2392:
2279:
2267:
2236:
2068:
1982:
1915:
1822:
1802:
1717:
1595:
1417:
1404:
229:
2470:
2452:
2152:
2086:
2050:
1750:
1745:
2404:
4794:
4420:
3086:
2710:
2440:
1987:
1723:
1711:
1200:
50:
2398:
2386:
2368:
4405:
2764:
2521:
2480:
2273:
2157:
1920:
1694:
973:
402:
62:
1853:
492:
3812:
3777:
3644:
3442:
3383:
3353:
2297:
2261:
2241:
2147:
2041:
1760:
1296:. Third Revised edition, Second impression. Sanibel Island, Florida: Ralph Curtis Books. 399 pp.
1289:
1164:
1141:
428:
255:
109:
3361:
3115:
662:
440:
4970:
4838:
4885:
4781:
4355:
3338:
3290:
3281:
3164:
2741:
2302:
2129:
2124:
1892:
1781:
1627:
1622:
1297:
1249:
1172:
1100:
1079:
1046:
337:
1195:
621:, a monotypic genus. Considerable debate has occurred about the existence of two subspecies,
4898:
4890:
4864:
4147:
3535:
3510:
3419:
3150:
2695:
2218:
1910:
1790:
1546:
1325:
1311:
1245:
1131:
1092:
1038:
968:
363:
316:
4595:
4243:
3972:
3940:
3577:
3486:
3262:
2848:
2593:
2462:
2330:
2020:
2010:
2005:
1806:
586:
471:
1383:
1042:
4490:
4480:
4377:
4227:
4067:
3769:
3714:
3456:
3391:
2931:
2922:
2891:
2799:
2688:
2602:
2317:
2097:
2030:
1689:
1215:
704:
283:
4987:
4859:
4670:
4553:
4453:
4442:
4413:
4327:
4211:
4195:
4155:
4089:
3999:
3964:
3908:
3892:
3876:
3850:
3838:
3822:
3800:
3792:
3599:
3310:
3143:
3004:
2951:
2899:
2307:
2210:
1556:
1489:
1365:
959:
950:
568:
186:
82:
77:
17:
1145:
4786:
4605:
4530:
4506:
4023:
3900:
3884:
3860:
3761:
3706:
3630:
3546:
3520:
3434:
3345:
3324:
3248:
3213:
3023:
2989:
2856:
2772:
2718:
2340:
2252:
1931:
1645:
1495:
1238:
696:
206:
54:
1318:. London: Trustees of the British Museum. (Edward Newman, printer). iv + 124 pp. (
4943:
4916:
4820:
4753:
4583:
4538:
4384:
4369:
4291:
4283:
4275:
4251:
4131:
4115:
4099:
4015:
4007:
3956:
3932:
3924:
3916:
3784:
3736:
3659:
3584:
3449:
3405:
3331:
3317:
3269:
3126:
3044:
2982:
2975:
2968:
2883:
2825:
2817:
2734:
2726:
2673:
2663:
2289:
2139:
1681:
1614:
692:
688:
654:
meaning "leopard" and refers to the leopard-like spots on the tortoise's shell.
290:
1096:
4662:
4575:
4561:
4522:
4299:
4219:
4187:
4171:
4139:
4077:
4050:
4031:
3948:
3753:
3673:
3666:
3614:
3591:
3501:
3368:
3100:
2907:
2840:
2757:
2749:
2652:
2643:
1974:
1902:
1538:
1483:
1477:
949:
Baker, P.J.; Kabigumila, J.; Leuteritz, T.; Hofmeyr, M.; Ngwava, J.M. (2015).
864:
759:
577:
415:
176:
4744:
4654:
4514:
4498:
4427:
4391:
4341:
4334:
4259:
4235:
4179:
4123:
3830:
3622:
3570:
3528:
3241:
3199:
3107:
3093:
3079:
3072:
3065:
3058:
3030:
2875:
2703:
739:
499:
126:
1104:
995:
715:
4937:
4738:
4469:
4398:
4318:
4307:
4203:
3868:
3745:
3637:
3479:
3464:
3426:
3375:
3301:
3206:
3185:
3171:
3157:
3013:
2940:
2809:
2788:
2780:
2562:
2359:
2228:
1997:
1884:
1515:
1459:
1440:
1136:
1119:
1088:
675:
582:
556:
196:
146:
4957:
4812:
4613:
4362:
4267:
4163:
4107:
3687:
3680:
3562:
3412:
3398:
3255:
3220:
3135:
3037:
2577:
2568:
2167:
2116:
1471:
1465:
700:
590:
560:
156:
4825:
4348:
3991:
2583:
2556:
2544:
1453:
166:
136:
4715:
1356:
1316:
Hand-list of the
Specimens of Shield Reptiles in the British Museum
703:. The species is generally absent from the humid forest regions of
4911:
3555:
2622:
1409:
722:
714:
661:
564:
99:
93:
3051:
1330:
Bulletin of the Museum of
Comparative Zoölogy at Harvard College
4799:
4719:
2517:
1413:
1275:(1828). "Descriptions of three new Species of Land Tortoises".
1366:"Tortoise Trust Web - Observations on Dehydration in Reptiles"
944:
942:
940:
938:
936:
934:
932:
930:
2513:
617:(2006). More recently, consensus appears to have settled on
1294:
Field Guide to Snakes and other
Reptiles of Southern Africa
1064:
1062:
1286:, new species, pp. 420–421). (in English and Latin).
1405:
IUCN/SSC Tortoise and
Freshwater Turtle Specialist Group
1190:
1188:
629:, but recent work does not support this distinction.
815:
Retracted fore legs of adult female leopard tortoise
4927:
4728:
4645:
4638:
4631:
4593:
4479:
4452:
4317:
4087:
4065:
4058:
4049:
3982:
3848:
3810:
3735:
3728:
3697:
3545:
3509:
3500:
3300:
3279:
3125:
3012:
3003:
2950:
2930:
2921:
2866:
2798:
2662:
2651:
2642:
2610:
2601:
2592:
2461:
2357:
2339:
2316:
2288:
2250:
2227:
2209:
2166:
2138:
2115:
2095:
2039:
1996:
1973:
1929:
1901:
1883:
1851:
1744:
1680:
1644:
1613:
1555:
1537:
1504:
1237:
575:, although in the past, it was commonly placed in
1171:. South Africa: Struik Publishers. p. 128.
974:10.2305/IUCN.UK.2015-4.RLTS.T163449A1009442.en
4692:Phylogenetic arrangement of turtles based on
2529:
2502:Phylogenetic arrangement of turtles based on
1425:
8:
1169:Tortoises, Terrapins & Turtles of Africa
571:. It is the only extant member of the genus
891:Leopard tortoise digging a hole to lay eggs
4716:
4642:
4635:
4062:
4055:
3732:
3506:
3009:
2927:
2659:
2648:
2616:
2607:
2598:
2536:
2522:
2514:
1508:
1432:
1418:
1410:
1244:. Smithsonian Institution Press. pp.
1236:Ernst, Carl H.; Barbour, Roger W. (1989).
915:Leopard tortoise closing hole with eggs in
71:
40:
31:
1196:"Psammobates pardalis (Leopard Tortoise)"
1135:
972:
803:Small 20-year-old leopard tortoise eating
1159:
1157:
1155:
666:Shell patterns fade in mature specimens.
1388:(Leopard Tortoise) Care – Misty Corton"
926:
769:
529:Le, Raxworthy, McCord & Mertz, 2006
1965:Saddle-backed Rodrigues giant tortoise
1959:Saddle-backed Mauritius giant tortoise
1120:"Checklist of Chelonians of the World"
1026:Turtle Taxonomy Working Group (2014).
1021:
1019:
1017:
1015:
719:Leopard tortoise eating plant material
5024:Taxa named by Thomas Bell (zoologist)
563:of eastern and southern Africa, from
555:) is a large and attractively marked
7:
4994:IUCN Red List least concern species
1043:10.3854/crm.5.000.checklist.v7.2014
960:IUCN Red List of Threatened Species
461:— Loveridge & E. Williams, 1957
25:
1118:Fritz, Uwe; Havaš, Peter (2007).
635:is a combination of Greek words:
49:On the S90 Road north of Satara,
1528:
1339:, new subspecies, pp. 4–5).
908:
896:
884:
872:
856:
844:
832:
820:
808:
796:
784:
772:
113:
1947:Domed Rodrigues giant tortoise
1941:Domed Mauritius giant tortoise
1813:Santiago Island giant tortoise
639:meaning "mark" or "point" and
516:Stigmochelys pardalis babcocki
507:Centrochelys pardalis babcocki
410:Stigmochelys pardalis pardalis
397:Centrochelys pardalis pardalis
1:
1766:Chatham Island giant tortoise
839:Three adult leopard tortoises
525:Geochelone pardalis babcockii
479:Geochelone paradalis babcocki
1833:Volcán Darwin giant tortoise
1828:Volcán Alcedo giant tortoise
1818:Pinzón Island giant tortoise
1308:, pp. 29–30 + Plate 4).
903:Leopard tortoise laying eggs
879:Three baby leopard tortoises
488:Geochelone pardalis baboocki
457:Geochelone pardalis babcocki
371:Geochelone pardalis pardalis
5009:Reptiles of Southern Africa
4705:List of Testudines families
2326:Flat-backed spider tortoise
2201:Speke's hinge-back tortoise
2191:Lobatse hinge-back tortoise
2016:Goode's thornscrub tortoise
1843:Western Santa Cruz tortoise
1823:Sierra Negra giant tortoise
1771:Eastern Santa Cruz tortoise
5040:
5019:Reptiles described in 1828
2186:Home's hinge-back tortoise
2181:Forest hinge-back tortoise
2176:Bell's hinge-back tortoise
1838:Volcán Wolf giant tortoise
1787:Hood Island giant tortoise
1776:Fernandina Island tortoise
1526:
1097:10.1016/j.zool.2007.02.003
4688:
2619:
2551:
2500:
2196:Natal hinge-back tortoise
1761:Cerro Azul giant tortoise
1604:Aldabrachelys grandidieri
1511:
1448:
1337:Testudo pardalis babcocki
448:Testudo pardalis babcocki
358:Testudo pardalis pardalis
261:
254:
235:
228:
110:Scientific classification
108:
91:
69:
60:
48:
39:
34:
2107:Hesperotestudo percrassa
1863:Cheirogaster bacharidisi
1797:Santa Fe Island tortoise
1782:Floreana Island tortoise
1736:Chelonoidis sombrerensis
1654:African spurred tortoise
1322:, new genus, p. 5).
683:Distribution and habitat
643:meaning "tortoise". The
2026:Sonoran Desert tortoise
779:One-month-old hatchling
734:Leopard tortoise eating
679:yellow, tan, or brown.
650:is from the Latin word
1953:Réunion giant tortoise
1875:Cheirogaster schafferi
1869:Cheirogaster gymnesica
1706:Chelonoidis alburyorum
1700:Yellow-footed tortoise
1672:Centrochelys vulcanica
1666:Centrochelys burchardi
1660:Centrochelys atlantica
1635:Astrochelys rogerbouri
1565:Aldabra giant tortoise
738:Leopard tortoises are
735:
720:
667:
597:Taxonomy and etymology
4760:Stigmochelys pardalis
4730:Stigmochelys pardalis
2486:Spur-thighed tortoise
2423:Stylemys nebrascensis
2417:Stylemys karakolensis
2375:Stylemys calaverensis
2280:Megalochelys sondaari
2268:Megalochelys cautleyi
2237:Asian forest tortoise
1983:Burmese star tortoise
1803:Pinta Island tortoise
1596:Aldabrachelys abrupta
1370:www.tortoisetrust.org
1352:Stigmochelys pardalis
967:: e.T163449A1009442.
953:Stigmochelys pardalis
733:
718:
665:
552:Stigmochelys pardalis
384:Stigmochelys pardalis
345:Megachersine pardalis
239:Stigmochelys pardalis
18:Stigmochelys pardalis
5014:Fauna of East Africa
2476:Kleinmann's tortoise
2435:Stylemys oregonensis
2381:Stylemys canetotiana
2075:Hadrianus schucherti
2057:Hadrianus majusculus
1988:Indian star tortoise
1730:Chelonoidis sellovii
1724:Chelonoidis monensis
1712:Chelonoidis cubensis
1357:The Reptile Database
1240:Turtles of the World
1201:Animal Diversity Web
1137:10.3897/vz.57.e30895
996:"Appendices | CITES"
863:Leopard tortoise on
851:Mature adult (faded)
711:Ecology and behavior
691:and Somalia, across
423:Psammobates pardalis
51:Kruger National Park
2481:Marginated tortoise
2411:Stylemys frizaciana
2274:Megalochelys margae
2158:Travancore tortoise
2081:Hadrianus utahensis
2063:Hadrianus octonaria
1751:Chelonoidis complex
1695:Red-footed tortoise
1386:Geochelone pardalis
1306:Geochelone pardalis
466:Geochelone babcocki
63:Conservation status
27:Species of tortoise
3813:Nanhsiungchelyidae
3778:Macrocephalochelys
2492:Testudo brevitesta
2471:Hermann's tortoise
2447:Stylemys uintensis
2429:Stylemys neglectus
2393:Stylemys conspecta
2298:Geometric tortoise
2262:Megalochelys atlas
2242:Impressed tortoise
2153:Forsten's tortoise
2148:Elongated tortoise
2069:Hadrianus robustus
1746:Galápagos tortoise
1718:Chelonoidis lutzae
1277:Zoological Journal
1216:"Leopard Tortoise"
1124:Vertebrate Zoology
827:Close-up of scutes
736:
721:
668:
375:— Loveridge &
312:Testudo bipunctata
5004:Turtles of Africa
4981:
4980:
4886:Open Tree of Life
4722:Taxon identifiers
4713:
4712:
4684:
4683:
4680:
4679:
4627:
4626:
4623:
4622:
4045:
4044:
4041:
4040:
3724:
3723:
3496:
3495:
2999:
2998:
2917:
2916:
2638:
2637:
2634:
2633:
2630:
2629:
2511:
2510:
2506:. Key: †=extinct.
2453:Stylemys undabuna
2303:Serrated tortoise
2087:Hadrianus vialovi
2051:Hadrianus corsoni
1921:Speckled padloper
1893:Angulate tortoise
1628:Radiated tortoise
1623:Angonoka tortoise
1524:
1523:
1178:978-1-77007-463-7
731:
543:
542:
530:
521:
512:
496:
484:
475:
462:
453:
444:
419:
406:
393:
380:
367:
354:
341:
320:
303:
287:
279:Testudo biguttata
275:
103:
86:
35:Leopard tortoise
16:(Redirected from
5031:
4974:
4973:
4961:
4960:
4948:
4947:
4946:
4929:Testudo pardalis
4920:
4919:
4907:
4906:
4894:
4893:
4881:
4880:
4868:
4867:
4855:
4854:
4842:
4841:
4829:
4828:
4816:
4815:
4803:
4802:
4790:
4789:
4777:
4776:
4764:
4763:
4762:
4749:
4748:
4747:
4717:
4643:
4636:
4148:Chupacabrachelys
4063:
4056:
3733:
3511:Carettochelyidae
3507:
3010:
2928:
2660:
2649:
2617:
2608:
2599:
2538:
2531:
2524:
2515:
2405:Stylemys emiliae
2349:Leopard tortoise
2219:Pancake tortoise
2130:Greater padloper
1547:Russian tortoise
1532:
1531:
1509:
1434:
1427:
1420:
1411:
1401:
1399:
1398:
1392:www.chelonia.org
1379:
1377:
1376:
1284:Testudo pardalis
1260:
1259:
1243:
1233:
1227:
1226:
1224:
1223:
1212:
1206:
1205:
1192:
1183:
1182:
1161:
1150:
1149:
1139:
1115:
1109:
1108:
1066:
1057:
1056:
1032:
1023:
1010:
1009:
1007:
1006:
992:
986:
985:
983:
981:
976:
946:
912:
900:
888:
876:
860:
848:
836:
824:
812:
800:
788:
776:
732:
567:to the southern
547:leopard tortoise
528:
519:
510:
491:
482:
469:
460:
451:
426:
413:
400:
387:
374:
361:
348:
335:
315:
301:
282:
273:
270:Testudo pardalis
241:
221:S. pardalis
118:
117:
97:
80:
75:
74:
44:
32:
21:
5039:
5038:
5034:
5033:
5032:
5030:
5029:
5028:
4984:
4983:
4982:
4977:
4969:
4964:
4956:
4951:
4942:
4941:
4936:
4923:
4915:
4910:
4902:
4897:
4889:
4884:
4876:
4871:
4863:
4858:
4850:
4845:
4837:
4832:
4824:
4819:
4811:
4806:
4798:
4793:
4785:
4780:
4772:
4767:
4758:
4757:
4752:
4743:
4742:
4737:
4724:
4714:
4709:
4676:
4619:
4596:Sahonachelyidae
4589:
4475:
4448:
4313:
4244:Phosphatochelys
4083:
4037:
3978:
3973:Terlinguachelys
3941:Pneumatoarthrus
3844:
3806:
3720:
3693:
3541:
3492:
3296:
3275:
3263:Siebenrockiella
3121:
2995:
2946:
2913:
2862:
2849:Mesodermochelys
2794:
2655:
2626:
2588:
2547:
2542:
2512:
2507:
2496:
2457:
2441:Stylemys pygmea
2353:
2335:
2331:Spider tortoise
2312:
2284:
2246:
2223:
2205:
2162:
2134:
2125:Common padloper
2111:
2091:
2035:
2021:Gopher tortoise
2011:Desert tortoise
2006:Bolson tortoise
1992:
1969:
1925:
1897:
1879:
1847:
1807:Lonesome George
1740:
1676:
1640:
1609:
1587:A. g. hololissa
1551:
1533:
1529:
1520:
1513:Species of the
1500:
1444:
1438:
1396:
1394:
1382:
1374:
1372:
1363:
1346:
1269:
1267:Further reading
1264:
1263:
1256:
1235:
1234:
1230:
1221:
1219:
1214:
1213:
1209:
1194:
1193:
1186:
1179:
1163:
1162:
1153:
1117:
1116:
1112:
1068:
1067:
1060:
1053:
1030:
1025:
1024:
1013:
1004:
1002:
994:
993:
989:
979:
977:
948:
947:
928:
923:
916:
913:
904:
901:
892:
889:
880:
877:
868:
861:
852:
849:
840:
837:
828:
825:
816:
813:
804:
801:
792:
789:
780:
777:
768:
752:
723:
713:
685:
660:
599:
539:
538:
527:
518:
509:
490:
481:
468:
459:
452:Loveridge, 1935
450:
425:
412:
399:
386:
373:
360:
347:
334:
314:
300:
281:
272:
265:
250:
243:
237:
224:
112:
104:
87:
76:
72:
65:
28:
23:
22:
15:
12:
11:
5:
5037:
5035:
5027:
5026:
5021:
5016:
5011:
5006:
5001:
4996:
4986:
4985:
4979:
4978:
4976:
4975:
4962:
4949:
4933:
4931:
4925:
4924:
4922:
4921:
4908:
4895:
4882:
4869:
4856:
4843:
4830:
4817:
4804:
4791:
4778:
4765:
4750:
4734:
4732:
4726:
4725:
4720:
4711:
4710:
4708:
4707:
4700:
4699:
4698:. † = extinct.
4689:
4686:
4685:
4682:
4681:
4678:
4677:
4675:
4674:
4666:
4658:
4649:
4647:
4640:
4633:
4629:
4628:
4625:
4624:
4621:
4620:
4618:
4617:
4609:
4600:
4598:
4591:
4590:
4588:
4587:
4579:
4572:
4565:
4557:
4549:
4542:
4534:
4526:
4518:
4510:
4502:
4494:
4491:Albertwoodemys
4485:
4483:
4481:Podocnemididae
4477:
4476:
4474:
4473:
4466:
4458:
4456:
4450:
4449:
4447:
4446:
4438:
4431:
4424:
4417:
4409:
4402:
4395:
4388:
4381:
4378:Lomalatachelys
4373:
4366:
4359:
4352:
4345:
4338:
4331:
4323:
4321:
4315:
4314:
4312:
4311:
4303:
4295:
4287:
4279:
4271:
4263:
4255:
4247:
4239:
4231:
4228:Labrostochelys
4223:
4215:
4207:
4199:
4191:
4183:
4175:
4167:
4159:
4151:
4143:
4135:
4127:
4119:
4111:
4103:
4094:
4092:
4085:
4084:
4082:
4081:
4072:
4070:
4068:Araripemydidae
4060:
4053:
4047:
4046:
4043:
4042:
4039:
4038:
4036:
4035:
4027:
4019:
4011:
4003:
3995:
3986:
3984:
3980:
3979:
3977:
3976:
3968:
3960:
3952:
3944:
3936:
3928:
3920:
3912:
3904:
3896:
3888:
3880:
3872:
3864:
3855:
3853:
3846:
3845:
3843:
3842:
3834:
3826:
3817:
3815:
3808:
3807:
3805:
3804:
3796:
3788:
3781:
3773:
3770:Emarginachelys
3765:
3757:
3749:
3741:
3739:
3730:
3726:
3725:
3722:
3721:
3719:
3718:
3715:Sinaspideretes
3710:
3701:
3699:
3695:
3694:
3692:
3691:
3684:
3677:
3670:
3663:
3655:
3648:
3641:
3634:
3626:
3618:
3610:
3603:
3595:
3588:
3581:
3574:
3566:
3559:
3551:
3549:
3543:
3542:
3540:
3539:
3532:
3524:
3515:
3513:
3504:
3498:
3497:
3494:
3493:
3491:
3490:
3483:
3475:
3468:
3460:
3453:
3446:
3438:
3430:
3423:
3416:
3409:
3402:
3395:
3392:Hesperotestudo
3387:
3379:
3372:
3365:
3357:
3349:
3342:
3335:
3328:
3321:
3314:
3306:
3304:
3298:
3297:
3295:
3294:
3286:
3284:
3282:Platysternidae
3277:
3276:
3274:
3273:
3266:
3259:
3252:
3245:
3238:
3231:
3224:
3217:
3210:
3203:
3196:
3189:
3182:
3175:
3168:
3161:
3154:
3147:
3139:
3131:
3129:
3123:
3122:
3120:
3119:
3111:
3104:
3097:
3090:
3083:
3076:
3069:
3062:
3055:
3048:
3041:
3034:
3027:
3018:
3016:
3007:
3001:
3000:
2997:
2996:
2994:
2993:
2986:
2979:
2972:
2964:
2956:
2954:
2948:
2947:
2945:
2944:
2936:
2934:
2932:Dermatemydidae
2925:
2923:Kinosternoidea
2919:
2918:
2915:
2914:
2912:
2911:
2903:
2895:
2892:Procolpochelys
2887:
2879:
2870:
2868:
2864:
2863:
2861:
2860:
2852:
2844:
2836:
2829:
2821:
2813:
2804:
2802:
2800:Dermochelyidae
2796:
2795:
2793:
2792:
2784:
2776:
2768:
2761:
2753:
2745:
2738:
2730:
2722:
2714:
2707:
2699:
2692:
2689:Carolinochelys
2684:
2677:
2668:
2666:
2657:
2646:
2640:
2639:
2636:
2635:
2632:
2631:
2628:
2627:
2620:
2614:
2605:
2596:
2590:
2589:
2587:
2586:
2580:
2571:
2565:
2559:
2552:
2549:
2548:
2543:
2541:
2540:
2533:
2526:
2518:
2509:
2508:
2501:
2498:
2497:
2495:
2494:
2488:
2483:
2478:
2473:
2467:
2465:
2459:
2458:
2456:
2455:
2449:
2443:
2437:
2431:
2425:
2419:
2413:
2407:
2401:
2399:Stylemys copei
2395:
2389:
2387:Stylemys capax
2383:
2377:
2371:
2369:Stylemys botti
2364:
2362:
2355:
2354:
2352:
2351:
2345:
2343:
2337:
2336:
2334:
2333:
2328:
2322:
2320:
2314:
2313:
2311:
2310:
2305:
2300:
2294:
2292:
2286:
2285:
2283:
2282:
2276:
2270:
2264:
2257:
2255:
2248:
2247:
2245:
2244:
2239:
2233:
2231:
2225:
2224:
2222:
2221:
2215:
2213:
2207:
2206:
2204:
2203:
2198:
2193:
2188:
2183:
2178:
2172:
2170:
2164:
2163:
2161:
2160:
2155:
2150:
2144:
2142:
2136:
2135:
2133:
2132:
2127:
2121:
2119:
2113:
2112:
2110:
2109:
2102:
2100:
2098:Hesperotestudo
2093:
2092:
2090:
2089:
2083:
2077:
2071:
2065:
2059:
2053:
2046:
2044:
2037:
2036:
2034:
2033:
2031:Texas tortoise
2028:
2023:
2018:
2013:
2008:
2002:
2000:
1994:
1993:
1991:
1990:
1985:
1979:
1977:
1971:
1970:
1968:
1967:
1961:
1955:
1949:
1943:
1936:
1934:
1927:
1926:
1924:
1923:
1918:
1916:Karoo padloper
1913:
1907:
1905:
1899:
1898:
1896:
1895:
1889:
1887:
1881:
1880:
1878:
1877:
1871:
1865:
1858:
1856:
1849:
1848:
1846:
1845:
1840:
1835:
1830:
1825:
1820:
1815:
1810:
1799:
1794:
1784:
1778:
1773:
1768:
1763:
1757:
1755:
1754:
1753:
1742:
1741:
1739:
1738:
1732:
1726:
1720:
1714:
1708:
1702:
1697:
1692:
1690:Chaco tortoise
1686:
1684:
1678:
1677:
1675:
1674:
1668:
1662:
1656:
1650:
1648:
1642:
1641:
1639:
1638:
1630:
1625:
1619:
1617:
1611:
1610:
1608:
1607:
1599:
1591:
1583:
1580:A. g. daudinii
1575:
1568:
1561:
1559:
1553:
1552:
1550:
1549:
1543:
1541:
1535:
1534:
1527:
1525:
1522:
1521:
1512:
1506:
1502:
1501:
1499:
1498:
1492:
1486:
1480:
1474:
1468:
1462:
1456:
1449:
1446:
1445:
1439:
1437:
1436:
1429:
1422:
1414:
1408:
1407:
1402:
1380:
1364:Edqvist, Ulf.
1360:
1359:
1345:
1344:External links
1342:
1341:
1340:
1323:
1309:
1287:
1268:
1265:
1262:
1261:
1254:
1228:
1218:. Maryland Zoo
1207:
1184:
1177:
1151:
1130:(2): 294–295.
1110:
1058:
1052:978-0965354097
1051:
1011:
987:
925:
924:
922:
919:
918:
917:
914:
907:
905:
902:
895:
893:
890:
883:
881:
878:
871:
869:
862:
855:
853:
850:
843:
841:
838:
831:
829:
826:
819:
817:
814:
807:
805:
802:
795:
793:
790:
783:
781:
778:
771:
767:
764:
751:
748:
712:
709:
705:Central Africa
684:
681:
659:
656:
627:S. p. babcocki
623:S. p. pardalis
598:
595:
541:
540:
537:
536:
522:
513:
511:— Vetter, 2002
504:
485:
476:
463:
454:
445:
420:
407:
394:
381:
368:
355:
342:
321:
309:
298:Testudo armata
295:
276:
266:
263:
262:
259:
258:
252:
251:
244:
233:
232:
226:
225:
218:
216:
212:
211:
204:
200:
199:
194:
190:
189:
184:
180:
179:
174:
170:
169:
164:
160:
159:
154:
150:
149:
144:
140:
139:
134:
130:
129:
124:
120:
119:
106:
105:
92:
89:
88:
70:
67:
66:
61:
58:
57:
46:
45:
37:
36:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
5036:
5025:
5022:
5020:
5017:
5015:
5012:
5010:
5007:
5005:
5002:
5000:
4997:
4995:
4992:
4991:
4989:
4972:
4967:
4963:
4959:
4954:
4950:
4945:
4939:
4935:
4934:
4932:
4930:
4926:
4918:
4913:
4909:
4905:
4900:
4896:
4892:
4887:
4883:
4879:
4874:
4870:
4866:
4861:
4857:
4853:
4848:
4844:
4840:
4835:
4831:
4827:
4822:
4818:
4814:
4809:
4805:
4801:
4796:
4792:
4788:
4783:
4779:
4775:
4770:
4766:
4761:
4755:
4751:
4746:
4740:
4736:
4735:
4733:
4731:
4727:
4723:
4718:
4706:
4702:
4701:
4697:
4696:
4691:
4690:
4687:
4673:
4672:
4671:Tacuarembemys
4667:
4665:
4664:
4659:
4657:
4656:
4651:
4650:
4648:
4644:
4641:
4637:
4634:
4630:
4616:
4615:
4610:
4608:
4607:
4606:Sahonachelys
4602:
4601:
4599:
4597:
4592:
4586:
4585:
4580:
4578:
4577:
4573:
4571:
4570:
4569:Peltocephalus
4566:
4564:
4563:
4558:
4556:
4555:
4554:Lapparentemys
4550:
4548:
4547:
4543:
4541:
4540:
4535:
4533:
4532:
4527:
4525:
4524:
4519:
4517:
4516:
4511:
4509:
4508:
4503:
4501:
4500:
4495:
4493:
4492:
4487:
4486:
4484:
4482:
4478:
4472:
4471:
4467:
4465:
4464:
4460:
4459:
4457:
4455:
4454:Pelomedusidae
4451:
4445:
4444:
4443:Yaminuechelys
4439:
4437:
4436:
4432:
4430:
4429:
4425:
4423:
4422:
4418:
4416:
4415:
4414:Prochelidella
4410:
4408:
4407:
4403:
4401:
4400:
4396:
4394:
4393:
4389:
4387:
4386:
4382:
4380:
4379:
4374:
4372:
4371:
4367:
4365:
4364:
4360:
4358:
4357:
4353:
4351:
4350:
4346:
4344:
4343:
4339:
4337:
4336:
4332:
4330:
4329:
4328:Acanthochelys
4325:
4324:
4322:
4320:
4316:
4310:
4309:
4304:
4302:
4301:
4296:
4294:
4293:
4288:
4286:
4285:
4280:
4278:
4277:
4272:
4270:
4269:
4264:
4262:
4261:
4256:
4254:
4253:
4248:
4246:
4245:
4240:
4238:
4237:
4232:
4230:
4229:
4224:
4222:
4221:
4216:
4214:
4213:
4212:Kinkonychelys
4208:
4206:
4205:
4200:
4198:
4197:
4196:Itapecuruemys
4192:
4190:
4189:
4184:
4182:
4181:
4176:
4174:
4173:
4168:
4166:
4165:
4160:
4158:
4157:
4156:Eotaphrosphys
4152:
4150:
4149:
4144:
4142:
4141:
4136:
4134:
4133:
4128:
4126:
4125:
4120:
4118:
4117:
4112:
4110:
4109:
4104:
4102:
4101:
4096:
4095:
4093:
4091:
4090:Bothremydidae
4086:
4080:
4079:
4074:
4073:
4071:
4069:
4064:
4061:
4057:
4054:
4052:
4048:
4034:
4033:
4028:
4026:
4025:
4020:
4018:
4017:
4012:
4010:
4009:
4004:
4002:
4001:
4000:Argillochelys
3996:
3994:
3993:
3988:
3987:
3985:
3981:
3975:
3974:
3969:
3967:
3966:
3965:Santanachelys
3961:
3959:
3958:
3953:
3951:
3950:
3945:
3943:
3942:
3937:
3935:
3934:
3929:
3927:
3926:
3921:
3919:
3918:
3913:
3911:
3910:
3909:Desmatochelys
3905:
3903:
3902:
3897:
3895:
3894:
3893:Calcarichelys
3889:
3887:
3886:
3881:
3879:
3878:
3877:Atlantochelys
3873:
3871:
3870:
3865:
3863:
3862:
3857:
3856:
3854:
3852:
3851:Protostegidae
3847:
3841:
3840:
3839:Jiangxichelys
3835:
3833:
3832:
3827:
3825:
3824:
3823:Anomalochelys
3819:
3818:
3816:
3814:
3809:
3803:
3802:
3801:Protochelydra
3797:
3795:
3794:
3793:Planiplastron
3789:
3787:
3786:
3782:
3780:
3779:
3774:
3772:
3771:
3766:
3764:
3763:
3758:
3756:
3755:
3750:
3748:
3747:
3743:
3742:
3740:
3738:
3734:
3731:
3727:
3717:
3716:
3711:
3709:
3708:
3703:
3702:
3700:
3696:
3690:
3689:
3685:
3683:
3682:
3678:
3676:
3675:
3671:
3669:
3668:
3664:
3662:
3661:
3656:
3654:
3653:
3649:
3647:
3646:
3642:
3640:
3639:
3635:
3633:
3632:
3627:
3625:
3624:
3619:
3617:
3616:
3611:
3609:
3608:
3604:
3602:
3601:
3600:Drazinderetes
3596:
3594:
3593:
3589:
3587:
3586:
3582:
3580:
3579:
3575:
3573:
3572:
3567:
3565:
3564:
3560:
3558:
3557:
3553:
3552:
3550:
3548:
3544:
3538:
3537:
3536:Carettochelys
3533:
3531:
3530:
3525:
3523:
3522:
3517:
3516:
3514:
3512:
3508:
3505:
3503:
3499:
3489:
3488:
3484:
3482:
3481:
3476:
3474:
3473:
3469:
3467:
3466:
3461:
3459:
3458:
3454:
3452:
3451:
3447:
3445:
3444:
3439:
3437:
3436:
3431:
3429:
3428:
3424:
3422:
3421:
3420:Malacochersus
3417:
3415:
3414:
3410:
3408:
3407:
3403:
3401:
3400:
3396:
3394:
3393:
3388:
3386:
3385:
3380:
3378:
3377:
3373:
3371:
3370:
3366:
3364:
3363:
3358:
3356:
3355:
3350:
3348:
3347:
3343:
3341:
3340:
3336:
3334:
3333:
3329:
3327:
3326:
3322:
3320:
3319:
3315:
3313:
3312:
3311:Aldabrachelys
3308:
3307:
3305:
3303:
3299:
3293:
3292:
3288:
3287:
3285:
3283:
3278:
3272:
3271:
3267:
3265:
3264:
3260:
3258:
3257:
3253:
3251:
3250:
3246:
3244:
3243:
3239:
3237:
3236:
3232:
3230:
3229:
3225:
3223:
3222:
3218:
3216:
3215:
3211:
3209:
3208:
3204:
3202:
3201:
3197:
3195:
3194:
3193:Leucocephalon
3190:
3188:
3187:
3183:
3181:
3180:
3176:
3174:
3173:
3169:
3167:
3166:
3162:
3160:
3159:
3155:
3153:
3152:
3148:
3146:
3145:
3144:Banhxeochelys
3140:
3138:
3137:
3133:
3132:
3130:
3128:
3124:
3118:
3117:
3112:
3110:
3109:
3105:
3103:
3102:
3098:
3096:
3095:
3091:
3089:
3088:
3084:
3082:
3081:
3077:
3075:
3074:
3070:
3068:
3067:
3063:
3061:
3060:
3056:
3054:
3053:
3049:
3047:
3046:
3042:
3040:
3039:
3035:
3033:
3032:
3028:
3026:
3025:
3020:
3019:
3017:
3015:
3011:
3008:
3006:
3005:Testudinoidea
3002:
2992:
2991:
2987:
2985:
2984:
2980:
2978:
2977:
2973:
2971:
2970:
2965:
2963:
2962:
2958:
2957:
2955:
2953:
2952:Kinosternidae
2949:
2943:
2942:
2938:
2937:
2935:
2933:
2929:
2926:
2924:
2920:
2910:
2909:
2904:
2902:
2901:
2900:Protosphargis
2896:
2894:
2893:
2888:
2886:
2885:
2880:
2878:
2877:
2872:
2871:
2869:
2865:
2859:
2858:
2853:
2851:
2850:
2845:
2843:
2842:
2837:
2835:
2834:
2830:
2828:
2827:
2822:
2820:
2819:
2814:
2812:
2811:
2806:
2805:
2803:
2801:
2797:
2791:
2790:
2785:
2783:
2782:
2777:
2775:
2774:
2769:
2767:
2766:
2762:
2760:
2759:
2754:
2752:
2751:
2746:
2744:
2743:
2739:
2737:
2736:
2731:
2729:
2728:
2723:
2721:
2720:
2715:
2713:
2712:
2708:
2706:
2705:
2700:
2698:
2697:
2693:
2691:
2690:
2685:
2683:
2682:
2678:
2676:
2675:
2670:
2669:
2667:
2665:
2661:
2658:
2656:(Sea turtles)
2654:
2650:
2647:
2645:
2641:
2625:
2624:
2618:
2615:
2613:
2609:
2606:
2604:
2600:
2597:
2595:
2591:
2585:
2581:
2579:
2575:
2572:
2570:
2566:
2564:
2560:
2558:
2554:
2553:
2550:
2546:
2539:
2534:
2532:
2527:
2525:
2520:
2519:
2516:
2505:
2499:
2493:
2489:
2487:
2484:
2482:
2479:
2477:
2474:
2472:
2469:
2468:
2466:
2464:
2460:
2454:
2450:
2448:
2444:
2442:
2438:
2436:
2432:
2430:
2426:
2424:
2420:
2418:
2414:
2412:
2408:
2406:
2402:
2400:
2396:
2394:
2390:
2388:
2384:
2382:
2378:
2376:
2372:
2370:
2366:
2365:
2363:
2361:
2356:
2350:
2347:
2346:
2344:
2342:
2338:
2332:
2329:
2327:
2324:
2323:
2321:
2319:
2315:
2309:
2308:Tent tortoise
2306:
2304:
2301:
2299:
2296:
2295:
2293:
2291:
2287:
2281:
2277:
2275:
2271:
2269:
2265:
2263:
2259:
2258:
2256:
2254:
2249:
2243:
2240:
2238:
2235:
2234:
2232:
2230:
2226:
2220:
2217:
2216:
2214:
2212:
2211:Malacochersus
2208:
2202:
2199:
2197:
2194:
2192:
2189:
2187:
2184:
2182:
2179:
2177:
2174:
2173:
2171:
2169:
2165:
2159:
2156:
2154:
2151:
2149:
2146:
2145:
2143:
2141:
2137:
2131:
2128:
2126:
2123:
2122:
2120:
2118:
2114:
2108:
2104:
2103:
2101:
2099:
2094:
2088:
2084:
2082:
2078:
2076:
2072:
2070:
2066:
2064:
2060:
2058:
2054:
2052:
2048:
2047:
2045:
2043:
2038:
2032:
2029:
2027:
2024:
2022:
2019:
2017:
2014:
2012:
2009:
2007:
2004:
2003:
2001:
1999:
1995:
1989:
1986:
1984:
1981:
1980:
1978:
1976:
1972:
1966:
1962:
1960:
1956:
1954:
1950:
1948:
1944:
1942:
1938:
1937:
1935:
1933:
1928:
1922:
1919:
1917:
1914:
1912:
1911:Nama padloper
1909:
1908:
1906:
1904:
1900:
1894:
1891:
1890:
1888:
1886:
1882:
1876:
1872:
1870:
1866:
1864:
1860:
1859:
1857:
1855:
1850:
1844:
1841:
1839:
1836:
1834:
1831:
1829:
1826:
1824:
1821:
1819:
1816:
1814:
1811:
1808:
1804:
1800:
1798:
1795:
1792:
1788:
1785:
1783:
1779:
1777:
1774:
1772:
1769:
1767:
1764:
1762:
1759:
1758:
1756:
1752:
1749:
1748:
1747:
1743:
1737:
1733:
1731:
1727:
1725:
1721:
1719:
1715:
1713:
1709:
1707:
1703:
1701:
1698:
1696:
1693:
1691:
1688:
1687:
1685:
1683:
1679:
1673:
1669:
1667:
1663:
1661:
1657:
1655:
1652:
1651:
1649:
1647:
1643:
1637:
1636:
1631:
1629:
1626:
1624:
1621:
1620:
1618:
1616:
1612:
1606:
1605:
1600:
1598:
1597:
1592:
1589:
1588:
1584:
1582:
1581:
1576:
1574:
1573:
1572:A. g. arnoldi
1569:
1566:
1563:
1562:
1560:
1558:
1557:Aldabrachelys
1554:
1548:
1545:
1544:
1542:
1540:
1536:
1518:
1517:
1510:
1507:
1503:
1497:
1493:
1491:
1490:Testudinoidea
1488:Superfamily:
1487:
1485:
1481:
1479:
1475:
1473:
1469:
1467:
1463:
1461:
1457:
1455:
1451:
1450:
1447:
1442:
1435:
1430:
1428:
1423:
1421:
1416:
1415:
1412:
1406:
1403:
1393:
1389:
1387:
1381:
1371:
1367:
1362:
1361:
1358:
1354:
1353:
1348:
1347:
1343:
1338:
1334:
1331:
1327:
1324:
1321:
1317:
1313:
1310:
1307:
1303:
1302:0-88359-042-5
1299:
1295:
1291:
1288:
1285:
1281:
1278:
1274:
1271:
1270:
1266:
1257:
1255:9780874744149
1251:
1247:
1242:
1241:
1232:
1229:
1217:
1211:
1208:
1203:
1202:
1197:
1191:
1189:
1185:
1180:
1174:
1170:
1166:
1160:
1158:
1156:
1152:
1147:
1143:
1138:
1133:
1129:
1125:
1121:
1114:
1111:
1106:
1102:
1098:
1094:
1090:
1086:
1082:
1081:
1076:
1072:
1065:
1063:
1059:
1054:
1048:
1044:
1040:
1036:
1029:
1022:
1020:
1018:
1016:
1012:
1001:
997:
991:
988:
975:
970:
966:
962:
961:
956:
954:
945:
943:
941:
939:
937:
935:
933:
931:
927:
920:
911:
906:
899:
894:
887:
882:
875:
870:
866:
859:
854:
847:
842:
835:
830:
823:
818:
811:
806:
799:
794:
787:
782:
775:
770:
765:
763:
761:
756:
749:
747:
743:
741:
717:
710:
708:
706:
702:
698:
694:
690:
682:
680:
677:
672:
664:
657:
655:
653:
649:
646:
645:specific name
642:
638:
634:
630:
628:
624:
620:
616:
612:
608:
604:
596:
594:
592:
588:
584:
580:
579:
574:
570:
569:Cape Province
566:
562:
559:found in the
558:
554:
553:
548:
534:
526:
523:
517:
514:
508:
505:
502:
501:
494:
489:
486:
480:
477:
473:
467:
464:
458:
455:
449:
446:
442:
438:
434:
430:
424:
421:
417:
411:
408:
404:
398:
395:
391:
385:
382:
378:
372:
369:
365:
359:
356:
352:
346:
343:
339:
333:
329:
325:
322:
318:
313:
310:
307:
299:
296:
293:
292:
285:
280:
277:
271:
268:
267:
260:
257:
253:
248:
242:
240:
234:
231:
230:Binomial name
227:
223:
222:
217:
214:
213:
210:
209:
205:
202:
201:
198:
195:
192:
191:
188:
187:Testudinoidea
185:
183:Superfamily:
182:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
116:
111:
107:
101:
95:
90:
84:
79:
78:Least Concern
68:
64:
59:
56:
52:
47:
43:
38:
33:
30:
19:
4999:Stigmochelys
4928:
4729:
4694:
4669:
4661:
4653:
4612:
4604:
4582:
4574:
4567:
4560:
4552:
4546:Erymnochelys
4544:
4537:
4531:Cerrejonemys
4529:
4521:
4513:
4507:Brontochelys
4505:
4497:
4489:
4468:
4461:
4441:
4433:
4426:
4421:Pseudemydura
4419:
4412:
4404:
4397:
4390:
4383:
4376:
4368:
4361:
4354:
4347:
4340:
4333:
4326:
4306:
4298:
4290:
4282:
4274:
4266:
4258:
4250:
4242:
4234:
4226:
4218:
4210:
4202:
4194:
4186:
4178:
4170:
4162:
4154:
4146:
4138:
4130:
4122:
4114:
4106:
4098:
4076:
4030:
4024:Prionochelys
4022:
4014:
4006:
3998:
3990:
3971:
3963:
3955:
3947:
3939:
3931:
3923:
3915:
3907:
3901:Cratochelone
3899:
3891:
3885:Bouliachelys
3883:
3875:
3867:
3861:Alienochelys
3859:
3837:
3829:
3821:
3799:
3791:
3783:
3776:
3768:
3762:Chelydropsis
3760:
3752:
3744:
3713:
3707:Basilochelys
3705:
3686:
3679:
3672:
3665:
3658:
3650:
3643:
3636:
3631:Khunnuchelys
3629:
3621:
3613:
3605:
3598:
3590:
3583:
3576:
3569:
3561:
3554:
3547:Trionychidae
3534:
3527:
3521:Allaeochelys
3519:
3485:
3478:
3472:Stigmochelys
3471:
3470:
3463:
3455:
3448:
3441:
3435:Megalochelys
3433:
3425:
3418:
3411:
3404:
3397:
3390:
3382:
3374:
3367:
3360:
3352:
3346:Cylindraspis
3344:
3337:
3330:
3325:Centrochelys
3323:
3316:
3309:
3302:Testudinidae
3291:Platysternon
3289:
3270:Vijayachelys
3268:
3261:
3254:
3249:Rhinoclemmys
3247:
3240:
3233:
3226:
3219:
3214:Melanochelys
3212:
3205:
3198:
3191:
3184:
3177:
3170:
3163:
3156:
3149:
3142:
3134:
3114:
3106:
3099:
3092:
3085:
3078:
3071:
3064:
3057:
3050:
3043:
3036:
3029:
3024:Acherontemys
3022:
2990:Sternotherus
2988:
2981:
2974:
2967:
2959:
2939:
2906:
2898:
2890:
2882:
2874:
2857:Psephophorus
2855:
2847:
2839:
2831:
2824:
2816:
2808:
2787:
2779:
2773:Pacifichelys
2771:
2763:
2756:
2748:
2742:Lepidochelys
2740:
2733:
2725:
2719:Gigantatypus
2717:
2711:Eretmochelys
2709:
2702:
2694:
2687:
2679:
2672:
2653:Chelonioidea
2621:
2573:
2348:
2341:Stigmochelys
2253:Megalochelys
1932:Cylindraspis
1854:Cheirogaster
1646:Centrochelys
1633:
1602:
1594:
1585:
1578:
1570:
1567:(subspecies:
1514:
1496:Testudinidae
1395:. Retrieved
1391:
1385:
1373:. Retrieved
1369:
1350:
1336:
1332:
1329:
1320:Stigmochelys
1319:
1315:
1305:
1293:
1290:Branch, Bill
1283:
1282:: 419–421. (
1279:
1276:
1239:
1231:
1220:. Retrieved
1210:
1199:
1168:
1165:Branch, Bill
1127:
1123:
1113:
1084:
1078:
1074:
1070:
1034:
1003:. Retrieved
999:
990:
978:. Retrieved
964:
958:
952:
757:
753:
750:Conservation
744:
737:
697:South Africa
686:
673:
669:
651:
647:
640:
636:
633:Stigmochelys
632:
631:
626:
622:
619:Stigmochelys
618:
614:
613:(2002), and
611:Centrochelys
610:
607:Stigmochelys
606:
602:
600:
576:
573:Stigmochelys
572:
551:
550:
546:
544:
532:
524:
520:— Bour, 2002
515:
506:
498:
487:
483:— Dadd, 1974
478:
465:
456:
447:
422:
409:
396:
383:
370:
357:
344:
331:
327:
323:
311:
305:
297:
289:
278:
269:
238:
236:
220:
219:
208:Stigmochelys
207:
197:Testudinidae
55:South Africa
29:
4821:iNaturalist
4754:Wikispecies
4584:Stupendemys
4539:Cordichelys
4385:Mesoclemmys
4370:Hydromedusa
4292:Taphrosphys
4284:Sankuchemys
4276:Rhothonemys
4252:Polysternon
4132:Cearachelys
4116:Azabbaremys
4100:Araiochelys
4016:Ctenochelys
4008:Bashuchelys
3957:Rhinochelys
3933:Ocepechelon
3925:Notochelone
3917:Iserosaurus
3785:Macrochelys
3737:Chelydridae
3660:Palaeoamyda
3585:Cyclanorbis
3450:Psammobates
3443:Oligopherus
3406:Indotestudo
3354:Cymatholcus
3332:Chelonoidis
3318:Astrochelys
3127:Geoemydidae
3045:Deirochelys
2983:Staurotypus
2976:Kinosternon
2969:Hoplochelys
2884:Peritresius
2833:Dermochelys
2826:Cosmochelys
2818:Corsochelys
2735:Itilochelys
2727:Glarichelys
2674:Allopleuron
2664:Cheloniidae
2603:Superfamily
2290:Psammobates
2140:Indotestudo
1682:Chelonoidis
1615:Astrochelys
1326:Loveridge A
1091:: 298–307.
980:12 November
740:herbivorous
693:East Africa
689:South Sudan
658:Description
615:Psammobates
377:E. Williams
306:nomen nudum
291:nomen nudum
96:Appendix II
4988:Categories
4663:Caririemys
4576:Podocnemis
4562:Latentemys
4523:Carbonemys
4463:Pelomedusa
4300:Ummulisani
4220:Kurmademys
4188:Inaechelys
4172:Galianemys
4140:Chedighaii
4078:Araripemys
4051:Pleurodira
4032:Toxochelys
3949:Protostega
3754:Chelydrops
3674:Pelodiscus
3667:Pelochelys
3615:Gilmoremys
3592:Cycloderma
3502:Trionychia
3369:Geochelone
3362:Floridemys
3228:Notochelys
3116:Wilburemys
3087:Malaclemys
2941:Dermatemys
2908:Puppigerus
2841:Eosphargis
2758:Miocaretta
2750:Mexichelys
2644:Cryptodira
2584:Testudines
2545:Testudines
1975:Geochelone
1903:Chersobius
1539:Agrionemys
1484:Cryptodira
1482:Suborder:
1478:Testudines
1470:Subclass:
1397:2016-04-11
1375:2016-04-11
1222:2021-01-30
1075:Geochelone
1005:2022-01-14
921:References
865:Azerbaijan
760:heartwater
603:Geochelone
587:succulents
578:Geochelone
328:Geochelone
324:Geochelone
302:Boie, 1831
274:Bell, 1828
177:Cryptodira
173:Suborder:
167:Testudines
4944:Q45735818
4703:See also
4655:Caribemys
4515:Caninemys
4499:Bauruemys
4428:Rheodytes
4392:Myuchelys
4335:Chelodina
4260:Puentemys
4236:Nigeremys
4180:Ilatardia
4124:Bothremys
3831:Basilemys
3645:Nilssonia
3623:Hutchemys
3571:Axestemys
3529:Anosteira
3384:Hadrianus
3242:Pangshura
3200:Malayemys
3165:Geoclemys
3108:Trachemys
3101:Terrapene
3094:Pseudemys
3080:Graptemys
3073:Glyptemys
3066:Emydoidea
3059:Actinemys
3031:Chrysemys
2876:Euclastes
2704:Eochelone
2555:Kingdom:
2042:Hadrianus
1452:Kingdom:
1335:: 1–19. (
1000:cites.org
533:ex errore
500:ex errore
493:Młynarski
472:Pritchard
433:Raxworthy
364:Loveridge
338:Fitzinger
215:Species:
133:Kingdom:
127:Eukaryota
4971:10676519
4938:Wikidata
4912:Species+
4904:pardalis
4839:10363364
4739:Wikidata
4470:Pelusios
4435:Rhinemys
4406:Platemys
4399:Phrynops
4319:Chelidae
4308:Zolhafah
4204:Jainemys
3869:Archelon
3746:Chelydra
3638:Lissemys
3480:Stylemys
3465:Solitudo
3427:Manouria
3376:Gopherus
3339:Chersina
3207:Mauremys
3186:Heosemys
3179:Hardella
3172:Geoemyda
3158:Cyclemys
3014:Emydidae
2961:Claudius
2810:Arabemys
2789:Tasbacka
2781:Syllomus
2696:Chelonia
2594:Suborder
2578:Diapsida
2569:Reptilia
2563:Chordata
2561:Phylum:
2557:Animalia
2360:Stylemys
2229:Manouria
1998:Gopherus
1885:Chersina
1516:tortoise
1494:Family:
1472:Anapsida
1466:Reptilia
1460:Chordata
1458:Phylum:
1454:Animalia
1441:Tortoise
1349:Species
1314:(1873).
1292:(2004).
1167:(2008).
1146:87809001
1105:17611092
1089:Elsevier
791:Juvenile
676:carapace
648:pardalis
609:(2001),
605:(1957),
591:thistles
583:aardvark
561:savannas
557:tortoise
332:pardalis
264:Synonyms
256:Synonyms
193:Family:
157:Reptilia
147:Chordata
143:Phylum:
137:Animalia
123:Domain:
83:IUCN 3.1
4958:2441763
4813:2441757
4745:Q640955
4614:Sokatra
4363:Emydura
4268:Rosasia
4164:Foxemys
4108:Arenila
3688:Trionyx
3681:Rafetus
3607:Dogania
3563:Apalone
3487:Testudo
3413:Kinixys
3399:Homopus
3256:Sacalia
3235:Orlitia
3221:Morenia
3136:Batagur
3038:Clemmys
2765:Natator
2681:Caretta
2582:Order:
2567:Class:
2463:Testudo
2168:Kinixys
2117:Homopus
1476:Order:
1464:Class:
1312:Gray JE
1246:248–249
1080:Zoology
1071:Testudo
766:Gallery
701:Namibia
641:chelone
390:Gerlach
249:, 1828)
203:Genus:
163:Order:
153:Class:
98: (
81: (
4891:776504
4878:569598
4865:163449
4852:551780
4800:794299
4774:751388
4646:
4639:
4632:
4356:Elusor
4349:Elseya
4342:Chelus
4059:
3992:Adocus
3983:
3729:
3698:
3578:Chitra
3280:
2867:
2612:Family
1519:family
1505:Genera
1443:family
1300:
1273:Bell T
1252:
1175:
1144:
1103:
1049:
652:pardus
637:stigma
495:, 1976
474:, 1967
443:, 2006
439:&
437:McCord
418:, 2002
405:, 2002
403:Vetter
392:, 2001
379:, 1957
366:, 1935
353:, 1933
351:Hewitt
340:, 1835
319:, 1831
286:, 1829
284:Cuvier
4966:IRMNG
4834:IRMNG
4826:40092
4787:52MQ5
3652:Palea
3556:Amyda
3457:Pyxis
3151:Cuora
2623:Genus
2574:Clade
2318:Pyxis
1791:Diego
1142:S2CID
1087:(4).
1031:(PDF)
867:stamp
695:, to
565:Sudan
441:Mertz
100:CITES
94:CITES
4953:GBIF
4917:8867
4873:NCBI
4860:IUCN
4847:ITIS
4808:GBIF
4769:BOLD
3052:Emys
1298:ISBN
1250:ISBN
1173:ISBN
1101:PMID
1073:and
1047:ISBN
982:2021
965:2015
699:and
674:The
625:and
589:and
545:The
416:Bour
317:Gray
247:Bell
4795:EoL
4782:CoL
1355:at
1304:. (
1132:doi
1093:doi
1085:110
1077:".
1039:doi
969:doi
4990::
4968::
4955::
4940::
4914::
4901::
4899:RD
4888::
4875::
4862::
4849::
4836::
4823::
4810::
4797::
4784::
4771::
4756::
4741::
2576::
1805:†(
1390:.
1368:.
1333:79
1248:.
1198:.
1187:^
1154:^
1140:.
1128:57
1126:.
1122:.
1099:.
1083:.
1061:^
1045:.
1033:.
1014:^
998:.
963:.
957:.
929:^
593:.
470:—
435:,
431:,
429:Le
427:—
414:—
401:—
388:—
362:—
349:—
336:—
330:)
53:,
4668:†
4660:†
4652:†
4611:†
4603:†
4594:†
4581:†
4559:†
4551:†
4536:†
4528:†
4520:†
4512:†
4504:†
4496:†
4488:†
4440:†
4411:†
4375:†
4305:†
4297:†
4289:†
4281:†
4273:†
4265:†
4257:†
4249:†
4241:†
4233:†
4225:†
4217:†
4209:†
4201:†
4193:†
4185:†
4177:†
4169:†
4161:†
4153:†
4145:†
4137:†
4129:†
4121:†
4113:†
4105:†
4097:†
4088:†
4075:†
4066:†
4029:†
4021:†
4013:†
4005:†
3997:†
3989:†
3970:†
3962:†
3954:†
3946:†
3938:†
3930:†
3922:†
3914:†
3906:†
3898:†
3890:†
3882:†
3874:†
3866:†
3858:†
3849:†
3836:†
3828:†
3820:†
3811:†
3798:†
3790:†
3775:†
3767:†
3759:†
3751:†
3712:†
3704:†
3657:†
3628:†
3620:†
3612:†
3597:†
3568:†
3526:†
3518:†
3477:†
3462:†
3440:†
3432:†
3389:†
3381:†
3359:†
3351:†
3141:†
3113:†
3021:†
2966:†
2905:†
2897:†
2889:†
2881:†
2873:†
2854:†
2846:†
2838:†
2823:†
2815:†
2807:†
2786:†
2778:†
2770:†
2755:†
2747:†
2732:†
2724:†
2716:†
2701:†
2686:†
2671:†
2537:e
2530:t
2523:v
2490:†
2451:†
2445:†
2439:†
2433:†
2427:†
2421:†
2415:†
2409:†
2403:†
2397:†
2391:†
2385:†
2379:†
2373:†
2367:†
2358:†
2278:†
2272:†
2266:†
2260:†
2251:†
2105:†
2096:†
2085:†
2079:†
2073:†
2067:†
2061:†
2055:†
2049:†
2040:†
1963:†
1957:†
1951:†
1945:†
1939:†
1930:†
1873:†
1867:†
1861:†
1852:†
1809:)
1801:†
1793:)
1789:(
1780:†
1734:†
1728:†
1722:†
1716:†
1710:†
1704:†
1670:†
1664:†
1658:†
1632:†
1601:†
1593:†
1590:)
1577:†
1433:e
1426:t
1419:v
1400:.
1384:"
1378:.
1280:3
1258:.
1225:.
1204:.
1181:.
1148:.
1134::
1107:.
1095::
1055:.
1041::
1008:.
984:.
971::
955:"
951:"
549:(
535:)
531:(
503:)
497:(
326:(
308:)
304:(
294:)
288:(
245:(
102:)
85:)
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
