472:
382:
4260:
397:
90:
225:
4245:
427:
4252:
4975:
4947:
50:
4942:
412:
442:
188:
457:
371:
520:
Female treeshrews have a gestation period of 45–50 days and give birth to up to three young in nests lined with dry leaves inside tree hollows. The young are born blind and hairless, but are able to leave the nest after about a month. During this period, the mother provides relatively little
486:
Treeshrews are slender animals with long tails and soft, greyish to reddish-brown fur. The terrestrial species tend to be larger than the arboreal forms, and to have larger claws, which they use for digging up insect prey. They have poorly developed canine teeth and unspecialised molars, with an
2025:
Meredith, Robert W.; Janečka, Jan E.; Gatesy, John; Ryder, Oliver A.; Fisher, Colleen A.; Teeling, Emma C.; Goodbla, Alisha; Eizirik, Eduardo; Simão, Taiz L. L. (28 October 2011). "Impacts of the
Cretaceous terrestrial revolution and KPg extinction on mammal diversification".
471:
1926:
Janecka, Jan E.; Miller, Webb; Pringle, Thomas H.; Wiens, Frank; Zitzmann, Annette; Helgen, Kristofer M.; Springer, Mark S.; Murphy, William J. (2 November 2007). "Molecular and genomic data identify the closest living relatives of the
Primates".
1981:
Zhou, Xuming; Sun, Fengming; Xu, Shixia; Yang, Guang; Li, Ming (1 March 2015). "The position of tree shrews in the mammalian tree: Comparing multi-gene analyses with phylogenomic results leaves monophyly of
Euarchonta doubtful".
2135:
Lin, J.; Chen, G.; Gu, L.; Shen, Y.; Zheng, M.; Zheng, W.; Hu, X.; Zhang, X.; Qiu, Y.; Liu, X.; Jiang, C. (2014). "Phylogenetic affinity of tree shrews to Glires is attributed to fast evolution rate".
1897:
909:
Several other arrangements of these orders have been proposed in the past, and the above tree is only a well-favored proposal. Although it is known that
Scandentia is one of the most
2336:
Roberts, T.E.; Lanier, H.C.; Sargis, E.J.; Olson, L.E. (2011). "Molecular phylogeny of treeshrews (Mammalia: Scandentia) and the timescale of diversification in
Southeast Asia".
716:
cannot be ruled out. Some studies place
Scandentia as sister of the Glires, which would invalidate Euarchonta: It is this organization that is shown in the tree diagram below.
1883:
5143:
5182:
1480:
of India. Most belong to the family
Tupaiidae; one fossil species described from the Oligocene of Yunnan is thought to be closer to the pen-tailed treeshrew.
3362:
1905:
4259:
5246:
5117:
5156:
1783:
1730:
1567:
2966:
524:
Treeshrews reach sexual maturity after around four months, and breed for much of the year, with no clear breeding season in most species.
1638:
918:
913:
Euarchontoglire clades, the exact phylogenetic position is not yet considered resolved: It may be a sister of Glires, Primatomorpha, or
584:
The pen-tailed treeshrew in
Malaysia is able to consume large amounts of naturally fermented nectar from flower buds of the bertam palm
381:
2496:
5241:
3078:
1223:
400:
3069:
1214:
1800:
5174:
3355:
5161:
2086:
5169:
5065:
1549:
89:
5187:
4978:
3348:
3111:
2959:
1253:
308:
4244:
3183:
1325:
445:
4946:
1491:
1485:
4180:
3120:
1509:
1262:
672:. Taxonomists continue to refine the treeshrews' relations to primates and to other closely related clades.
4995:
4894:
4318:
1623:
396:
4675:
5208:
3192:
2489:
1877:
1456:
1334:
910:
661:
156:
1515:
5070:
4251:
5091:
5042:
4941:
4659:
4308:
3310:
3301:
3291:
3255:
2387:
2240:
2101:
2035:
1936:
1837:
1437:
1388:
1154:
1136:
541:
475:
282:
4844:
4796:
3788:
3781:
3774:
3753:
1497:
5236:
4917:
4782:
4588:
4474:
4465:
4435:
4338:
4038:
3802:
3596:
3528:
3479:
3330:
2952:
2944:
675:
552:
297:
4837:
4816:
4748:
4696:
4648:
4558:
4444:
4393:
4028:
3816:
3795:
3760:
3507:
1161:; the two families are thought to have separated 60 million years ago. The former Tupaiidae genus
4775:
4710:
4682:
4509:
4328:
4197:
4108:
3918:
3911:
3823:
3809:
3767:
3746:
3708:
3680:
3673:
3652:
3638:
3500:
3219:
3210:
3147:
3138:
3102:
2460:
2295:
2294:
Kriegs, Jan Ole; Churakov, Gennady; Jurka, Jerzy; Brosius, Jürgen; Schmitz, Jürgen (2007-02-20).
2067:
1960:
1688:
1352:
1289:
1280:
1244:
586:
533:
415:
84:
4703:
4537:
4495:
4451:
4414:
4370:
4115:
4048:
3904:
3897:
3687:
3645:
3552:
3538:
3493:
3472:
3451:
2085:
Pettigrew, J.D.; Jamieson, B.G.; Robson, S.K.; Hall, L.S.; McAnally, K.I.; Cooper, H.M. (1989).
1775:
1768:
1722:
1557:
5213:
4858:
4641:
4627:
4574:
4523:
4488:
4481:
4421:
4407:
4400:
4160:
4139:
3883:
3860:
3853:
3722:
3659:
3514:
3465:
1657:
5195:
5078:
4668:
4502:
3846:
3715:
3694:
3666:
3545:
3246:
3237:
3228:
3165:
3129:
2857:
2415:
2353:
2315:
2276:
2258:
2209:
2191:
2152:
2117:
2059:
2051:
2007:
1999:
1952:
1865:
1779:
1749:
A new universal etymological technological, and pronouncing dictionary of the
English Language
1726:
1680:
1573:
1563:
1379:
1370:
1361:
1307:
1271:
590:(with up to 3.8% alcohol content) the entire year without it having any effects on behaviour.
575:, feeding on insects, small vertebrates, fruit, and seeds. Among other things, treeshrews eat
505:
323:
4567:
3925:
3701:
5200:
5148:
4968:
4613:
4011:
3607:
3422:
3371:
3156:
3045:
3036:
3016:
2878:
2619:
2556:
2482:
2450:
2405:
2395:
2345:
2307:
2266:
2248:
2199:
2183:
2144:
2109:
2043:
1991:
1944:
1855:
1845:
1672:
1503:
1298:
1193:
1149:, which are divided into two families. The majority are in the "ordinary" treeshrew family,
1071:
928:
853:
723:
709:
624:
509:
430:
385:
359:
270:
5033:
224:
4055:
3952:
3736:
3273:
3264:
3201:
3174:
3093:
2771:
2672:
2640:
1658:"The tree shrews: Adjuncts and alternatives to primates as models for biomedical research"
1617:
1406:
1397:
1343:
1316:
679:
617:
460:
263:
255:
2889:(Rats, guinea pigs, squirrels, beavers, chinchillas, porcupines, capybaras and relatives)
521:
maternal care, visiting her young only for a few minutes every other day to suckle them.
315:, but high ratios are not uncommon for animals weighing less than 1 kg (2 lb).
2391:
2376:"An early Oligocene fossil demonstrates treeshrews are slowly evolving "living fossils""
2244:
2105:
2039:
1940:
1841:
1747:
1642:
1468:, China, but the identity of this animal is uncertain. Other fossils have come from the
4735:
4153:
4101:
3945:
3874:
2817:
2763:
2664:
2648:
2410:
2375:
2271:
2228:
2204:
2171:
1860:
1825:
1715:
1238:
653:
because of certain internal similarities to primates (for example, similarities in the
426:
351:
340:
259:
58:
5230:
4883:
4807:
4759:
4384:
4353:
4129:
4070:
2808:
2680:
2229:"Coalescent-based genome analyses resolve the early branches of the Euarchontoglires"
1676:
1553:
1461:
1420:
1158:
1075:
1015:
816:
713:
683:
631:
278:
215:
203:
177:
39:
2464:
2071:
1964:
1824:
Wiens F, Zitzmann A, Lachance MA, Yegles M, Pragst F, Wurst FM; et al. (2008).
1692:
5047:
4768:
4597:
4291:
4278:
4212:
4190:
4091:
2825:
2755:
2656:
2611:
1810:
49:
5083:
2253:
5130:
5027:
4963:
4830:
4823:
4789:
4363:
3991:
3837:
3585:
3568:
3441:
3407:
2865:
2787:
2705:
2349:
2148:
1528:
1431:
1079:
921:(SINEs) offer strong evidence for scandentia belonging to the euarchonta group:
712:. However, the alternative placement of treeshrews as sister to both Glires and
646:
556:
289:
411:
370:
4906:
4717:
4689:
4606:
4544:
4530:
4516:
4146:
4079:
4018:
3968:
3622:
3486:
3060:
2894:
2868:(Pigs, camels, hippos, deer, buffalo, gazelles, giraffes, whales and dolphins)
2747:
2590:
2547:
2531:
2455:
2434:
2311:
1208:
989:
951:
914:
769:
706:
698:
690:
687:
605:
537:
441:
322:, and have been used as an alternative to primates in experimental studies of
187:
141:
5018:
2262:
2195:
2055:
2003:
604:
reduces their pain response to capsaicinoids, which scientists believe is an
288:
Though called 'treeshrews', and despite having previously been classified in
5104:
4851:
4620:
4002:
3026:
2849:
2726:
2718:
2598:
2564:
2047:
1995:
1948:
1850:
1577:
1176:
1150:
665:
627:
594:
577:
545:
488:
327:
274:
209:
197:
172:
101:
2419:
2357:
2319:
2280:
2213:
2187:
2156:
2113:
2063:
2011:
1956:
1869:
1684:
1591:
668:", however they were soon split from the primates and moved into their own
2121:
678:
studies have suggested that the treeshrews should be given the same rank (
497:
494:
456:
17:
5057:
5012:
4634:
3890:
3458:
3389:
3010:
2998:
2841:
2697:
2525:
1898:"See Why Tree Shrews Are Only the Second Known Mammal to Seek Spicy Food"
1805:
1477:
1187:
650:
601:
572:
355:
319:
301:
121:
1454:
The fossil record of treeshrews is poor. The oldest putative treeshrew,
4874:
3631:
2918:
2795:
2627:
2537:
2474:
1469:
1163:
1142:
1067:
868:
657:
635:
2400:
2170:
Foley, Nicole M.; Springer, Mark S.; Teeling, Emma C. (19 July 2016).
1616:
593:
Treeshrews have also been observed intentionally eating foods high in
5135:
4171:
3936:
3401:
3395:
3383:
3004:
2992:
2910:
2886:
2734:
2581:
2519:
2506:
1473:
1035:
967:
938:
829:
785:
756:
702:
694:
693:. According to this classification, the Euarchonta are sister to the
251:
131:
111:
4989:
917:, or separate from and sister to all other Euarchontoglires. Shared
5122:
5109:
3340:
346:
2779:
2087:"Phylogenetic relations between microbats, megabats, and primates"
1562:(3rd ed.). Johns Hopkins University Press. pp. 104–109.
1465:
1146:
669:
654:
639:
598:
560:
470:
455:
440:
425:
410:
395:
380:
312:
293:
74:
68:
1559:
Mammal
Species of the World: A Taxonomic and Geographic Reference
2296:"Evolutionary history of 7SL RNA-derived SINEs in Supraprimates"
4993:
4276:
3420:
3344:
2948:
2478:
2227:
Kumar, Vikas; Hallström, Björn M.; Janke, Axel (1 April 2013).
1167:
was disbanded in 2011 when the Mindanao treeshrew was moved to
597:, a behavior unique among mammals other than humans. A single
2833:
2435:"Tupaiine tree shrews (Scandentia, Mammalia) from the Yuanmou
2172:"Mammal madness: Is the mammal tree of life not yet resolved?"
1826:"Chronic intake of fermented floral nectar by wild treeshrews"
1157:, is different enough to warrant placement in its own family,
5096:
1641:. The Genome Institute. Washington University. Archived from
608:
to be able to consume spicy foods in their natural habitats.
532:
Treeshrews live in small family groups, which defend their
318:
Among orders of mammals, treeshrews are closely related to
1721:. Baltimore, MD: Johns Hopkins University Press. p.
1656:
Cao, J.; Yang, E.B.; Su, J.-J.; Li, Y.; Chow, P. (2003).
4732:
4585:
4555:
4462:
4432:
4381:
4350:
4334:
4324:
4314:
4304:
4186:
4126:
4088:
4044:
4034:
4024:
3871:
3834:
3733:
3619:
3603:
3592:
3581:
3525:
3438:
5002:
4905:
4882:
4872:
4806:
4758:
4731:
4658:
4596:
4584:
4554:
4461:
4431:
4380:
4349:
4289:
4211:
4170:
4125:
4087:
4078:
4069:
4001:
3977:
3966:
3935:
3870:
3833:
3732:
3618:
3566:
3524:
3437:
3299:
3290:
3091:
3058:
3034:
3025:
2877:
2816:
2807:
2746:
2717:
2696:
2639:
2610:
2589:
2580:
2555:
2546:
232:The distribution and density of treeshrew species.
2921:(lorises, lemurs, tarsiers, monkeys, apes, humans)
1767:
1714:
2176:Philosophical Transactions of the Royal Society B
2094:Philosophical Transactions of the Royal Society B
682:) as the primates and, with the primates and the
1801:"Tiny tree shrew can drink you under the table"
1766:Martin, Robert D. (1984). Macdonald, D. (ed.).
2331:
2329:
3356:
2960:
2490:
277:(19 species, "ordinary" treeshrews), and the
8:
1882:: CS1 maint: multiple names: authors list (
705:), and the two groups are combined into the
304:; among other things, treeshrews eat fruit.
4990:
4879:
4740:
4593:
4297:
4286:
4273:
4084:
4075:
3983:
3974:
3574:
3434:
3430:
3417:
3363:
3349:
3341:
3296:
3031:
2967:
2953:
2945:
2813:
2714:
2607:
2586:
2552:
2497:
2483:
2475:
512:in the case of the more arboreal species.
223:
186:
48:
31:
2454:
2409:
2399:
2369:
2367:
2270:
2252:
2203:
1976:
1974:
1859:
1849:
2828:(Hedgehogs, shrews, moles and relatives)
1774:. New York, NY: Facts on File. pp.
634:, the pen-tailed treeshrew. The 20
369:
1540:
563:, depending on the particular species.
1875:
2338:Molecular Phylogenetics and Evolution
2137:Molecular Phylogenetics and Evolution
2096:. Mammalia: Chiroptera and Primates.
1761:
1759:
645:Treeshrews were moved from the order
7:
1060:
1028:
1021:
995:
944:
934:
924:
919:short interspersed nuclear elements
845:
822:
762:
736:
729:
719:
3079:Northern smooth-tailed treeshrew (
1472:of Thailand, Pakistan, India, and
664:), and classified as a "primitive
25:
3070:Bornean smooth-tailed treeshrew (
2860:(Horses, rhinoceroses and tapirs)
1615:Rines, George Edwin, ed. (1920).
311:than any other mammal, including
4974:
4973:
4945:
4940:
4258:
4250:
4243:
1677:10.1034/j.1600-0684.2003.00022.x
1639:"an article on Tupaia belangeri"
1224:Northern smooth-tailed treeshrew
1171:based on a molecular phylogeny.
401:Northern smooth-tailed treeshrew
88:
5247:Extant Eocene first appearances
1215:Bornean smooth-tailed treeshrew
2443:Swiss Journal of Palaeontology
1665:Journal of Medical Primatology
1:
1717:Walker's Mammals of the World
1483:Named fossil species include
2254:10.1371/journal.pone.0060019
2350:10.1016/j.ympev.2011.04.021
2149:10.1016/j.ympev.2013.12.001
1770:The Encyclopedia of Mammals
326:, psychosocial stress, and
5263:
3112:Golden-bellied treeshrew (
2897:(Rabbits, hares and pikas)
2852:(Dogs, cats and relatives)
2758:(Tenrecs and golden moles)
2439:locality of Yunnan, China"
1134:
358:, and was provided by Sir
262:. They make up the entire
4958:
4938:
4743:
4300:
4285:
4272:
4241:
3986:
3577:
3433:
3429:
3416:
3378:
3327:
3121:Bangka Island treeshrew (
2987:
2683:(Kangaroos and relatives)
2514:
2456:10.1007/s13358-011-0029-0
2312:10.1016/j.tig.2007.02.002
1799:Moscowicz, Clara (2008).
1065:
1058:
1033:
1026:
1019:
1000:
993:
965:
949:
942:
932:
866:
850:
843:
827:
820:
783:
767:
760:
741:
734:
727:
686:(colugos), belong to the
536:from intruders. Most are
307:Treeshrews have a higher
231:
222:
194:
185:
169:
164:
85:Scientific classification
83:
66:
56:
47:
34:
2675:(Bilbies and bandicoots)
2433:Ni, X.; Qiu, Z. (2012).
1830:Proc Natl Acad Sci U S A
1476:, China, as well as the
1254:Golden-bellied treeshrew
954:(rabbits, hares, pikas)
772:(rabbits, hares, pikas)
616:They make up the entire
309:brain to body mass ratio
5242:Fauna of Southeast Asia
3193:Long-footed treeshrew (
3184:Horsfield's treeshrew (
2567:(Platypus and echidnas)
2374:Li, Q.; Ni, X. (2016).
2048:10.1126/science.1211028
1996:10.1111/1749-4877.12116
1949:10.1126/science.1147555
1851:10.1073/pnas.0801628105
1492:Prodendrogale engesseri
1486:Prodendrogale yunnanica
1263:Bangka Island treeshrew
1153:, but one species, the
606:evolutionary adaptation
269:, which split into two
3311:Pen-tailed treeshrew (
3256:Kalimantan treeshrew (
2798:(Dugongs and manatees)
2737:(Anteaters and sloths)
2188:10.1098/rstb.2015.0140
2114:10.1098/rstb.1989.0102
1902:nationalgeographic.com
1624:Encyclopedia Americana
1556:; Reeder, D.M (eds.).
1510:Palaeotupaia sivalicus
676:Molecular phylogenetic
630:, the treeshrews, and
553:mark their territories
483:
468:
453:
438:
423:
408:
393:
378:
296:, and not all species
5209:Paleobiology Database
2667:(Quolls and dunnarts)
1548:Helgen, K.M. (2005).
1335:Long-footed treeshrew
1326:Horsfield's treeshrew
662:Wilfrid Le Gros Clark
660:, highlighted by Sir
504:Treeshrews have good
474:
459:
446:Horsfield's treeshrew
444:
429:
414:
399:
384:
373:
3220:Mountain treeshrew (
3211:Calamian treeshrew (
3148:Sumatran treeshrew (
3139:Mindanao treeshrew (
3103:Northern treeshrew (
1746:Craig, John (1849).
1713:Nowak, R.M. (1999).
1592:"Appendices | CITES"
1438:Pen-tailed treeshrew
1389:Kalimantan treeshrew
1155:pen-tailed treeshrew
1137:List of scandentians
542:pen-tailed treeshrew
476:Pen-tailed treeshrew
292:, they are not true
283:pen-tailed treeshrew
3247:Painted treeshrew (
3238:Palawan treeshrew (
3229:Nicobar treeshrew (
3166:Slender treeshrew (
3130:Striped treeshrew (
2392:2016NatSR...618627L
2245:2013PLoSO...860019K
2106:1989RSPTB.325..489P
2040:2011Sci...334..521M
1984:Integrative Zoology
1941:2007Sci...318..792J
1842:2008PNAS..10510426W
1145:are placed in four
638:are placed in five
478:(1850 depiction of
3157:Common treeshrew (
3046:Madras treeshrew (
2651:(Monito del monte)
2380:Scientific Reports
2300:Trends in Genetics
2182:(1699): 20150140.
1908:on August 27, 2018
1618:"Tree-shrew"
1550:"Order Scandentia"
1457:Eodendrogale parva
1353:Mountain treeshrew
1290:Sumatran treeshrew
1281:Mindanao treeshrew
1245:Northern treeshrew
587:Eugeissona tristis
484:
469:
454:
439:
424:
416:Northern treeshrew
409:
394:
379:
281:(one species, the
5224:
5223:
5196:Open Tree of Life
4996:Taxon identifiers
4987:
4986:
4954:
4953:
4936:
4935:
4932:
4931:
4928:
4927:
4868:
4867:
4727:
4726:
4268:
4267:
4239:
4238:
4235:
4234:
4231:
4230:
4207:
4206:
4065:
4064:
3962:
3961:
3562:
3561:
3338:
3337:
3323:
3322:
3286:
3285:
3274:Large treeshrew (
3265:Ruddy treeshrew (
3202:Pygmy treeshrew (
3175:Javan treeshrew (
2982:species by family
2980:
2942:
2941:
2938:
2937:
2934:
2933:
2930:
2929:
2922:
2914:
2906:
2898:
2890:
2869:
2861:
2853:
2845:
2837:
2829:
2799:
2791:
2783:
2775:
2767:
2766:(Elephant shrews)
2759:
2738:
2730:
2711:
2692:
2691:
2684:
2676:
2668:
2660:
2659:(Marsupial moles)
2657:Notoryctemorphia
2652:
2631:
2623:
2620:Paucituberculata
2604:
2576:
2575:
2568:
2401:10.1038/srep18627
2100:(1229): 489–559.
2034:(6055): 521–524.
1935:(5851): 792–794.
1813:on 15 March 2014.
1785:978-0-87196-871-5
1732:978-0-8018-5789-8
1569:978-0-8018-8221-0
1516:Ptilocercus kylin
1380:Painted treeshrew
1371:Palawan treeshrew
1362:Nicobar treeshrew
1308:Slender treeshrew
1272:Striped treeshrew
1127:
1126:
1118:
1117:
1109:
1108:
1100:
1099:
1091:
1090:
1047:
1046:
979:
978:
906:
905:
897:
896:
888:
887:
879:
878:
806:
805:
797:
796:
623:, split into the
480:Ptilocercus lowii
405:Dendrogale murina
236:
235:
160:
78:
16:(Redirected from
5254:
5217:
5216:
5204:
5203:
5191:
5190:
5178:
5177:
5165:
5164:
5152:
5151:
5139:
5138:
5126:
5125:
5113:
5112:
5100:
5099:
5087:
5086:
5074:
5073:
5061:
5060:
5051:
5050:
5038:
5037:
5036:
5023:
5022:
5021:
4991:
4977:
4976:
4969:Plesiadapiformes
4949:
4944:
4880:
4741:
4734:
4594:
4587:
4557:
4464:
4434:
4383:
4352:
4336:
4326:
4316:
4306:
4298:
4287:
4274:
4263:
4262:
4255:
4254:
4247:
4188:
4128:
4090:
4085:
4076:
4046:
4036:
4026:
3984:
3975:
3873:
3836:
3735:
3621:
3608:Zalambdalestidae
3605:
3594:
3583:
3575:
3527:
3440:
3435:
3431:
3423:Euarchontoglires
3418:
3372:Euarchontoglires
3365:
3358:
3351:
3342:
3297:
3213:T. moellendorffi
3032:
3017:Euarchontoglires
2978:
2969:
2962:
2955:
2946:
2920:
2912:
2904:
2896:
2888:
2879:Euarchontoglires
2867:
2859:
2851:
2843:
2835:
2827:
2814:
2797:
2789:
2781:
2773:
2765:
2757:
2736:
2728:
2715:
2710:
2709:
2701:
2682:
2674:
2673:Peramelemorphia
2666:
2658:
2650:
2629:
2628:Didelphimorphia
2622:(Shrew opossums)
2621:
2608:
2603:
2602:
2594:
2587:
2566:
2557:Australosphenida
2553:
2499:
2492:
2485:
2476:
2469:
2468:
2458:
2430:
2424:
2423:
2413:
2403:
2371:
2362:
2361:
2333:
2324:
2323:
2291:
2285:
2284:
2274:
2256:
2224:
2218:
2217:
2207:
2167:
2161:
2160:
2132:
2126:
2125:
2091:
2082:
2076:
2075:
2022:
2016:
2015:
1978:
1969:
1968:
1923:
1917:
1916:
1914:
1913:
1904:. Archived from
1894:
1888:
1887:
1881:
1873:
1863:
1853:
1836:(30): 10426–31.
1821:
1815:
1814:
1809:. Archived from
1796:
1790:
1789:
1773:
1763:
1754:
1753:
1743:
1737:
1736:
1720:
1710:
1704:
1703:
1701:
1699:
1662:
1653:
1647:
1646:
1635:
1629:
1628:
1620:
1612:
1606:
1605:
1603:
1602:
1588:
1582:
1581:
1545:
1504:Tupaia miocenica
1299:Common treeshrew
1194:Madras treeshrew
1131:Order Scandentia
1072:Plesiadapiformes
1061:
1029:
1022:
996:
945:
935:
929:Euarchontoglires
925:
854:Plesiadapiformes
846:
823:
763:
737:
730:
724:Euarchontoglires
720:
710:Euarchontoglires
501:
500:
499:
496:
431:Common treeshrew
420:Tupaia belangeri
390:Anathana ellioti
386:Madras treeshrew
360:Stamford Raffles
344:is derived from
256:tropical forests
227:
190:
155:
93:
92:
72:
52:
38:Temporal range:
32:
27:Order of mammals
21:
5262:
5261:
5257:
5256:
5255:
5253:
5252:
5251:
5227:
5226:
5225:
5220:
5212:
5207:
5199:
5194:
5186:
5181:
5173:
5168:
5160:
5155:
5147:
5142:
5134:
5129:
5121:
5116:
5108:
5103:
5095:
5090:
5082:
5077:
5069:
5064:
5056:
5054:
5046:
5041:
5032:
5031:
5026:
5017:
5016:
5011:
4998:
4988:
4983:
4950:
4924:
4901:
4864:
4802:
4754:
4723:
4660:Palaechthonidae
4654:
4580:
4550:
4457:
4427:
4376:
4345:
4309:Asioplesiadapis
4294:
4281:
4264:
4257:
4249:
4227:
4203:
4166:
4121:
4061:
3997:
3971:
3958:
3953:Simplicidentata
3931:
3866:
3829:
3737:Arctostylopidae
3728:
3614:
3571:
3558:
3520:
3425:
3412:
3374:
3369:
3339:
3334:
3319:
3282:
3240:T. palawanensis
3114:T. chrysogaster
3087:
3054:
3021:
2983:
2973:
2943:
2926:
2873:
2858:Perissodactyla
2803:
2742:
2703:
2702:
2700:
2688:
2665:Dasyuromorphia
2649:Microbiotheria
2641:Australidelphia
2635:
2596:
2595:
2593:
2572:
2542:
2510:
2503:
2473:
2472:
2432:
2431:
2427:
2373:
2372:
2365:
2335:
2334:
2327:
2293:
2292:
2288:
2226:
2225:
2221:
2169:
2168:
2164:
2134:
2133:
2129:
2089:
2084:
2083:
2079:
2024:
2023:
2019:
1980:
1979:
1972:
1925:
1924:
1920:
1911:
1909:
1896:
1895:
1891:
1874:
1823:
1822:
1818:
1798:
1797:
1793:
1786:
1765:
1764:
1757:
1745:
1744:
1740:
1733:
1712:
1711:
1707:
1697:
1695:
1660:
1655:
1654:
1650:
1645:on 1 June 2010.
1637:
1636:
1632:
1614:
1613:
1609:
1600:
1598:
1590:
1589:
1585:
1570:
1547:
1546:
1542:
1537:
1525:
1513:from India and
1507:from Thailand,
1452:
1407:Large treeshrew
1398:Ruddy treeshrew
1375:T. palawanensis
1344:Pygmy treeshrew
1317:Javan treeshrew
1258:T. chrysogaster
1139:
1133:
1128:
1119:
1110:
1101:
1092:
1048:
980:
907:
898:
889:
880:
807:
798:
649:into the order
614:
571:Treeshrews are
569:
540:, although the
530:
518:
493:
492:
461:Pygmy treeshrew
368:
336:
207:, Purple: both
154:
87:
79:
43:
36:
28:
23:
22:
15:
12:
11:
5:
5260:
5258:
5250:
5249:
5244:
5239:
5229:
5228:
5222:
5221:
5219:
5218:
5205:
5192:
5179:
5166:
5153:
5140:
5127:
5114:
5101:
5088:
5075:
5062:
5052:
5039:
5024:
5008:
5006:
5000:
4999:
4994:
4985:
4984:
4982:
4981:
4971:
4966:
4959:
4956:
4955:
4952:
4951:
4939:
4937:
4934:
4933:
4930:
4929:
4926:
4925:
4923:
4922:
4921:
4920:
4911:
4909:
4903:
4902:
4900:
4899:
4898:
4897:
4888:
4886:
4877:
4870:
4869:
4866:
4865:
4863:
4862:
4855:
4848:
4845:Elphidotarsius
4841:
4834:
4827:
4820:
4812:
4810:
4804:
4803:
4801:
4800:
4797:Pronothodectes
4793:
4786:
4779:
4772:
4764:
4762:
4756:
4755:
4753:
4752:
4744:
4738:
4736:Plesiadapoidea
4729:
4728:
4725:
4724:
4722:
4721:
4714:
4707:
4700:
4693:
4686:
4679:
4672:
4664:
4662:
4656:
4655:
4653:
4652:
4645:
4638:
4631:
4624:
4617:
4610:
4602:
4600:
4591:
4582:
4581:
4579:
4578:
4571:
4563:
4561:
4552:
4551:
4549:
4548:
4541:
4534:
4527:
4520:
4513:
4506:
4499:
4492:
4485:
4478:
4470:
4468:
4459:
4458:
4456:
4455:
4448:
4440:
4438:
4429:
4428:
4426:
4425:
4418:
4411:
4404:
4397:
4389:
4387:
4378:
4377:
4375:
4374:
4367:
4359:
4357:
4347:
4346:
4344:
4343:
4332:
4322:
4312:
4301:
4295:
4290:
4283:
4282:
4277:
4270:
4269:
4266:
4265:
4242:
4240:
4237:
4236:
4233:
4232:
4229:
4228:
4226:
4225:
4224:
4223:
4217:
4215:
4209:
4208:
4205:
4204:
4202:
4201:
4194:
4184:
4176:
4174:
4172:Cynocephalidae
4168:
4167:
4165:
4164:
4157:
4154:Planetetherium
4150:
4143:
4135:
4133:
4123:
4122:
4120:
4119:
4112:
4105:
4102:Dracontolestes
4097:
4095:
4082:
4073:
4067:
4066:
4063:
4062:
4060:
4059:
4052:
4042:
4032:
4022:
4015:
4007:
4005:
3999:
3998:
3996:
3995:
3987:
3981:
3972:
3967:
3964:
3963:
3960:
3959:
3957:
3956:
3949:
3946:Duplicidentata
3941:
3939:
3933:
3932:
3930:
3929:
3922:
3915:
3908:
3901:
3894:
3887:
3879:
3877:
3875:Pseudictopidae
3868:
3867:
3865:
3864:
3857:
3850:
3842:
3840:
3831:
3830:
3828:
3827:
3820:
3813:
3806:
3799:
3792:
3789:Kazachostylops
3785:
3782:Enantiostylops
3778:
3775:Bothriostylops
3771:
3764:
3757:
3754:Anatolostylops
3750:
3742:
3740:
3730:
3729:
3727:
3726:
3719:
3712:
3705:
3698:
3691:
3684:
3677:
3670:
3663:
3656:
3649:
3642:
3635:
3627:
3625:
3616:
3615:
3613:
3612:
3601:
3590:
3578:
3572:
3567:
3564:
3563:
3560:
3559:
3557:
3556:
3549:
3542:
3534:
3532:
3522:
3521:
3519:
3518:
3511:
3504:
3497:
3490:
3483:
3476:
3469:
3462:
3455:
3447:
3445:
3427:
3426:
3421:
3414:
3413:
3411:
3410:
3404:
3398:
3392:
3386:
3379:
3376:
3375:
3370:
3368:
3367:
3360:
3353:
3345:
3336:
3335:
3328:
3325:
3324:
3321:
3320:
3318:
3317:
3307:
3305:
3294:
3288:
3287:
3284:
3283:
3281:
3280:
3271:
3267:T. splendidula
3262:
3253:
3244:
3235:
3226:
3217:
3208:
3199:
3190:
3181:
3172:
3163:
3154:
3145:
3136:
3127:
3118:
3109:
3099:
3097:
3089:
3088:
3086:
3085:
3076:
3066:
3064:
3056:
3055:
3053:
3052:
3042:
3040:
3029:
3023:
3022:
3020:
3019:
3013:
3007:
3001:
2995:
2988:
2985:
2984:
2974:
2972:
2971:
2964:
2957:
2949:
2940:
2939:
2936:
2935:
2932:
2931:
2928:
2927:
2925:
2924:
2916:
2908:
2900:
2892:
2883:
2881:
2875:
2874:
2872:
2871:
2863:
2855:
2847:
2839:
2831:
2822:
2820:
2818:Laurasiatheria
2811:
2805:
2804:
2802:
2801:
2793:
2785:
2777:
2772:Tubulidentata
2769:
2764:Macroscelidea
2761:
2752:
2750:
2744:
2743:
2741:
2740:
2732:
2723:
2721:
2712:
2694:
2693:
2690:
2689:
2687:
2686:
2681:Diprotodontia
2678:
2670:
2662:
2654:
2645:
2643:
2637:
2636:
2634:
2633:
2625:
2616:
2614:
2605:
2584:
2578:
2577:
2574:
2573:
2571:
2570:
2561:
2559:
2550:
2544:
2543:
2541:
2540:
2534:
2528:
2522:
2515:
2512:
2511:
2504:
2502:
2501:
2494:
2487:
2479:
2471:
2470:
2437:Lufengpithecus
2425:
2363:
2344:(3): 358–372.
2325:
2306:(4): 158–161.
2286:
2219:
2162:
2127:
2077:
2017:
1990:(2): 186–198.
1970:
1918:
1889:
1816:
1791:
1784:
1755:
1738:
1731:
1705:
1671:(3): 123–130.
1648:
1630:
1607:
1583:
1568:
1539:
1538:
1536:
1533:
1532:
1531:
1524:
1521:
1498:Tupaia storchi
1460:, is from the
1451:
1448:
1447:
1446:
1445:
1444:
1426:
1425:
1423:
1416:
1415:
1414:
1413:
1404:
1402:T. splendidula
1395:
1386:
1377:
1368:
1359:
1350:
1341:
1332:
1323:
1314:
1305:
1296:
1287:
1278:
1269:
1260:
1251:
1233:
1232:
1231:
1230:
1221:
1203:
1202:
1201:
1200:
1182:
1181:
1179:
1135:Main article:
1132:
1129:
1125:
1124:
1121:
1120:
1116:
1115:
1112:
1111:
1107:
1106:
1103:
1102:
1098:
1097:
1094:
1093:
1089:
1088:
1085:
1084:
1064:
1059:
1057:
1054:
1053:
1050:
1049:
1045:
1044:
1041:
1040:
1032:
1027:
1025:
1020:
1018:
1012:
1011:
1008:
1007:
999:
994:
992:
986:
985:
982:
981:
977:
976:
973:
972:
964:
961:
960:
957:
956:
948:
943:
941:
933:
931:
923:
904:
903:
900:
899:
895:
894:
891:
890:
886:
885:
882:
881:
877:
876:
873:
872:
865:
862:
861:
858:
857:
849:
844:
842:
839:
838:
835:
834:
826:
821:
819:
813:
812:
809:
808:
804:
803:
800:
799:
795:
794:
791:
790:
782:
779:
778:
775:
774:
766:
761:
759:
753:
752:
749:
748:
740:
735:
733:
728:
726:
718:
613:
610:
568:
565:
555:using various
529:
526:
517:
514:
489:dental formula
367:
364:
335:
332:
260:Southeast Asia
254:native to the
234:
233:
229:
228:
220:
219:
216:Ptilocercidae
204:Ptilocercidae
192:
191:
183:
182:
181:
180:
175:
167:
166:
162:
161:
149:
145:
144:
139:
135:
134:
129:
125:
124:
119:
115:
114:
109:
105:
104:
99:
95:
94:
81:
80:
67:
64:
63:
54:
53:
45:
44:
37:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
5259:
5248:
5245:
5243:
5240:
5238:
5235:
5234:
5232:
5215:
5210:
5206:
5202:
5197:
5193:
5189:
5184:
5180:
5176:
5171:
5167:
5163:
5158:
5154:
5150:
5145:
5141:
5137:
5132:
5128:
5124:
5119:
5115:
5111:
5106:
5102:
5098:
5093:
5089:
5085:
5080:
5076:
5072:
5067:
5063:
5059:
5053:
5049:
5044:
5040:
5035:
5029:
5025:
5020:
5014:
5010:
5009:
5007:
5005:
5001:
4997:
4992:
4980:
4972:
4970:
4967:
4965:
4961:
4960:
4957:
4948:
4943:
4919:
4915:
4914:
4913:
4912:
4910:
4908:
4904:
4896:
4895:Strepsirrhini
4892:
4891:
4890:
4889:
4887:
4885:
4884:Strepsirrhini
4881:
4878:
4876:
4871:
4861:
4860:
4856:
4854:
4853:
4849:
4847:
4846:
4842:
4840:
4839:
4835:
4833:
4832:
4828:
4826:
4825:
4821:
4819:
4818:
4814:
4813:
4811:
4809:
4808:Carpolestidae
4805:
4799:
4798:
4794:
4792:
4791:
4787:
4785:
4784:
4783:Platychoerops
4780:
4778:
4777:
4773:
4771:
4770:
4766:
4765:
4763:
4761:
4760:Plesiadapidae
4757:
4751:
4750:
4746:
4745:
4742:
4739:
4737:
4730:
4720:
4719:
4715:
4713:
4712:
4708:
4706:
4705:
4701:
4699:
4698:
4694:
4692:
4691:
4687:
4685:
4684:
4680:
4678:
4677:
4673:
4671:
4670:
4666:
4665:
4663:
4661:
4657:
4651:
4650:
4646:
4644:
4643:
4639:
4637:
4636:
4632:
4630:
4629:
4625:
4623:
4622:
4618:
4616:
4615:
4611:
4609:
4608:
4604:
4603:
4601:
4599:
4595:
4592:
4590:
4589:Paromomyoidea
4583:
4577:
4576:
4572:
4570:
4569:
4565:
4564:
4562:
4560:
4553:
4547:
4546:
4542:
4540:
4539:
4535:
4533:
4532:
4528:
4526:
4525:
4521:
4519:
4518:
4514:
4512:
4511:
4507:
4505:
4504:
4500:
4498:
4497:
4493:
4491:
4490:
4486:
4484:
4483:
4479:
4477:
4476:
4475:Arctodontomys
4472:
4471:
4469:
4467:
4466:Microsyopidae
4460:
4454:
4453:
4449:
4447:
4446:
4442:
4441:
4439:
4437:
4436:Picromomyidae
4430:
4424:
4423:
4419:
4417:
4416:
4412:
4410:
4409:
4405:
4403:
4402:
4398:
4396:
4395:
4391:
4390:
4388:
4386:
4385:Micromomyidae
4379:
4373:
4372:
4368:
4366:
4365:
4361:
4360:
4358:
4355:
4354:Purgatoriidae
4348:
4341:
4340:
4339:Picrodontidae
4333:
4331:
4330:
4323:
4321:
4320:
4313:
4311:
4310:
4303:
4302:
4299:
4296:
4293:
4288:
4284:
4280:
4275:
4271:
4261:
4256:
4253:
4246:
4221:
4220:
4219:
4218:
4216:
4214:
4210:
4200:
4199:
4195:
4193:
4192:
4185:
4183:
4182:
4178:
4177:
4175:
4173:
4169:
4163:
4162:
4158:
4156:
4155:
4151:
4149:
4148:
4144:
4142:
4141:
4137:
4136:
4134:
4131:
4130:Plagiomenidae
4124:
4118:
4117:
4113:
4111:
4110:
4106:
4104:
4103:
4099:
4098:
4096:
4093:
4086:
4083:
4081:
4077:
4074:
4072:
4071:Primatomorpha
4068:
4058:
4057:
4053:
4051:
4050:
4043:
4041:
4040:
4039:Prodendrogale
4033:
4031:
4030:
4023:
4021:
4020:
4016:
4014:
4013:
4009:
4008:
4006:
4004:
4000:
3994:
3993:
3989:
3988:
3985:
3982:
3980:
3976:
3973:
3970:
3965:
3955:
3954:
3950:
3948:
3947:
3943:
3942:
3940:
3938:
3934:
3928:
3927:
3923:
3921:
3920:
3916:
3914:
3913:
3909:
3907:
3906:
3902:
3900:
3899:
3895:
3893:
3892:
3888:
3886:
3885:
3881:
3880:
3878:
3876:
3869:
3863:
3862:
3858:
3856:
3855:
3851:
3849:
3848:
3844:
3843:
3841:
3839:
3832:
3826:
3825:
3821:
3819:
3818:
3814:
3812:
3811:
3807:
3805:
3804:
3803:Palaeostylops
3800:
3798:
3797:
3793:
3791:
3790:
3786:
3784:
3783:
3779:
3777:
3776:
3772:
3770:
3769:
3765:
3763:
3762:
3758:
3756:
3755:
3751:
3749:
3748:
3744:
3743:
3741:
3738:
3731:
3725:
3724:
3720:
3718:
3717:
3713:
3711:
3710:
3706:
3704:
3703:
3699:
3697:
3696:
3692:
3690:
3689:
3685:
3683:
3682:
3678:
3676:
3675:
3671:
3669:
3668:
3664:
3662:
3661:
3657:
3655:
3654:
3650:
3648:
3647:
3643:
3641:
3640:
3636:
3634:
3633:
3629:
3628:
3626:
3624:
3617:
3610:
3609:
3602:
3599:
3598:
3597:Picrodontidae
3591:
3588:
3587:
3580:
3579:
3576:
3573:
3570:
3565:
3555:
3554:
3550:
3548:
3547:
3543:
3541:
3540:
3536:
3535:
3533:
3530:
3529:Picrodontidae
3523:
3517:
3516:
3512:
3510:
3509:
3505:
3503:
3502:
3498:
3496:
3495:
3491:
3489:
3488:
3484:
3482:
3481:
3480:Frugivastodon
3477:
3475:
3474:
3470:
3468:
3467:
3463:
3461:
3460:
3456:
3454:
3453:
3449:
3448:
3446:
3443:
3436:
3432:
3428:
3424:
3419:
3415:
3409:
3405:
3403:
3399:
3397:
3393:
3391:
3387:
3385:
3381:
3380:
3377:
3373:
3366:
3361:
3359:
3354:
3352:
3347:
3346:
3343:
3333:
3332:
3326:
3316:
3314:
3309:
3308:
3306:
3304:
3303:
3298:
3295:
3293:
3292:Ptilocercidae
3289:
3279:
3277:
3272:
3270:
3268:
3263:
3261:
3259:
3254:
3252:
3250:
3245:
3243:
3241:
3236:
3234:
3232:
3231:T. nicobarica
3227:
3225:
3223:
3218:
3216:
3214:
3209:
3207:
3205:
3200:
3198:
3196:
3191:
3189:
3187:
3182:
3180:
3178:
3177:T. hypochrysa
3173:
3171:
3169:
3164:
3162:
3160:
3155:
3153:
3151:
3150:T. ferruginea
3146:
3144:
3142:
3137:
3135:
3133:
3128:
3126:
3124:
3119:
3117:
3115:
3110:
3108:
3106:
3101:
3100:
3098:
3096:
3095:
3090:
3084:
3082:
3077:
3075:
3073:
3068:
3067:
3065:
3063:
3062:
3057:
3051:
3049:
3044:
3043:
3041:
3039:
3038:
3033:
3030:
3028:
3024:
3018:
3014:
3012:
3008:
3006:
3002:
3000:
2996:
2994:
2990:
2989:
2986:
2981:
2970:
2965:
2963:
2958:
2956:
2951:
2950:
2947:
2923:
2917:
2915:
2909:
2907:
2901:
2899:
2893:
2891:
2885:
2884:
2882:
2880:
2876:
2870:
2866:Artiodactyla
2864:
2862:
2856:
2854:
2848:
2846:
2840:
2838:
2832:
2830:
2826:Eulipotyphla
2824:
2823:
2821:
2819:
2815:
2812:
2810:
2809:Boreoeutheria
2806:
2800:
2794:
2792:
2786:
2784:
2778:
2776:
2770:
2768:
2762:
2760:
2756:Afrosoricida
2754:
2753:
2751:
2749:
2745:
2739:
2733:
2731:
2725:
2724:
2722:
2720:
2716:
2713:
2707:
2699:
2695:
2685:
2679:
2677:
2671:
2669:
2663:
2661:
2655:
2653:
2647:
2646:
2644:
2642:
2638:
2632:
2626:
2624:
2618:
2617:
2615:
2613:
2609:
2606:
2600:
2592:
2588:
2585:
2583:
2579:
2569:
2563:
2562:
2560:
2558:
2554:
2551:
2549:
2545:
2539:
2535:
2533:
2529:
2527:
2523:
2521:
2517:
2516:
2513:
2508:
2500:
2495:
2493:
2488:
2486:
2481:
2480:
2477:
2466:
2462:
2457:
2452:
2448:
2444:
2440:
2438:
2429:
2426:
2421:
2417:
2412:
2407:
2402:
2397:
2393:
2389:
2385:
2381:
2377:
2370:
2368:
2364:
2359:
2355:
2351:
2347:
2343:
2339:
2332:
2330:
2326:
2321:
2317:
2313:
2309:
2305:
2301:
2297:
2290:
2287:
2282:
2278:
2273:
2268:
2264:
2260:
2255:
2250:
2246:
2242:
2239:(4): e60019.
2238:
2234:
2230:
2223:
2220:
2215:
2211:
2206:
2201:
2197:
2193:
2189:
2185:
2181:
2177:
2173:
2166:
2163:
2158:
2154:
2150:
2146:
2142:
2138:
2131:
2128:
2123:
2119:
2115:
2111:
2107:
2103:
2099:
2095:
2088:
2081:
2078:
2073:
2069:
2065:
2061:
2057:
2053:
2049:
2045:
2041:
2037:
2033:
2029:
2021:
2018:
2013:
2009:
2005:
2001:
1997:
1993:
1989:
1985:
1977:
1975:
1971:
1966:
1962:
1958:
1954:
1950:
1946:
1942:
1938:
1934:
1930:
1922:
1919:
1907:
1903:
1899:
1893:
1890:
1885:
1879:
1871:
1867:
1862:
1857:
1852:
1847:
1843:
1839:
1835:
1831:
1827:
1820:
1817:
1812:
1808:
1807:
1802:
1795:
1792:
1787:
1781:
1777:
1772:
1771:
1762:
1760:
1756:
1751:
1750:
1742:
1739:
1734:
1728:
1724:
1719:
1718:
1709:
1706:
1694:
1690:
1686:
1682:
1678:
1674:
1670:
1666:
1659:
1652:
1649:
1644:
1640:
1634:
1631:
1626:
1625:
1619:
1611:
1608:
1597:
1593:
1587:
1584:
1579:
1575:
1571:
1565:
1561:
1560:
1555:
1551:
1544:
1541:
1534:
1530:
1527:
1526:
1522:
1520:
1519:from Yunnan.
1518:
1517:
1512:
1511:
1506:
1505:
1501:from Yunnan,
1500:
1499:
1494:
1493:
1488:
1487:
1481:
1479:
1475:
1471:
1467:
1463:
1462:Middle Eocene
1459:
1458:
1450:Fossil record
1449:
1443:
1439:
1436:
1435:
1434:
1433:
1428:
1427:
1424:
1422:
1421:Ptilocercidae
1418:
1417:
1412:
1408:
1405:
1403:
1399:
1396:
1394:
1390:
1387:
1385:
1381:
1378:
1376:
1372:
1369:
1367:
1366:T. nicobarica
1363:
1360:
1358:
1354:
1351:
1349:
1345:
1342:
1340:
1336:
1333:
1331:
1327:
1324:
1322:
1321:T. hypochrysa
1318:
1315:
1313:
1309:
1306:
1304:
1300:
1297:
1295:
1294:T. ferruginea
1291:
1288:
1286:
1282:
1279:
1277:
1273:
1270:
1268:
1264:
1261:
1259:
1255:
1252:
1250:
1246:
1243:
1242:
1241:
1240:
1235:
1234:
1229:
1225:
1222:
1220:
1216:
1213:
1212:
1211:
1210:
1205:
1204:
1199:
1195:
1192:
1191:
1190:
1189:
1184:
1183:
1180:
1178:
1174:
1173:
1172:
1170:
1166:
1165:
1160:
1159:Ptilocercidae
1156:
1152:
1148:
1144:
1138:
1130:
1123:
1122:
1114:
1113:
1105:
1104:
1096:
1095:
1087:
1086:
1083:
1081:
1077:
1076:Strepsirrhini
1073:
1069:
1063:
1062:
1056:
1055:
1052:
1051:
1043:
1042:
1039:
1037:
1031:
1030:
1024:
1023:
1017:
1016:Primatomorpha
1014:
1013:
1010:
1009:
1006:
1005:(treeshrews)
1004:
998:
997:
991:
988:
987:
984:
983:
975:
974:
971:
969:
963:
962:
959:
958:
955:
953:
947:
946:
940:
937:
936:
930:
927:
926:
922:
920:
916:
912:
902:
901:
893:
892:
884:
883:
875:
874:
871:
870:
864:
863:
860:
859:
856:
855:
848:
847:
841:
840:
837:
836:
833:
831:
825:
824:
818:
817:Primatomorpha
815:
814:
811:
810:
802:
801:
793:
792:
789:
787:
781:
780:
777:
776:
773:
771:
765:
764:
758:
755:
754:
751:
750:
747:
746:(treeshrews)
745:
739:
738:
732:
731:
725:
722:
721:
717:
715:
714:Primatomorpha
711:
708:
704:
700:
696:
692:
689:
685:
684:flying lemurs
681:
677:
673:
671:
667:
663:
659:
656:
652:
648:
643:
641:
637:
633:
632:Ptilocercidae
629:
626:
622:
619:
611:
609:
607:
603:
600:
596:
591:
589:
588:
582:
580:
579:
574:
566:
564:
562:
558:
554:
549:
547:
543:
539:
535:
527:
525:
522:
515:
513:
511:
507:
502:
490:
481:
477:
473:
466:
462:
458:
451:
447:
443:
436:
432:
428:
421:
417:
413:
406:
402:
398:
391:
387:
383:
377:
374:Dentition of
372:
365:
363:
361:
357:
353:
349:
348:
343:
342:
333:
331:
329:
325:
321:
316:
314:
310:
305:
303:
299:
298:live in trees
295:
291:
286:
284:
280:
279:Ptilocercidae
276:
272:
268:
265:
261:
258:of South and
257:
253:
249:
245:
242:(also called
241:
230:
226:
221:
218:
217:
212:
211:
206:
205:
200:
199:
193:
189:
184:
179:
178:Ptilocercidae
176:
174:
171:
170:
168:
163:
158:
153:
150:
147:
146:
143:
140:
137:
136:
133:
130:
127:
126:
123:
120:
117:
116:
113:
110:
107:
106:
103:
100:
97:
96:
91:
86:
82:
76:
70:
65:
61:
60:
55:
51:
46:
41:
40:Middle Eocene
33:
30:
19:
5003:
4857:
4850:
4843:
4838:Carpomegodon
4836:
4829:
4822:
4817:Carpocristes
4815:
4795:
4788:
4781:
4774:
4769:Chiromyoides
4767:
4749:Chronolestes
4747:
4716:
4709:
4702:
4697:Plesiolestes
4695:
4688:
4681:
4674:
4667:
4649:Phenacolemur
4647:
4640:
4633:
4626:
4619:
4612:
4605:
4598:Paromomyidae
4573:
4566:
4559:Toliapinidae
4543:
4536:
4529:
4522:
4515:
4508:
4501:
4494:
4487:
4480:
4473:
4450:
4445:Alveojunctus
4443:
4420:
4413:
4406:
4399:
4394:Chalicomomys
4392:
4369:
4362:
4337:
4327:
4317:
4307:
4292:Pan-Primates
4279:Pan-Primates
4248:
4213:Pan-Primates
4196:
4191:Dermotherium
4189:
4181:Cynocephalus
4179:
4159:
4152:
4145:
4138:
4114:
4107:
4100:
4092:Mixodectidae
4054:
4047:
4037:
4029:Palaeotupaia
4027:
4017:
4010:
3990:
3978:
3951:
3944:
3924:
3917:
3910:
3903:
3896:
3889:
3882:
3859:
3852:
3845:
3822:
3817:Stenostylops
3815:
3808:
3801:
3796:Migrostylops
3794:
3787:
3780:
3773:
3766:
3761:Arctostylops
3759:
3752:
3745:
3721:
3714:
3707:
3700:
3693:
3686:
3679:
3672:
3665:
3658:
3651:
3644:
3637:
3630:
3606:
3595:
3584:
3551:
3544:
3537:
3513:
3508:Sinclairella
3506:
3499:
3492:
3485:
3478:
3471:
3464:
3457:
3450:
3406:Infraclass:
3329:
3312:
3300:
3275:
3266:
3257:
3248:
3239:
3230:
3221:
3212:
3203:
3194:
3185:
3176:
3167:
3158:
3149:
3140:
3131:
3122:
3113:
3105:T. belangeri
3104:
3092:
3080:
3071:
3059:
3047:
3035:
2979:(Treeshrews)
2976:
2905:(Treeshrews)
2902:
2788:Proboscidea
2729:(Armadillos)
2612:Ameridelphia
2565:Monotremata
2446:
2442:
2436:
2428:
2386:(1): 18627.
2383:
2379:
2341:
2337:
2303:
2299:
2289:
2236:
2232:
2222:
2179:
2175:
2165:
2140:
2136:
2130:
2097:
2093:
2080:
2031:
2027:
2020:
1987:
1983:
1932:
1928:
1921:
1910:. Retrieved
1906:the original
1901:
1892:
1878:cite journal
1833:
1829:
1819:
1811:the original
1804:
1794:
1769:
1748:
1741:
1716:
1708:
1696:. Retrieved
1668:
1664:
1651:
1643:the original
1633:
1622:
1610:
1599:. Retrieved
1595:
1586:
1558:
1554:Wilson, D.E.
1543:
1514:
1508:
1502:
1496:
1490:
1484:
1482:
1455:
1453:
1441:
1430:
1410:
1401:
1392:
1383:
1374:
1365:
1356:
1347:
1338:
1329:
1320:
1311:
1302:
1293:
1284:
1275:
1266:
1257:
1249:T. belangeri
1248:
1237:
1227:
1218:
1207:
1197:
1186:
1168:
1162:
1141:The 23
1140:
1066:
1034:
1002:
1001:
966:
950:
908:
867:
851:
828:
784:
768:
743:
742:
674:
644:
620:
615:
592:
585:
583:
576:
570:
557:scent glands
550:
531:
523:
519:
516:Reproduction
503:
485:
479:
464:
449:
434:
419:
404:
389:
375:
345:
339:
337:
317:
306:
287:
266:
250:) are small
247:
243:
239:
237:
214:
208:
202:
196:
151:
138:Grandorder:
57:
29:
5131:iNaturalist
5028:Wikispecies
4964:Anagaloidea
4831:Carpolestes
4824:Carpodaptes
4790:Plesiadapis
4776:Nannodectes
4711:Talpohenach
4683:Palenochtha
4676:Palaechthon
4510:Megadelphus
4364:Purgatorius
4329:Russellodon
4319:Pandemonium
4198:Galeopterus
4109:Eudaemonema
3992:Ptilocercus
3919:Pseudictops
3912:Paranictops
3838:Astigalidae
3824:Wanostylops
3810:Sinostylops
3768:Asiostylops
3747:Allostylops
3709:Stenanagale
3681:Huaiyangale
3674:Hsiuannania
3653:Chianshania
3639:Anagalopsis
3586:Apatemyidae
3569:Gliriformes
3501:Labidolemur
3442:Apatemyidae
3408:Placentalia
3302:Ptilocercus
3258:T. salatana
3195:T. longipes
3186:T. javanica
3168:T. gracilis
3141:T. everetti
3132:T. dorsalis
3123:T. discolor
3072:D. melanura
3015:Superorder
3009:Infraclass
2977:Scandentia
2911:Dermoptera
2903:Scandentia
2895:Lagomorpha
2844:(Pangolins)
2834:Chiroptera
2790:(Elephants)
2780:Hyracoidea
2536:(unranked)
2143:: 193–200.
1529:Purgatorius
1432:Ptilocercus
1393:T. salatana
1339:T. longipes
1330:T. javanica
1312:T. gracilis
1285:T. everetti
1276:T. dorsalis
1267:T. discolor
1219:D. melanura
1080:Haplorrhini
647:Insectivora
508:, which is
450:T. javanica
366:Description
300:. They are
290:Insectivora
244:tree shrews
71:Appendix II
5237:Treeshrews
5231:Categories
5048:Scandentia
5034:Scandentia
5004:Scandentia
4962:See also:
4918:Haplorhini
4907:Haplorhini
4718:Torrejonia
4704:Premnoides
4690:Phoxomylus
4607:Acidomomys
4545:Uintasorex
4538:Niptomomys
4531:Navajovius
4517:Microsyops
4496:Choctawius
4452:Picromomys
4415:Micromomys
4371:Ursolestes
4222:see below↓
4147:Plagiomene
4116:Mixodectes
4080:Dermoptera
4049:Sivatupaia
4019:Dendrogale
3979:Scandentia
3969:Euarchonta
3905:Haltictops
3898:Cartictops
3688:Interogale
3646:Anaptogale
3623:Anagalidae
3553:Zanycteris
3539:Draconodus
3494:Jepsenella
3487:Heterohyus
3473:Carcinella
3452:Aethomylos
3400:Subclass:
3222:T. montana
3061:Dendrogale
3048:A. ellioti
2850:Carnivora
2842:Pholidota
2774:(Aardvark)
2748:Afrotheria
2727:Cingulata
2708:inclusive)
2630:(Opossums)
2601:inclusive)
2591:Metatheria
2548:Yinotheria
2532:Vertebrata
2530:Subphylum
1912:2018-08-26
1601:2022-01-14
1535:References
1357:T. montana
1209:Dendrogale
1198:A. ellioti
1038:(colugos)
1036:Dermoptera
1003:Scandentia
990:Euarchonta
970:(rodents)
952:Lagomorpha
915:Dermoptera
832:(Colugos)
830:Dermoptera
788:(rodents)
770:Lagomorpha
744:Scandentia
707:superorder
699:lagomorphs
691:Euarchonta
688:grandorder
621:Scandentia
573:omnivorous
267:Scandentia
240:treeshrews
152:Scandentia
142:Euarchonta
35:Treeshrews
18:Scandentia
4859:Subengius
4852:Saxonella
4642:Paromomys
4628:Elwynella
4621:Edworthia
4575:Toliapina
4524:Nanomomys
4489:Berruvius
4482:Bartelsia
4422:Tinimomys
4408:Foxomomys
4401:Dryomomys
4161:Worlandia
4140:Ellesmene
4003:Tupaiidae
3884:Allictops
3861:Zhujegale
3854:Yupingale
3723:Zofiagale
3660:Diacronus
3515:Unuchinia
3466:Asiapator
3382:Kingdom:
3081:D. murina
3027:Tupaiidae
2919:Primates
2913:(Colugos)
2887:Rodentia
2782:(Hyraxes)
2719:Xenarthra
2706:Placental
2599:Marsupial
2449:: 51–60.
2263:1932-6203
2196:0962-8436
2056:0036-8075
2004:1749-4877
1698:1 January
1596:cites.org
1228:D. murina
1177:Tupaiidae
1151:Tupaiidae
666:prosimian
628:Tupaiidae
595:capsaicin
578:Rafflesia
546:nocturnal
534:territory
510:binocular
354:word for
338:The name
328:hepatitis
302:omnivores
275:Tupaiidae
248:banxrings
210:Tupaiidae
198:Tupaiidae
173:Tupaiidae
165:Families
108:Kingdom:
102:Eukaryota
5175:11900001
5055:BioLib:
5013:Wikidata
4979:Category
4875:Primates
4669:Anasazia
4635:Ignacius
4503:Craseops
4012:Anathana
3891:Anictops
3847:Astigale
3716:Wanogale
3695:Linnania
3667:Eosigale
3546:Picrodus
3459:Apatemys
3396:Mammalia
3390:Chordata
3388:Phylum:
3384:Animalia
3331:Category
3313:P. lowii
3249:T. picta
3204:T. minor
3037:Anathana
3011:Eutheria
3005:Mammalia
2999:Chordata
2993:Animalia
2991:Kingdom
2796:Sirenia
2698:Eutheria
2526:Chordata
2520:Animalia
2518:Kingdom
2465:85271080
2420:26766238
2358:21565274
2320:17307271
2281:23560065
2233:PLOS ONE
2214:27325836
2157:24333622
2072:38120449
2064:21940861
2012:25311886
1965:12251814
1957:17975064
1870:18663222
1806:NBC News
1693:16910445
1685:12823622
1578:62265494
1523:See also
1478:Pliocene
1442:P. lowii
1384:T. picta
1348:T. minor
1188:Anathana
1068:Primates
968:Rodentia
869:Primates
786:Rodentia
651:Primates
625:families
612:Taxonomy
602:mutation
528:Behavior
487:overall
465:T. minor
356:squirrel
320:primates
271:families
201:, Blue:
132:Mammalia
122:Chordata
118:Phylum:
112:Animalia
98:Domain:
42:– Recent
5019:Q231550
4568:Avenius
3926:Suyinia
3702:Qipania
3632:Anagale
3394:Class:
3276:T. tana
3159:T. glis
2997:Phylum
2975:Extant
2735:Pilosa
2538:Amniota
2524:Phylum
2505:Extant
2411:4725336
2388:Bibcode
2272:3613385
2241:Bibcode
2205:4920340
2122:2575767
2102:Bibcode
2036:Bibcode
2028:Science
1937:Bibcode
1929:Science
1861:2492458
1838:Bibcode
1776:440–445
1470:Miocene
1419:Family
1411:T. tana
1303:T. glis
1175:Family
1164:Urogale
1143:species
703:rodents
658:anatomy
636:species
581:fruit.
538:diurnal
498:3.1.3.3
495:2.1.3.3
435:T. glis
252:mammals
148:Order:
128:Class:
73: (
5201:574734
5162:552291
5110:1SCANO
4873:Crown-
4614:Arcius
4056:Tupaia
3937:Glires
3402:Theria
3094:Tupaia
3003:Class
2836:(Bats)
2582:Theria
2509:orders
2507:mammal
2463:
2418:
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1495:, and
1474:Yunnan
1429:Genus
1239:Tupaia
1236:Genus
1206:Genus
1185:Genus
1169:Tupaia
1147:genera
939:Glires
757:Glires
695:Glires
640:genera
506:vision
376:Tupaia
350:, the
341:Tupaia
324:myopia
313:humans
294:shrews
273:: the
159:, 1855
157:Wagner
59:Tupaia
5214:40294
5149:12640
5144:IRMNG
5136:46280
5058:31936
2461:S2CID
2090:(PDF)
2068:S2CID
1961:S2CID
1689:S2CID
1661:(PDF)
1552:. In
1466:Henan
911:basal
680:order
670:clade
655:brain
618:order
599:TRPV1
561:urine
551:They
352:Malay
347:tupai
264:order
195:Red:
75:CITES
69:CITES
5188:9392
5183:NCBI
5157:ITIS
5118:GBIF
5105:EPPO
5097:8705
5066:BOLD
4916:see
4893:see
2416:PMID
2354:PMID
2316:PMID
2277:PMID
2259:ISSN
2210:PMID
2192:ISSN
2153:PMID
2118:PMID
2060:PMID
2052:ISSN
2008:PMID
2000:ISSN
1953:PMID
1884:link
1866:PMID
1780:ISBN
1727:ISBN
1700:2012
1681:PMID
1574:OCLC
1564:ISBN
701:and
567:Diet
334:Name
238:The
213:and
62:sp.
5170:MSW
5123:801
5092:EoL
5084:426
5079:CoL
5071:349
5043:ADW
2451:doi
2447:131
2406:PMC
2396:doi
2346:doi
2308:doi
2267:PMC
2249:doi
2200:PMC
2184:doi
2180:371
2145:doi
2110:doi
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2044:doi
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1992:doi
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