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Sclerocarya birrea

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and elephants all browse on the marula tree, with elephants in particular being a major consumer. Elephants eat the bark, branches and fruits of the marula, which may limit the spread of the trees. The damaged bark, due to browsing, can be used to identify marula trees as elephants preferentially
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On an industrial level the fruit of the marula tree is collected from the wild by members of rural communities on whose land the trees grow. This harvest and sale of fruit only occur over two to three months, but is an important source of income to poor rural people, especially women. The fruit is
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The tree is a single-stemmed species with a broad, spreading crown. It is distinguished by its grey mottled bark and can grow up to 18 meters tall, primarily in low altitudes and open woodlands. The distribution of this species throughout Africa and Madagascar has followed the
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from eating fermented marula fruit, as do reports in the popular press. While the fruit is commonly eaten by elephants, the animals would need a huge amount of fermented marulas to have any effect on them, and other animals prefer the ripe fruit.
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with a single seed encased within their endocarp, although up to four seeds can be present. The seeds have a delicate nutty flavour and are much sought-after, especially by small rodents who know to gnaw exactly where the plugs are located.
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Leakey, Roger; Shackleton, Sheona; Du Plessis, Pierre (2005). "Domestication potential of Marula (Sclerocarya birrea subsp caffra) in South Africa and Namibia: 1. Phenotypic variation in fruit traits".
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The fruit is traditionally used for food in Africa, and has considerable socioeconomic importance. The fruit juice and pulp are mixed with water and stored in a container over 1–3 days of
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Mansourian, Suzan; Enjin, Anders V.; Jirle, Erling; Ramesh, Vedika; Rehermann, Guillermo; Becher, Paul G.; Pool, John E.; Stensmyr, Marcus C. (December 6, 2018).
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in their migrations. There is some evidence of human domestication of marula trees, as trees found on farm lands tend to have larger fruit size.
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delivered to processing plants where fruit pulp, pips, kernels and kernel oil are extracted and stored for processing throughout the year.
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is derived from the Ancient Greek words 'skleros' meaning 'hard' and 'karyon' meaning 'nut'. This refers to the hard pit of the fruit. The
1134:"Livelihood benefits from the local level commercialization of savanna resources: a case study of the new and expanding trade in marula ( 305: 1876: 1801: 889: 881: 863: 410:). They fall to the ground when unripe and green in colour, and then ripen to a yellow colour on the ground. They are succulent and 1861: 1851: 634: 1866: 1724: 1494: 1836: 1821: 1811: 1512: 562:. The alcoholic distilled beverage (morula) made from the fruit is referenced in the stories of the South African writer 1831: 1806: 1133: 630: 606: 422:). These stones, when dry, expose the seeds by shedding 2 (sometimes 3) small circular plugs at one end. The fruits are 1403: 1390: 1058:
Engelter & Wehmeyer; Wehmeyer, A.S. (1970). "Fatty acid composition of oils of some edible seeds of wild plants".
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The marula fruit has been suggested to be the food of choice for the ancestral forest-dwelling form of the fruit fly
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Common names include jelly plum, cat thorn, morula, cider tree, marula, maroola nut/plum, and in
248: 97: 1564: 1037: 963:. Department of Water Affairs and Forestry, Republic of South Africa. 3 May 2013. Archived from 1525: 1685: 1546: 1416: 1290: 1190: 1075: 1013: 1003: 895: 885: 655: 1626: 1551: 1280: 1272: 1182: 1067: 995: 836: 447: 316: 1118: 747: 647:, which was much more selective about which fruit they preferred than the flies that have 367: 236: 140: 705: 1577: 1268: 832: 1572: 1285: 1255: 1246: 501: 127: 1785: 1520: 756: 509: 396: 186: 87: 82: 1322: 1202: 989: 848: 1768: 1690: 1639: 1421: 555: 348:, "nut", in reference to the stone inside the fleshy fruit), commonly known as the 310: 1395: 1382: 17: 1677: 1666: 604:
target them. Elephants distribute marula seeds in their dung. In the documentary
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These subspecies are differentiated by changes in leaf shape and size. Subsp.
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with a strong and distinctive flavour. Inside is a walnut-sized, thick-walled
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The fruits, which ripen between December and March, have a light yellow skin (
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to live near to humans. The ancestral fruit flies are triggered by the
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with emphasis on its importance as a NTFP in South and Southern Africa
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The marula fruit is eaten by various animals in Southern Africa.
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in Southern Africa. The marula tree is protected in South Africa.
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Marula oil for sale at Ongwediva Annual Trade Fair 2016, Namibia
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to moisturise the skin, and as an edible oil in the diet of
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with toothed margins, features not present in adult plants
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per flower. On rare occasion a male flower can produce a
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Morris, Steve; Humphreys, David; Reynolds, Dan (2006).
1650: 1340: 438:. These have red sepals and petals, and about 20 8: 742:IUCN SSC Global Tree Specialist Group & 1328: 1060:Journal of Agricultural and Food Chemistry 923: 921: 919: 917: 915: 744:Botanic Gardens Conservation International 76: 49: 40: 31: 1284: 1251:Are Seasonal Specialists on Marula Fruit" 1157: 1155: 876:. Non-Wood Forest Products. Vol. 5. 508:. The marula belongs to the same family, 469:is divided into three subspecies: subsp. 991:Lost Crops of Africa: Volume III: Fruits 614:, released in 1974, some scenes portray 489:is found in southern Africa, and subsp. 1038:"The marvellous brewers of marula beer" 734: 665: 1138:) beer in Bushbuckridge, South Africa" 1114: 1104: 1093:Shackleton, S.E.; et al. (2002). 952: 950: 675:Marula windfalls in Ongwediva, Namibia 528:, and is closely related to the genus 1323:Swazisecrets.com: Marula African oils 1175:Physiological and Biochemical Zoology 7: 1318:West African plants – A Photo Guide. 485:is found in northern Africa, subsp. 1792:IUCN Red List least concern species 757:IUCN Red List of Threatened Species 558:to make marula beer, a traditional 1148:(11): 651–657 – via ingenta. 25: 882:Food and Agriculture Organization 864:Food and Agriculture Organization 1142:South African Journal of Science 716: 704: 692: 687:Marula cider, Ongwediva, Namibia 680: 668: 101: 1842:Protected trees of South Africa 1743:urn:lsid:ipni.org:names:71444-1 1604:urn:lsid:ipni.org:names:71162-1 370:, the Sudano-Sahelian range of 1223:Couper, Ross (24 April 2014). 1: 1872:Taxa named by Achille Richard 1827:Fruits originating in Africa 607:Animals Are Beautiful People 1132:Sheona, Shackleton (2004). 493:is only found in Tanzania. 1893: 1095:A summary of knowledge on 1040:. Eco products. 2016-03-14 798:Royal Botanic Gardens, Kew 793:Plants of the World Online 341: 333: 1277:10.1016/j.cub.2018.10.033 982:National Research Council 841:10.1007/s10457-005-2419-z 254: 247: 225: 218: 98:Scientific classification 96: 74: 65: 61:with and without foliage 57: 48: 39: 34: 1877:Plants described in 1831 1802:Flora of Southern Africa 1000:National Academies Press 764:: e.T208166817A208357864 1862:Drought-tolerant plants 1852:Garden plants of Africa 1249:Drosophila melanogaster 868:"Potential Edible Nuts" 644:Drosophila melanogaster 337:⟨sklērós⟩ 1867:Drought-tolerant trees 462:Taxonomy and etymology 346:⟨káryon⟩ 339:, meaning "hard", and 320: 302: 294: 286: 1837:Trees of South Africa 1822:Edible nuts and seeds 1812:Flora of South Africa 658:in the marula fruit. 591:Uses by other species 564:Herman Charles Bosman 406:), with white flesh ( 308: 300: 292: 284: 1263:(24): 3960–3968.e3. 821:Agroforestry Systems 362:, indigenous to the 352:, is a medium-sized 1832:Trees of Madagascar 1807:Flora of Madagascar 1269:2018CBio...28E3960M 1072:10.1021/jf60167a025 833:2005AgrSy..64...25L 68:Conservation status 1565:Sclerocarya birrea 1372:Sclerocarya birrea 1342:Sclerocarya birrea 1313:Sclerocarya birrea 1166:Loxodonta africana 1136:Sclerocarya birrea 1097:Sclerocarya birrea 931:Sclerocarya birrea 786:Sclerocarya birrea 750:Sclerocarya birrea 560:alcoholic beverage 467:Sclerocarya birrea 325:Sclerocarya birrea 321: 303: 301:Green marula fruit 295: 287: 229:Sclerocarya birrea 18:Sclerocarya caffra 1779: 1778: 1547:Open Tree of Life 1334:Taxon identifiers 1009:978-0-309-10596-5 958:"Protected Trees" 788:(A.Rich.) Hochst" 656:ethyl isovalerate 649:self-domesticated 534:from Madagascar. 496:The generic name 279: 278: 273: 262: 91: 16:(Redirected from 1884: 1857:Ornamental trees 1847:Dioecious plants 1772: 1771: 1759: 1758: 1746: 1745: 1733: 1732: 1720: 1719: 1707: 1706: 1694: 1693: 1681: 1680: 1671: 1670: 1669: 1643: 1642: 1630: 1629: 1620: 1619: 1607: 1606: 1594: 1593: 1581: 1580: 1568: 1567: 1555: 1554: 1542: 1541: 1529: 1528: 1516: 1515: 1503: 1502: 1490: 1489: 1477: 1476: 1464: 1463: 1451: 1450: 1438: 1437: 1425: 1424: 1412: 1411: 1399: 1398: 1386: 1385: 1376: 1375: 1374: 1361: 1360: 1359: 1329: 1299: 1298: 1288: 1242: 1236: 1235: 1233: 1231: 1220: 1214: 1213: 1211: 1209: 1172: 1159: 1150: 1149: 1129: 1123: 1122: 1116: 1112: 1110: 1102: 1090: 1084: 1083: 1055: 1049: 1048: 1046: 1045: 1034: 1028: 1027: 1025: 1024: 996:Washington, D.C. 978: 972: 971: 969: 962: 954: 945: 944: 942: 940: 925: 910: 909: 907: 906: 862:Wickens, G. E.; 859: 853: 852: 815: 809: 808: 806: 804: 780: 774: 773: 771: 769: 739: 720: 708: 696: 684: 672: 550:Traditional uses 502:specific epithet 347: 344: 343: 338: 335: 268: 266:Poupartia birrea 260: 231: 106: 105: 85: 80: 79: 53: 44: 32: 21: 1892: 1891: 1887: 1886: 1885: 1883: 1882: 1881: 1782: 1781: 1780: 1775: 1767: 1762: 1754: 1749: 1741: 1736: 1728: 1723: 1715: 1710: 1702: 1697: 1689: 1684: 1676: 1674: 1665: 1664: 1659: 1652:Spondias birrea 1646: 1638: 1633: 1625: 1623: 1615: 1610: 1602: 1597: 1589: 1584: 1576: 1571: 1563: 1558: 1550: 1545: 1537: 1532: 1524: 1519: 1511: 1506: 1498: 1493: 1485: 1480: 1472: 1467: 1459: 1454: 1446: 1441: 1433: 1428: 1420: 1415: 1407: 1402: 1394: 1389: 1381: 1379: 1370: 1369: 1364: 1355: 1354: 1349: 1336: 1308: 1303: 1302: 1244: 1243: 1239: 1229: 1227: 1222: 1221: 1217: 1207: 1205: 1170: 1161: 1160: 1153: 1131: 1130: 1126: 1113: 1103: 1092: 1091: 1087: 1057: 1056: 1052: 1043: 1041: 1036: 1035: 1031: 1022: 1020: 1010: 994:. Vol. 3. 980: 979: 975: 970:on 5 July 2010. 967: 960: 956: 955: 948: 938: 936: 927: 926: 913: 904: 902: 892: 861: 860: 856: 817: 816: 812: 802: 800: 782: 781: 777: 767: 765: 746:(BGCI) (2024). 741: 740: 736: 731: 724: 721: 712: 709: 700: 697: 688: 685: 676: 673: 664: 593: 584: 582:Commercial uses 552: 547: 479:multifoliolata. 464: 392: 368:Southern Africa 345: 336: 258:Spondias birrea 243: 233: 227: 214: 100: 92: 81: 77: 70: 28: 27:Species of tree 23: 22: 15: 12: 11: 5: 1890: 1888: 1880: 1879: 1874: 1869: 1864: 1859: 1854: 1849: 1844: 1839: 1834: 1829: 1824: 1819: 1814: 1809: 1804: 1799: 1794: 1784: 1783: 1777: 1776: 1774: 1773: 1769:wfo-0000435345 1760: 1747: 1734: 1721: 1708: 1695: 1682: 1672: 1656: 1654: 1648: 1647: 1645: 1644: 1640:wfo-0000434908 1631: 1621: 1608: 1595: 1582: 1569: 1556: 1543: 1530: 1517: 1504: 1491: 1478: 1465: 1452: 1439: 1426: 1413: 1400: 1387: 1377: 1362: 1346: 1344: 1338: 1337: 1332: 1326: 1325: 1320: 1307: 1306:External links 1304: 1301: 1300: 1247:"Wild African 1237: 1215: 1187:10.1086/499983 1181:(2): 363–369. 1151: 1124: 1085: 1050: 1029: 1008: 984:(2008-01-25). 973: 946: 911: 890: 854: 810: 775: 733: 732: 730: 727: 726: 725: 722: 715: 713: 710: 703: 701: 698: 691: 689: 686: 679: 677: 674: 667: 663: 660: 592: 589: 583: 580: 551: 548: 546: 543: 491:multifoliolata 463: 460: 430:The trees are 391: 388: 293:Female flowers 277: 276: 275: 274: 263: 252: 251: 245: 244: 234: 223: 222: 216: 215: 211:S. birrea 208: 206: 202: 201: 194: 190: 189: 184: 180: 179: 174: 170: 169: 164: 157: 156: 151: 144: 143: 138: 131: 130: 125: 118: 117: 112: 108: 107: 94: 93: 75: 72: 71: 66: 63: 62: 55: 54: 46: 45: 37: 36: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 1889: 1878: 1875: 1873: 1870: 1868: 1865: 1863: 1860: 1858: 1855: 1853: 1850: 1848: 1845: 1843: 1840: 1838: 1835: 1833: 1830: 1828: 1825: 1823: 1820: 1818: 1817:Desert fruits 1815: 1813: 1810: 1808: 1805: 1803: 1800: 1798: 1795: 1793: 1790: 1789: 1787: 1770: 1765: 1761: 1757: 1752: 1748: 1744: 1739: 1735: 1731: 1726: 1722: 1718: 1713: 1709: 1705: 1700: 1696: 1692: 1687: 1683: 1679: 1673: 1668: 1662: 1658: 1657: 1655: 1653: 1649: 1641: 1636: 1632: 1628: 1622: 1618: 1613: 1609: 1605: 1600: 1596: 1592: 1587: 1583: 1579: 1574: 1570: 1566: 1561: 1557: 1553: 1548: 1544: 1540: 1535: 1531: 1527: 1522: 1518: 1514: 1509: 1505: 1501: 1496: 1492: 1488: 1483: 1479: 1475: 1470: 1466: 1462: 1457: 1453: 1449: 1444: 1440: 1436: 1431: 1427: 1423: 1418: 1414: 1410: 1405: 1401: 1397: 1392: 1388: 1384: 1378: 1373: 1367: 1363: 1358: 1352: 1348: 1347: 1345: 1343: 1339: 1335: 1330: 1324: 1321: 1319: 1315: 1314: 1310: 1309: 1305: 1296: 1292: 1287: 1282: 1278: 1274: 1270: 1266: 1262: 1258: 1257: 1252: 1250: 1241: 1238: 1226: 1219: 1216: 1204: 1200: 1196: 1192: 1188: 1184: 1180: 1176: 1169: 1167: 1158: 1156: 1152: 1147: 1143: 1139: 1137: 1128: 1125: 1120: 1108: 1100: 1096: 1089: 1086: 1081: 1077: 1073: 1069: 1065: 1061: 1054: 1051: 1039: 1033: 1030: 1019: 1015: 1011: 1005: 1001: 997: 993: 992: 987: 983: 977: 974: 966: 959: 953: 951: 947: 935:. 15 May 2018 934: 932: 924: 922: 920: 918: 916: 912: 901: 897: 893: 891:92-5-103748-5 887: 883: 879: 875: 874: 869: 865: 858: 855: 850: 846: 842: 838: 834: 830: 826: 822: 814: 811: 799: 795: 794: 789: 787: 779: 776: 763: 759: 758: 753: 751: 745: 738: 735: 728: 719: 714: 707: 702: 699:Marula stones 695: 690: 683: 678: 671: 666: 661: 659: 657: 654: 650: 646: 645: 639: 636: 632: 629: 625: 621: 617: 613: 609: 608: 602: 598: 590: 588: 581: 579: 577: 573: 569: 565: 561: 557: 549: 544: 542: 540: 535: 533: 532: 527: 523: 519: 515: 511: 510:Anacardiaceae 507: 503: 499: 494: 492: 488: 484: 480: 476: 472: 468: 461: 459: 457: 453: 449: 445: 441: 437: 433: 428: 425: 421: 417: 413: 409: 405: 400: 398: 389: 387: 385: 381: 378:woodlands of 377: 373: 369: 366:woodlands of 365: 361: 358: 357:fruit-bearing 355: 351: 331: 330:Ancient Greek 327: 326: 318: 315: 312: 309:Sapling with 307: 299: 291: 283: 272: 267: 264: 259: 256: 255: 253: 250: 246: 242: 238: 232: 230: 224: 221: 220:Binomial name 217: 213: 212: 207: 204: 203: 200: 199: 195: 192: 191: 188: 187:Anacardiaceae 185: 182: 181: 178: 175: 172: 171: 168: 165: 162: 159: 158: 155: 152: 149: 146: 145: 142: 139: 136: 133: 132: 129: 128:Tracheophytes 126: 123: 120: 119: 116: 113: 110: 109: 104: 99: 95: 89: 84: 83:Least Concern 73: 69: 64: 60: 56: 52: 47: 43: 38: 33: 30: 19: 1651: 1341: 1312: 1260: 1254: 1248: 1240: 1228:. Retrieved 1218: 1206:. Retrieved 1178: 1174: 1165: 1145: 1141: 1135: 1127: 1098: 1094: 1088: 1066:(1): 25–26. 1063: 1059: 1053: 1042:. Retrieved 1032: 1021:. Retrieved 990: 976: 965:the original 937:. Retrieved 930: 903:. Retrieved 872: 857: 827:(1): 25–35. 824: 820: 813: 801:. Retrieved 791: 785: 778: 766:. Retrieved 761: 755: 749: 737: 711:Marula seeds 642: 640: 605: 601:rhinoceroses 594: 585: 556:fermentation 553: 536: 529: 497: 495: 490: 486: 482: 478: 474: 470: 466: 465: 429: 401: 393: 349: 324: 323: 322: 285:Marula trunk 265: 257: 228: 226: 210: 209: 197: 160: 147: 134: 121: 58: 29: 1797:Sclerocarya 1578:kew-2479830 1482:iNaturalist 1366:Wikispecies 1115:|work= 873:Edible Nuts 635:intoxicated 541:, maroela. 498:Sclerocarya 477:and subsp. 390:Description 380:East Africa 372:West Africa 311:distinctive 269:(A. Rich.) 198:Sclerocarya 141:Angiosperms 1786:Categories 1573:Plant List 1208:26 October 1044:2019-02-14 1023:2008-07-17 905:2008-11-10 729:References 576:San people 568:Marula oil 456:elliptical 452:staminodes 384:Madagascar 314:emarginate 261:(A. Rich.) 177:Sapindales 1667:Q38423597 1526:208166817 1117:ignored ( 1107:cite book 633:becoming 631:allegedly 620:ostriches 616:elephants 612:Jamie Uys 572:topically 539:Afrikaans 531:Poupartia 522:pistachio 512:, as the 473:, subsp. 450:and have 444:gynoecium 432:dioecious 354:deciduous 205:Species: 111:Kingdom: 59:S. birrea 1756:50091089 1751:Tropicos 1661:Wikidata 1612:Tropicos 1351:Wikidata 1295:30528579 1230:25 April 1203:36629801 1195:16555195 1018:34344933 986:"Marula" 939:19 April 900:34529770 866:(1995). 849:21601141 624:warthogs 597:Giraffes 570:is used 420:endocarp 408:mesocarp 317:leaflets 249:Synonyms 183:Family: 154:Eudicots 88:IUCN 3.1 1730:71444-1 1704:3661572 1627:5854675 1617:1300356 1500:71162-1 1461:7160810 1357:Q675778 1286:7065024 1265:Bibcode 1080:5524461 829:Bibcode 803:27 July 768:27 July 662:Gallery 628:baboons 506:Senegal 448:pedicel 440:stamens 404:exocarp 376:savanna 334:σκληρός 271:Aubrév. 241:Hochst. 237:A.Rich. 193:Genus: 173:Order: 115:Plantae 86: ( 35:Marula 1675:APDB: 1624:uBio: 1586:PLANTS 1552:456294 1539:289766 1513:895125 1487:340245 1435:483490 1409:416922 1396:222419 1383:174216 1380:APDB: 1293:  1283:  1201:  1193:  1078:  1016:  1006:  898:  888:  847:  518:cashew 483:birrea 471:birrea 436:raceme 424:drupes 374:, the 364:miombo 350:marula 342:κάρυον 167:Rosids 1717:35329 1691:6ZCNL 1678:30812 1591:SCBI3 1474:33385 1448:SLCBB 1422:4VP9L 1199:S2CID 1171:(PDF) 968:(PDF) 961:(PDF) 845:S2CID 653:ester 526:sumac 514:mango 416:stone 397:Bantu 161:Clade 148:Clade 135:Clade 122:Clade 1738:POWO 1725:IPNI 1712:GRIN 1699:GBIF 1599:POWO 1560:PfaF 1534:NCBI 1521:IUCN 1508:ITIS 1495:IPNI 1469:GRIN 1456:GBIF 1443:EPPO 1404:BOLD 1391:APNI 1291:PMID 1256:Cell 1232:2014 1210:2015 1191:PMID 1119:help 1076:PMID 1014:OCLC 1004:ISBN 941:2019 896:OCLC 886:ISBN 878:Rome 805:2024 770:2024 762:2024 626:and 545:Uses 524:and 487:afra 475:afra 412:tart 382:and 360:tree 1764:WFO 1686:CoL 1635:WFO 1430:EoL 1417:CoL 1316:in 1281:PMC 1273:doi 1183:doi 1146:100 1068:doi 837:doi 610:by 1788:: 1766:: 1753:: 1740:: 1727:: 1714:: 1701:: 1688:: 1663:: 1637:: 1614:: 1601:: 1588:: 1575:: 1562:: 1549:: 1536:: 1523:: 1510:: 1497:: 1484:: 1471:: 1458:: 1445:: 1432:: 1419:: 1406:: 1393:: 1368:: 1353:: 1289:. 1279:. 1271:. 1261:28 1259:. 1253:. 1197:. 1189:. 1179:79 1177:. 1173:. 1154:^ 1144:. 1140:. 1111:: 1109:}} 1105:{{ 1074:. 1064:18 1062:. 1012:. 1002:. 998:: 988:. 949:^ 914:^ 894:. 884:. 880:: 870:. 843:. 835:. 825:64 823:. 796:. 790:. 760:. 754:. 622:, 618:, 599:, 566:. 520:, 516:, 458:. 386:. 332:: 239:) 163:: 150:: 137:: 124:: 1297:. 1275:: 1267:: 1234:. 1212:. 1185:: 1121:) 1101:. 1082:. 1070:: 1047:. 1026:. 943:. 933:" 929:" 908:. 851:. 839:: 831:: 807:. 784:" 772:. 752:" 748:" 418:( 328:( 235:( 90:) 20:)

Index

Sclerocarya caffra


Conservation status
Least Concern
IUCN 3.1
Scientific classification
Edit this classification
Plantae
Tracheophytes
Angiosperms
Eudicots
Rosids
Sapindales
Anacardiaceae
Sclerocarya
Binomial name
A.Rich.
Hochst.
Synonyms
Aubrév.




distinctive
emarginate
leaflets
Ancient Greek
deciduous

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