739:
725:
321:
46:
224:
608:
594:
33:
340:
284:
357:
302:
257:
262:
652:. The stem is 4–10 cm (1.6–3.9 in) long by 2–4 cm (0.8–1.6 in) thick, and is equal in width throughout its length, slightly thicker towards the base, or somewhat thicker in the middle. It is the same color as the cap, or paler. The surface texture of the stem is usually smooth, although some specimens may be slightly reticulated near the top. The
239:
671:
found on the tube faces) are irregularly club-shaped, with dimensions of 20–36 by 7–10 μm, while cheilocystidia (found on the tube edge) are club-shaped, rare, occur singly, and measure 18–32 by 7–9 μm. Although rare, there are also caulocystidia (occurring on the stem) that are arranged in
425:
that measures up to 15 cm (5.9 in) in diameter. The caps of young specimens have a velvety texture and are covered with a whitish to gray powdery coating; this texture and coating is gradually lost as the mushroom matures, and the cap often develops cracks. The pores on the underside of the
647:
that are arranged in a vertically arranged layer of minute tubes on the underside of the cap that create a surface of pores. This surface is whitish when young before turning dull pink or flesh-colored in maturity. When bruised, the pore surface initially stains reddish and slowly turns black. The
629:(developing a network of cracks and small crevices). The cap color is initially black to dark grayish brown; young specimens can have a whitish bloom (resembling a dusting of fine powder) on the surface. Fruit bodies, especially young specimens, tend to be free of
624:
is initially convex before later becoming broadly convex to eventually flattened in maturity; the diameter of the cap is typically between 3 and 15 cm (1.2 and 5.9 in). The cap surface is dry, with a velvet-like texture, although in age it can become
698:—a genus that is usually associated with bitter-tasting, unpalatable species. Frying slices of the mushroom brings out a "delicate, earthy, nutty flavor"; longer frying times make the cap "pleasantly crisp". The mushrooms can be used in
783:
produces smaller fruit bodies with caps up to 9 cm (3.5 in) in diameter, and its whitish flesh directly stains black without any intermediate reddish phase when injured. It is known from only from western
648:
shape of the pores is angular to irregular, and they are small, with roughly two pores per millimeter. The tubes are 5–10 mm (0.2–0.4 in) deep, and usually sunken around the area of attachment to the
1622:
546:
in 1916, and noted that Peck's description was made from young material obtained "before the white tubes had been colored by mature spores". Several later authorities have treated Peck's species as a
1867:
430:
is bluish purple to black, and measures up to 10 cm (3.9 in) long by 4 cm (1.6 in) thick. Both the pore surface and the whitish cap
1957:
1815:
223:
1439:
1123:
828:, but the latter has yellow cystidia when mounted in KOH, while the cystidia of the former are brownish yellow under similar conditions.
633:
and other insect larvae. As the mushroom matures, the bloom disappears and the color fades to become grayish to grayish brown. The cap
1005:. Schriften der naturforschenden Gesellschaft in Leipzig (in Latin). Vol. 1. Johann Ambrosius Barth. pp. 20–131 (see p. 95).
1685:
1496:
1469:
1365:
983:
1570:
2008:
2018:
919:
812:
by the strong orange to red discoloration that results when cutting or damaging the flesh of a fresh specimen. Furthermore,
843:
species, and its fruit bodies grow on the ground solitarily, scattered, or in groups under deciduous trees, particularly
1221:
Singer R. (1947). "The
Boletoideae of Florida. The Boletineae of Florida with notes on extralimital species. III".
45:
1872:
1648:
493:
459:
763:
498:
161:
2023:
775:
1767:
637:
is whitish, but after it is cut or injured, it will stain pink to reddish gray, and eventually turn black.
2013:
1729:
801:
570:
508:
2003:
1718:
140:
434:
will stain pink to reddish gray, and eventually turn black after being cut or injured. The mushroom is
1949:
1807:
1789:
1702:
1287:
1183:
1859:
535:
527:
451:
738:
724:
1998:
1714:
1546:
1238:
1203:
1140:
1065:
805:
555:
467:
170:
40:
667:. The basidia are club-shaped, four-spored, and measure 15–24 by 6–7.5 μm. Pleurocystidia (
1115:
1931:
1880:
1776:
1681:
1492:
1465:
1435:
1431:
1424:
1404:
1361:
1355:
1316:
1269:
1119:
1088:
979:
973:
938:
852:
769:
621:
547:
539:
422:
407:
267:
1895:
1486:
1000:
1885:
1538:
1230:
1195:
1107:
1057:
848:
673:
649:
634:
613:
One hour after bisection, this fruit body shows the black staining that eventually develops.
431:
427:
403:
307:
607:
593:
32:
863:
691:
470:
435:
391:
363:
1596:
1944:
1802:
1708:
1295:
785:
559:
463:
399:
87:
694:
with a pleasant odor and a mild taste. It is considered one of the best of the edible
1992:
1108:
77:
1781:
856:
699:
599:
The pore surface is whitish to dull pink, and initially stains reddish with injury.
411:
201:
1936:
1163:
398:
species, it grows solitarily, scattered, or in groups on the ground usually under
1922:
1854:
1828:
1761:
1032:
871:
653:
577:
482:
418:
347:
327:
190:
1752:
1020:(in Latin). Vol. 1. Lundin, Sweden: Ex Officiana Berlingiana. p. 127.
975:
The
Macrofungus Flora of China's Guangdong Province (Chinese University Press)
840:
664:
395:
387:
107:
1273:
1092:
580:
known as the "black velvet bolete"; Murrill called it the "blackish bolete".
1168:. North American Flora. Vol. 9. New York Botanical Garden. p. 135.
899:
503:
117:
97:
57:
1529:, a new species from the North American Atlantic and Gulf Coastal plain".
796:, and has stems that are minutely velvety and almost black near the base.
516:
1970:
1916:
1841:
1746:
1324:
946:
883:
879:
668:
563:
289:
248:
229:
157:
890:. It is also found in Mexico. In Asia, it has been recorded from China (
1962:
1820:
1550:
1242:
1207:
1069:
907:
903:
875:
660:
644:
487:
394:, and in eastern Asia, including China, Japan, Taiwan, and Thailand. A
820:, and typically has a more prominently reticulated stem. Specimens of
1975:
1846:
1833:
1360:. Lexington, Kentucky: University Press of Kentucky. pp. 337–8.
895:
867:
630:
383:
67:
1723:
1542:
1234:
1199:
1061:
1678:
North
American Boletes: A Color Guide to the Fleshy Pored Mushrooms
1430:. Ann Arbor, Michigan: The University of Michigan Press. pp.
891:
887:
677:
640:
862:
In North
America, the mushroom is widely distributed east of the
1794:
1491:. Syracuse, New York: Syracuse University Press. pp. 52–3.
1727:
978:. New York, New York: Columbia University Press. p. 485.
844:
1145:
Annual Report on the New York State Museum of
Natural History
672:
groups, and which measure individually 24–30 by 6–9 μm.
859:. Its dark color makes it difficult to notice in the field.
554:; this synonymy, however, is not indicated by either of the
1048:
Murrill WA. (1909). "The
Boletaceae of North America – 1".
1488:
The
Rainbow Beneath my Feet: A Mushroom Dyer's Field Guide
1357:
Mushrooms of West
Virginia and the Central Appalachians
1037:. Vol. 3. Leipzig, Germany: A. Felix. p. 535.
792:, has a brownish cap that lacks the velvety texture of
402:, particularly oak, although it has been recorded from
390:. The species is found in North America east of the
1906:
1736:
1106:Kirk PM, Cannon PF, Minter DW, Stalpers JA (2008).
659:The spores are oval to ellipsoid in shape, smooth,
511:transferred the species to his newly created genus
354:
337:
317:
299:
281:
254:
236:
217:
1571:"Four species of boletes newly recorded to Taiwan"
1423:
438:, and generally considered one of the best edible
1680:. Syracuse, New York: Syracuse University Press.
1464:. Buffalo, New York: Firefly Books. p. 282.
816:lacks the whitish bloom present on young caps of
1656:Songklanakarin Journal of Science and Technology
1114:(10th ed.). Wallingford, UK: CABI. p.
808:of North America, is readily distinguished from
1629:. Les champignons du Québec. www.mycoquebec.org
1455:
1453:
1451:
824:that are paler than usual can be confused with
1395:
1393:
1391:
1389:
1387:
1385:
1383:
1381:
1379:
1377:
1177:
1175:
663:(translucent), and measure 7–11 by 3.5–5
656:can range from pinkish to a deep flesh color.
1085:Les Livres du Mycologue, Tome III: Les Bolets
8:
1647:Seehanan S, Petcharat V, Te-chato S (2007).
1569:Chen C-M, Ho Y-S, Peng J-J, Lin T-C (2002).
1349:
1347:
1345:
1343:
1341:
967:
965:
963:
1564:
1562:
1560:
1724:
1676:Bessette AR, Bessette A, Roody WC (2000).
1462:Mushrooms and Other Fungi of North America
1417:
1415:
1413:
757:species have a superficial resemblance to
31:
20:
1260:. 1. Taxa described by Charles H. Peck".
1087:(in French). Paris, France: Le François.
761:and might be confused with it, including
1327:. International Mycological Association
949:. International Mycological Association
930:
1256:Wolfe CB Jr. (1981). "Type studies in
1002:Synopsis fungorum Carolinae superioris
214:
1165:(Agaricales) Polyporaceae–Agaricaceae
7:
972:Zhishu B, Zheng G, Taihui L (1993).
507:in 1909. In 1931, French mycologist
788:. The "false black velvet bolete",
14:
1525:Wolfe CB Jr, Halling RE (1989). "
477:. The species was one of several
1485:Bessette A, Bessette AR (2001).
910:), Japan, Taiwan, and Thailand.
737:
723:
606:
592:
355:
338:
319:
300:
282:
260:
255:
237:
222:
44:
1223:The American Midland Naturalist
1141:"Report of the Botanist (1875)"
866:. The distribution ranges from
576:means "white and black". It is
538:. Murrill reduced this name to
534:from specimens he collected in
515:, but this name has since been
462:from specimens he collected in
426:cap are small and pinkish. The
1578:Taiwan Journal of Biodiversity
1184:"Illustrations of fungi–XXXII"
920:List of North American boletes
421:have a black to grayish-brown
1:
501:transferred it to the genus
999:von Schweinitz LD. (1822).
230:Mycological characteristics
2040:
1649:"Some boletes of Thailand"
1595:Kuo M. (September 2003).
1034:Revisio Generum Plantarum
494:Revisio Generum Plantarum
460:Lewis David de Schweinitz
176:
169:
146:
139:
41:Scientific classification
39:
30:
23:
943:(Schwein.) Murrill 1909"
870:in Canada, south to the
832:Distribution and habitat
499:William Alphonso Murrill
2009:Fungi described in 1822
1658:(in Thai and English).
1621:Labbé R. (March 2010).
1527:Tylopileus griseocarnus
1110:Dictionary of the Fungi
2019:Fungi of North America
790:T. atronicotianus
764:T. atronicotianus
497:. American mycologist
473:this name in his 1821
450:The species was first
1719:New York State Museum
1288:"Homotypic synonyms:
1083:Gilbert J-E. (1931).
847:. Fruiting occurs in
814:T. griseocarneus
798:T. griseocarneus
776:T. griseocarneus
386:fungus in the family
197:Porphyrellus alboater
1603:. MushroomExpert.Com
1460:Phillips R. (2005).
1426:100 Edible Mushrooms
1182:Murrill WA. (1920).
1162:Murrill WA. (1916).
1151:: 29–82 (see p. 44).
878:, extending west to
676:are absent from the
509:Jean-Edouard Gilbert
1403:. (2000), pp.
1298:. CAB International
1018:Systema Mycologicum
826:T. ferrugineus
806:Gulf Coastal Plains
536:Sand Lake, New York
528:Charles Horton Peck
475:Systema Mycologicum
446:Taxonomy and naming
380:black velvet bolete
1768:Tylopilus alboater
1738:Tylopilus alboater
1715:Wax mushroom model
1703:Tylopilus alboater
1625:Tylopilus alboater
1599:Tylopilus alboater
1354:Roody WC. (2003).
1031:Kuntze O. (1898).
1016:Fries EM. (1821).
941:Tylopilus alboater
837:Tylopilus alboater
688:Tylopilus alboater
552:Tylopilus alboater
468:Elias Magnus Fries
382:, by some, is a
375:Tylopilus alboater
219:Tylopilus alboater
150:Tylopilus alboater
25:Tylopilus alboater
1986:
1985:
1881:Open Tree of Life
1730:Taxon identifiers
1627:/ Bolet noirâtre"
1441:978-0-472-03126-9
1319:Boletus nigrellus
1290:Boletus nigrellus
1139:Peck CH. (1878).
1125:978-0-85199-826-8
731:T. atronicotianus
674:Clamp connections
620:The shape of the
532:Boletus nigrellus
371:
370:
212:Species of fungus
210:
209:
205:
194:
183:
16:Species of fungus
2031:
1979:
1978:
1966:
1965:
1953:
1952:
1940:
1939:
1927:
1926:
1925:
1908:Boletus alboater
1899:
1898:
1889:
1888:
1876:
1875:
1863:
1862:
1850:
1849:
1837:
1836:
1824:
1823:
1811:
1810:
1798:
1797:
1785:
1784:
1772:
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1770:
1757:
1756:
1755:
1725:
1691:
1664:
1663:
1653:
1644:
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1634:
1618:
1612:
1611:
1609:
1608:
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1586:
1585:
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1566:
1555:
1554:
1522:
1516:
1509:
1503:
1502:
1482:
1476:
1475:
1457:
1446:
1445:
1429:
1419:
1408:
1397:
1372:
1371:
1351:
1336:
1335:
1333:
1332:
1313:
1307:
1306:
1304:
1303:
1284:
1278:
1277:
1253:
1247:
1246:
1218:
1212:
1211:
1179:
1170:
1169:
1159:
1153:
1152:
1136:
1130:
1129:
1113:
1103:
1097:
1096:
1080:
1074:
1073:
1045:
1039:
1038:
1028:
1022:
1021:
1013:
1007:
1006:
996:
990:
989:
969:
958:
957:
955:
954:
935:
822:T. alboater
818:T. alboater
810:T. alboater
794:T. alboater
759:T. alboater
745:T. griseocarneus
741:
727:
682:T. alboater
643:are produced in
610:
596:
571:specific epithet
544:T. alboater
456:Boletus alboater
359:
358:
342:
341:
323:
322:
304:
303:
286:
285:
264:
263:
259:
258:
241:
240:
226:
215:
199:
188:
186:Suillus alboater
181:
179:Boletus alboater
152:
132:T. alboater
49:
48:
35:
21:
2039:
2038:
2034:
2033:
2032:
2030:
2029:
2028:
1989:
1988:
1987:
1982:
1974:
1969:
1961:
1956:
1948:
1943:
1935:
1930:
1921:
1920:
1915:
1902:
1894:
1892:
1884:
1879:
1871:
1866:
1858:
1853:
1845:
1840:
1832:
1827:
1819:
1814:
1806:
1801:
1793:
1788:
1780:
1775:
1766:
1765:
1760:
1751:
1750:
1745:
1732:
1698:
1688:
1675:
1672:
1667:
1651:
1646:
1645:
1641:
1632:
1630:
1620:
1619:
1615:
1606:
1604:
1594:
1593:
1589:
1573:
1568:
1567:
1558:
1543:10.2307/3760072
1524:
1523:
1519:
1515:(2000), p. 259.
1510:
1506:
1499:
1484:
1483:
1479:
1472:
1459:
1458:
1449:
1442:
1422:Kuo M. (2007).
1421:
1420:
1411:
1398:
1375:
1368:
1353:
1352:
1339:
1330:
1328:
1315:
1314:
1310:
1301:
1299:
1286:
1285:
1281:
1255:
1254:
1250:
1235:10.2307/2421647
1220:
1219:
1215:
1200:10.2307/3753406
1181:
1180:
1173:
1161:
1160:
1156:
1138:
1137:
1133:
1126:
1105:
1104:
1100:
1082:
1081:
1077:
1062:10.2307/3753167
1047:
1046:
1042:
1030:
1029:
1025:
1015:
1014:
1010:
998:
997:
993:
986:
971:
970:
961:
952:
950:
937:
936:
932:
928:
916:
874:states down to
864:Rocky Mountains
834:
800:, found in the
781:T. atratus
770:T. atratus
751:
750:
749:
748:
747:
742:
734:
733:
728:
719:
718:
708:
706:Similar species
692:edible mushroom
618:
617:
616:
615:
614:
611:
602:
601:
600:
597:
586:
485:transferred to
448:
400:deciduous trees
392:Rocky Mountains
356:
339:
324:
320:
301:
283:
261:
256:
238:
232:
227:
213:
195:
184:
182:Schwein. (1822)
165:
154:
148:
135:
43:
17:
12:
11:
5:
2037:
2035:
2027:
2026:
2024:Fungus species
2021:
2016:
2011:
2006:
2001:
1991:
1990:
1984:
1983:
1981:
1980:
1967:
1954:
1941:
1928:
1912:
1910:
1904:
1903:
1901:
1900:
1890:
1877:
1864:
1851:
1838:
1825:
1812:
1799:
1786:
1773:
1758:
1742:
1740:
1734:
1733:
1728:
1722:
1721:
1712:
1709:Index Fungorum
1697:
1696:External links
1694:
1693:
1692:
1686:
1671:
1668:
1666:
1665:
1639:
1613:
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1504:
1497:
1477:
1470:
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1440:
1409:
1373:
1366:
1337:
1308:
1296:Index Fungorum
1279:
1248:
1213:
1171:
1154:
1131:
1124:
1098:
1075:
1040:
1023:
1008:
991:
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959:
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830:
786:New York state
743:
736:
735:
729:
722:
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720:
712:
711:
710:
709:
707:
704:
700:mushroom dying
612:
605:
604:
603:
598:
591:
590:
589:
588:
587:
585:
582:
560:Index Fungorum
464:North Carolina
447:
444:
369:
368:
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352:
351:
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335:
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88:Agaricomycetes
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36:
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9:
6:
4:
3:
2:
2036:
2025:
2022:
2020:
2017:
2015:
2014:Fungi of Asia
2012:
2010:
2007:
2005:
2002:
2000:
1997:
1996:
1994:
1977:
1972:
1968:
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1959:
1955:
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1778:
1774:
1769:
1763:
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1743:
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1739:
1735:
1731:
1726:
1720:
1716:
1713:
1711:
1710:
1705:
1704:
1700:
1699:
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1689:
1687:0-8156-0588-9
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1498:0-8156-0680-X
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558:authorities
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513:Porphyrellus
512:
502:
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491:in his 1898
486:
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419:fruit bodies
416:
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374:
373:
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1855:NatureServe
1829:iNaturalist
1762:Wikispecies
1268:: 199–213.
1056:(1): 4–18.
872:New England
841:mycorrhizal
654:spore print
584:Description
483:Otto Kuntze
454:in 1822 as
396:mycorrhizal
348:mycorrhizal
345:Ecology is
328:Spore print
200:(Schwein.)
189:(Schwein.)
1993:Categories
1670:Cited text
1633:2011-11-14
1607:2011-11-13
1584:(2): 51–8.
1331:2011-11-12
1321:Peck 1878"
1302:2011-11-12
953:2011-11-13
926:References
853:coniferous
717:lookalikes
713:Potential
530:described
471:sanctioned
408:coniferous
388:Boletaceae
108:Boletaceae
74:Division:
1999:Tylopilus
1923:Q59578720
1860:2.1073834
1531:Mycologia
1511:Bessette
1399:Bessette
1274:0082-0598
1258:Tylopilus
1188:Mycologia
1093:490436586
1050:Mycologia
900:Guangdong
849:deciduous
755:Tylopilus
715:Tylopilus
696:Tylopilus
556:taxonomic
526:In 1875,
521:Tylopilus
504:Tylopilus
452:described
442:species.
440:Tylopilus
404:deciduous
126:Species:
119:Tylopilus
98:Boletales
64:Kingdom:
58:Eukaryota
1971:MycoBank
1945:Fungorum
1917:Wikidata
1842:MycoBank
1803:Fungorum
1753:Q3546045
1747:Wikidata
1325:MycoBank
947:MycoBank
914:See also
884:Michigan
880:Missouri
802:Atlantic
669:cystidia
578:commonly
574:alboater
564:MycoBank
540:synonymy
517:subsumed
290:Hymenium
249:hymenium
171:Synonyms
158:Schwein.
104:Family:
54:Domain:
1963:7240630
1896:1133879
1821:3354662
1795:6672285
1717:at the
1551:3760072
1262:Sydowia
1243:2421647
1208:3753406
1070:3753167
908:Sichuan
904:Guangxi
876:Florida
661:hyaline
645:basidia
631:maggots
548:synonym
488:Suillus
479:Boletus
162:Murrill
114:Genus:
94:Order:
84:Class:
1976:168982
1950:168982
1893:uBio:
1886:876660
1847:100204
1808:100204
1684:
1549:
1513:et al.
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896:Fujian
886:, and
868:Quebec
855:, and
773:, and
690:is an
678:hyphae
641:Spores
627:rimose
436:edible
410:, and
384:bolete
365:edible
294:adnate
272:convex
204:(1931)
193:(1898)
191:Kuntze
164:(1909)
1873:80672
1834:63488
1782:7CZYF
1652:(PDF)
1574:(PDF)
1547:JSTOR
1432:227–9
1405:256–7
1401:et al
1292:Peck"
1239:JSTOR
1204:JSTOR
1066:JSTOR
892:Anhui
888:Texas
839:is a
753:Some
635:flesh
542:with
519:into
432:flesh
308:Stipe
245:Pores
68:Fungi
1958:GBIF
1937:MC9F
1868:NCBI
1816:GBIF
1682:ISBN
1493:ISBN
1466:ISBN
1436:ISBN
1362:ISBN
1270:ISSN
1120:ISBN
1089:OCLC
980:ISBN
804:and
650:stem
569:The
428:stem
417:The
332:pink
312:bare
276:flat
1932:CoL
1790:EoL
1777:CoL
1706:in
1539:doi
1231:doi
1196:doi
1116:559
1058:doi
845:oak
680:of
622:cap
562:or
550:of
458:by
423:cap
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310:is
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268:Cap
247:on
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