297:
274:
171:
196:
303:
202:
1447:
50:
310:
209:
1282:
Haren L, Merdes A (May 2002). "Direct binding of NuMA to tubulin is mediated by a novel sequence motif in the tail domain that bundles and stabilizes microtubules".
817:
798:
132:
1257:
Hsu HL, Yeh NH (February 1996). "Dynamic changes of NuMA during the cell cycle and possible appearance of a truncated form of NuMA during apoptosis".
1516:
1492:
1511:
1220:
1202:
47:
296:
1040:
1033:
1354:
Fava F, Raynaud-Messina B, Leung-Tack J, Mazzolini L, Li M, Guillemot JC, Cachot D, Tollon Y, Ferrara P, Wright M (November 1999).
273:
1189:
1168:
195:
170:
1185:
1238:
1164:
112:
309:
208:
1399:"Centrosomal proteins CG-NAP and kendrin provide microtubule nucleation sites by anchoring gamma-tubulin ring complex"
302:
201:
1485:
1311:
Murphy SM, Preble AM, Patel UK, O'Connell KL, Dias DP, Moritz M, Agard D, Stults JT, Stearns T (November 2001).
862:
120:
843:
1104:
1478:
184:
99:
1012:
987:
957:
932:
144:
1016:
991:
961:
936:
1428:
1385:
1342:
1299:
1270:
92:
40:
1462:
140:
1418:
1410:
1375:
1367:
1332:
1324:
1291:
1262:
389:
320:
264:
219:
364:
1380:
1355:
1423:
1398:
732:
727:
722:
717:
712:
707:
702:
697:
692:
687:
671:
666:
661:
656:
651:
646:
641:
636:
631:
626:
621:
616:
611:
595:
590:
585:
580:
1505:
1458:
1337:
1312:
567:
124:
382:
161:
148:
1225:
National Center for
Biotechnology Information, U.S. National Library of Medicine
1207:
National Center for
Biotechnology Information, U.S. National Library of Medicine
465:
1108:
281:
178:
128:
103:, 76P, GCP-4, GCP4, Grip76, MCCRP3, tubulin gamma complex associated protein 4
1414:
762:
525:
403:
348:
335:
247:
234:
136:
1432:
1389:
1346:
1328:
1303:
1295:
1371:
1274:
1266:
1080:
1075:
1064:
907:
888:
1356:"Human 76p: A new member of the gamma-tubulin-associated protein family"
1397:
Takahashi M, Yamagiwa A, Nishimura T, Mukai H, Ono Y (September 2002).
1140:
1135:
1130:
1125:
1120:
1096:
874:
829:
69:
1048:
784:
1446:
1313:"GCP5 and GCP6: two new members of the human gamma-tubulin complex"
688:
microtubule nucleation by interphase microtubule organizing center
747:
743:
1454:
1100:
116:
1103:. It is part of the gamma tubulin complex, which required for
1239:"Entrez Gene: Tubulin, gamma complex associated protein 4"
372:
1466:
537:
1005:
980:
950:
925:
1181:
1179:
1177:
1160:
1158:
1156:
319:
218:
16:Protein-coding gene in the species Homo sapiens
1186:GRCm38: Ensembl release 89: ENSMUSG00000027263
1486:
8:
1165:GRCh38: Ensembl release 89: ENSG00000137822
1093:Tubulin, gamma complex associated protein 4
1493:
1479:
758:
563:
360:
259:
156:
58:
1422:
1379:
1336:
1099:in humans that is encoded by the TUBGCP4
647:equatorial microtubule organizing center
1152:
581:structural constituent of cytoskeleton
18:
693:microtubule cytoskeleton organization
324:
285:
280:
223:
182:
177:
7:
1443:
1441:
708:cytoplasmic microtubule organization
733:protein-containing complex assembly
1465:. You can help Knowledge (XXG) by
1002:
977:
947:
922:
898:
879:
853:
834:
808:
789:
542:
460:
398:
377:
14:
1445:
308:
301:
295:
272:
207:
200:
194:
169:
1517:Human chromosome 15 gene stubs
526:More reference expression data
1:
1403:Molecular Biology of the Cell
1317:Molecular Biology of the Cell
617:microtubule organizing center
586:microtubule minus-end binding
293:
192:
1512:Genes on human chromosome 15
1360:The Journal of Cell Biology
1261:. 109 ( Pt 2) (2): 277–88.
1533:
1440:
1221:"Mouse PubMed Reference:"
1203:"Human PubMed Reference:"
1079:
1074:
1070:
1063:
1047:
1028:
1009:
984:
973:
954:
929:
918:
905:
901:
886:
882:
873:
860:
856:
841:
837:
828:
815:
811:
796:
792:
783:
768:
761:
757:
741:
566:
562:
550:
545:
536:
523:
472:
463:
410:
401:
371:
363:
359:
342:
329:
292:
271:
262:
258:
241:
228:
191:
168:
159:
155:
110:
107:
97:
90:
85:
66:
61:
44:
39:
34:
30:
26:
21:
1034:Chr 15: 43.37 – 43.41 Mb
698:mitotic spindle assembly
662:microtubule cytoskeleton
442:tibialis anterior muscle
1415:10.1091/mbc.E02-02-0112
1284:Journal of Cell Science
1259:Journal of Cell Science
504:internal carotid artery
1329:10.1091/mbc.12.11.3340
1296:10.1242/jcs.115.9.1815
1105:microtubule nucleation
1041:Chr 2: 121 – 121.03 Mb
713:microtubule nucleation
703:centrosome duplication
500:neural layer of retina
1372:10.1083/jcb.147.4.857
1267:10.1242/jcs.109.2.277
622:gamma-tubulin complex
596:gamma-tubulin binding
450:cerebellar hemisphere
185:Chromosome 15 (human)
326:2 E5|2 60.37 cM
287:Chromosome 2 (mouse)
62:List of PDB id codes
35:Available structures
446:epithelium of colon
426:ganglionic eminence
149:TUBGCP4 - orthologs
1453:This article on a
863:ENSMUSG00000027263
723:mitotic cell cycle
681:Biological process
642:recycling endosome
605:Cellular component
574:Molecular function
496:supraoptic nucleus
422:buccal mucosa cell
1474:
1473:
1290:(Pt 9): 1815–24.
1090:
1089:
1086:
1085:
1059:
1058:
1024:
1023:
999:
998:
969:
968:
944:
943:
914:
913:
895:
894:
869:
868:
850:
849:
824:
823:
805:
804:
753:
752:
632:spindle pole body
558:
557:
554:
553:
532:
531:
519:
518:
457:
456:
355:
354:
254:
253:
81:
80:
77:
76:
45:Ortholog search:
1524:
1495:
1488:
1481:
1449:
1442:
1436:
1426:
1393:
1383:
1350:
1340:
1307:
1278:
1243:
1242:
1235:
1229:
1228:
1217:
1211:
1210:
1199:
1193:
1183:
1172:
1162:
1072:
1071:
1043:
1036:
1019:
1003:
994:
978:
974:RefSeq (protein)
964:
948:
939:
923:
899:
880:
854:
835:
809:
790:
759:
728:spindle assembly
564:
543:
528:
488:genital tubercle
468:
466:Top expressed in
461:
430:ventricular zone
418:secondary oocyte
406:
404:Top expressed in
399:
378:
361:
351:
338:
327:
312:
305:
299:
288:
276:
260:
250:
237:
226:
211:
204:
198:
187:
173:
157:
151:
102:
95:
72:
59:
53:
32:
31:
19:
1532:
1531:
1527:
1526:
1525:
1523:
1522:
1521:
1502:
1501:
1500:
1499:
1439:
1396:
1353:
1323:(11): 3340–52.
1310:
1281:
1256:
1252:
1250:Further reading
1247:
1246:
1237:
1236:
1232:
1219:
1218:
1214:
1201:
1200:
1196:
1184:
1175:
1163:
1154:
1149:
1117:
1081:View/Edit Mouse
1076:View/Edit Human
1039:
1032:
1029:Location (UCSC)
1015:
1011:
990:
986:
960:
956:
935:
931:
844:ENSG00000137822
737:
676:
600:
591:protein binding
524:
515:
510:
506:
502:
498:
494:
490:
486:
482:
478:
476:ascending aorta
464:
453:
448:
444:
440:
436:
432:
428:
424:
420:
416:
402:
346:
333:
325:
315:
314:
313:
306:
286:
263:Gene location (
245:
232:
224:
214:
213:
212:
205:
183:
160:Gene location (
111:
98:
91:
68:
46:
17:
12:
11:
5:
1530:
1528:
1520:
1519:
1514:
1504:
1503:
1498:
1497:
1490:
1483:
1475:
1472:
1471:
1450:
1438:
1437:
1409:(9): 3235–45.
1394:
1351:
1308:
1279:
1253:
1251:
1248:
1245:
1244:
1230:
1212:
1194:
1173:
1151:
1150:
1148:
1145:
1144:
1143:
1138:
1133:
1128:
1123:
1116:
1113:
1088:
1087:
1084:
1083:
1078:
1068:
1067:
1061:
1060:
1057:
1056:
1054:
1052:
1045:
1044:
1037:
1030:
1026:
1025:
1022:
1021:
1007:
1006:
1000:
997:
996:
982:
981:
975:
971:
970:
967:
966:
952:
951:
945:
942:
941:
927:
926:
920:
916:
915:
912:
911:
903:
902:
896:
893:
892:
884:
883:
877:
871:
870:
867:
866:
858:
857:
851:
848:
847:
839:
838:
832:
826:
825:
822:
821:
813:
812:
806:
803:
802:
794:
793:
787:
781:
780:
775:
770:
766:
765:
755:
754:
751:
750:
739:
738:
736:
735:
730:
725:
720:
715:
710:
705:
700:
695:
690:
684:
682:
678:
677:
675:
674:
669:
664:
659:
654:
649:
644:
639:
634:
629:
624:
619:
614:
608:
606:
602:
601:
599:
598:
593:
588:
583:
577:
575:
571:
570:
560:
559:
556:
555:
552:
551:
548:
547:
540:
534:
533:
530:
529:
521:
520:
517:
516:
514:
513:
509:
505:
501:
497:
493:
489:
485:
484:tail of embryo
481:
477:
473:
470:
469:
458:
455:
454:
452:
451:
447:
443:
439:
435:
431:
427:
423:
419:
415:
411:
408:
407:
395:
394:
386:
375:
369:
368:
365:RNA expression
357:
356:
353:
352:
344:
340:
339:
331:
328:
323:
317:
316:
307:
300:
294:
290:
289:
284:
278:
277:
269:
268:
256:
255:
252:
251:
243:
239:
238:
230:
227:
222:
216:
215:
206:
199:
193:
189:
188:
181:
175:
174:
166:
165:
153:
152:
109:
105:
104:
96:
88:
87:
83:
82:
79:
78:
75:
74:
64:
63:
55:
54:
43:
37:
36:
28:
27:
24:
23:
15:
13:
10:
9:
6:
4:
3:
2:
1529:
1518:
1515:
1513:
1510:
1509:
1507:
1496:
1491:
1489:
1484:
1482:
1477:
1476:
1470:
1468:
1464:
1460:
1459:chromosome 15
1456:
1451:
1448:
1444:
1434:
1430:
1425:
1420:
1416:
1412:
1408:
1404:
1400:
1395:
1391:
1387:
1382:
1377:
1373:
1369:
1366:(4): 857–68.
1365:
1361:
1357:
1352:
1348:
1344:
1339:
1334:
1330:
1326:
1322:
1318:
1314:
1309:
1305:
1301:
1297:
1293:
1289:
1285:
1280:
1276:
1272:
1268:
1264:
1260:
1255:
1254:
1249:
1240:
1234:
1231:
1226:
1222:
1216:
1213:
1208:
1204:
1198:
1195:
1191:
1187:
1182:
1180:
1178:
1174:
1170:
1166:
1161:
1159:
1157:
1153:
1146:
1142:
1139:
1137:
1134:
1132:
1129:
1127:
1124:
1122:
1119:
1118:
1114:
1112:
1110:
1106:
1102:
1098:
1094:
1082:
1077:
1073:
1069:
1066:
1062:
1055:
1053:
1050:
1046:
1042:
1038:
1035:
1031:
1027:
1020:
1018:
1014:
1008:
1004:
1001:
995:
993:
989:
983:
979:
976:
972:
965:
963:
959:
953:
949:
946:
940:
938:
934:
928:
924:
921:
919:RefSeq (mRNA)
917:
910:
909:
904:
900:
897:
891:
890:
885:
881:
878:
876:
872:
865:
864:
859:
855:
852:
846:
845:
840:
836:
833:
831:
827:
820:
819:
814:
810:
807:
801:
800:
795:
791:
788:
786:
782:
779:
776:
774:
771:
767:
764:
760:
756:
749:
745:
740:
734:
731:
729:
726:
724:
721:
719:
716:
714:
711:
709:
706:
704:
701:
699:
696:
694:
691:
689:
686:
685:
683:
680:
679:
673:
670:
668:
665:
663:
660:
658:
655:
653:
650:
648:
645:
643:
640:
638:
635:
633:
630:
628:
625:
623:
620:
618:
615:
613:
610:
609:
607:
604:
603:
597:
594:
592:
589:
587:
584:
582:
579:
578:
576:
573:
572:
569:
568:Gene ontology
565:
561:
549:
544:
541:
539:
535:
527:
522:
511:
507:
503:
499:
495:
491:
487:
483:
479:
475:
474:
471:
467:
462:
459:
449:
445:
441:
437:
433:
429:
425:
421:
417:
413:
412:
409:
405:
400:
397:
396:
393:
391:
387:
385:
384:
380:
379:
376:
374:
370:
366:
362:
358:
350:
345:
341:
337:
332:
322:
318:
311:
304:
298:
291:
283:
279:
275:
270:
266:
261:
257:
249:
244:
240:
236:
231:
221:
217:
210:
203:
197:
190:
186:
180:
176:
172:
167:
163:
158:
154:
150:
146:
142:
138:
134:
130:
126:
122:
118:
114:
106:
101:
94:
89:
84:
73:
71:
65:
60:
57:
56:
52:
49:
42:
38:
33:
29:
25:
20:
1467:expanding it
1452:
1406:
1402:
1363:
1359:
1320:
1316:
1287:
1283:
1258:
1233:
1224:
1215:
1206:
1197:
1092:
1091:
1013:NP_001277753
1010:
988:NP_001273343
985:
958:NM_001290824
955:
933:NM_001286414
930:
906:
887:
861:
842:
816:
797:
777:
772:
667:cytoskeleton
637:spindle pole
492:aortic valve
480:spermatocyte
388:
381:
347:121,029,251
334:121,001,135
108:External IDs
67:
657:microtubule
246:43,409,771
233:43,369,221
86:Identifiers
1506:Categories
1192:, May 2017
1171:, May 2017
1147:References
1109:centrosome
672:centrosome
392:(ortholog)
129:HomoloGene
1457:on human
1017:NP_700436
992:NP_055259
962:NM_153387
937:NM_014444
763:Orthologs
612:cytoplasm
512:spermatid
137:GeneCards
1433:12221128
1390:10562286
1347:11694571
1304:11956313
1188:–
1167:–
1115:See also
1065:Wikidata
742:Sources:
652:membrane
438:testicle
1381:2156165
1275:8838651
1190:Ensembl
1169:Ensembl
1141:TUBGCP6
1136:TUBGCP5
1131:TUBGCP3
1126:TUBGCP2
1121:Tubulin
1107:at the
1097:protein
875:UniProt
830:Ensembl
769:Species
748:QuickGO
718:meiosis
627:cytosol
367:pattern
225:15q15.3
141:TUBGCP4
125:1196293
100:TUBGCP4
93:Aliases
22:TUBGCP4
1431:
1424:124155
1421:
1388:
1378:
1345:
1335:
1302:
1273:
1051:search
1049:PubMed
908:Q9D4F8
889:Q9UGJ1
785:Entrez
538:BioGPS
508:thymus
117:609610
1461:is a
1338:60259
1095:is a
818:51885
799:27229
778:Mouse
773:Human
744:Amigo
434:gonad
414:sperm
390:Mouse
383:Human
330:Start
265:Mouse
229:Start
162:Human
1463:stub
1455:gene
1429:PMID
1386:PMID
1343:PMID
1300:PMID
1271:PMID
1101:gene
373:Bgee
321:Band
282:Chr.
220:Band
179:Chr.
133:8690
113:OMIM
70:3RIP
51:RCSB
48:PDBe
1419:PMC
1411:doi
1376:PMC
1368:doi
1364:147
1333:PMC
1325:doi
1292:doi
1288:115
1263:doi
546:n/a
343:End
242:End
145:OMA
121:MGI
41:PDB
1508::
1427:.
1417:.
1407:13
1405:.
1401:.
1384:.
1374:.
1362:.
1358:.
1341:.
1331:.
1321:12
1319:.
1315:.
1298:.
1286:.
1269:.
1223:.
1205:.
1176:^
1155:^
1111:.
746:/
349:bp
336:bp
248:bp
235:bp
143:;
139::
135:;
131::
127:;
123::
119:;
115::
1494:e
1487:t
1480:v
1469:.
1435:.
1413::
1392:.
1370::
1349:.
1327::
1306:.
1294::
1277:.
1265::
1241:.
1227:.
1209:.
267:)
164:)
147::
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.