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Takakia ceratophylla

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The capsule expands and swells during development, in consequence, a spiral suture line called capsule dehiscence becomes evident by breaking down of the dextrorsely-arranged exothecial cells on the outer cell wall. The separation begins from the middle of the capsule and extend towards the apex and
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and depends on passive spore release. The spores production site is located in the inner space of the capsule. During spore release, the seta twists toward the right direction and the capsule dehiscence widens and becomes flattened, such that the spores can pass through the split and disperse over
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is protected by a small hood-like calyptra. The thickness of the calyptra gradually thins down from the apex towards the base. The seta and capsules are initially green, gradually turning into brown or black colour, and elongate or expand prior to the maturation of the sporangium. The long-tapered
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are irregularly spiral arranged and contain simple oil droplets and axillary hairs. The leaves are multitratose, and composed of 13-20 thick-walled cells. The unique colourless intercalary rhizomatous branches functions to produce new stems and shoots.
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gives rise to the capsule wall, while the endothecium gives rise to the archesporium and the dome-shaped columella, where the cells of the columella and spore sac break down during spore maturation.
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for nearly a century. Later rediscovered in Japan and claimed to be sharing traits from both the liverworts and mosses based on genetic sequences and morphological characters.
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foot is responsible for transferring nutrients from the gametophyte toward the sporophyte. And a well-defined conducting strand that contains
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are moss-like, and can be found buried in between the leaves on the lateral side of the stem. The stem of
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that experiences late snow cover. The species can be found at low to moderate elevations (70–700 m).
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time. The wideness of the capsule dehiscence depends on the age and dryness of the outer cells.
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for this species are rocky areas (eg. inland or mountain cliffs) and cold
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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Renzaglia, Karen Sue; McFarland, Ken D.; Smith, David K. (October 1997).
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is vulnerable and threatened by habitat loss due to human activities.
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mechanism of the species. The dried specimens do not contain odour.
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Some features that are similar to mosses include the presence of
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and peristome teeth are not differentiated in this species.
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is one of the two species of toothless mosses in the genus
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are rigid and green, the distal part of the shoots are
1523:"SPOROPHYTES OF TAKAKIA CERATOPHYLLA FOUND IN CHINA" 1437:"SPOROPHYTES OF TAKAKIA CERATOPHYLLA FOUND IN CHINA" 1406:"SPOROPHYTES OF TAKAKIA CERATOPHYLLA FOUND IN CHINA" 1265:"SPOROPHYTES OF TAKAKIA CERATOPHYLLA FOUND IN CHINA" 781:"SPOROPHYTES OF TAKAKIA CERATOPHYLLA FOUND IN CHINA" 750:"SPOROPHYTES OF TAKAKIA CERATOPHYLLA FOUND IN CHINA" 1671: 250:was first described as a liverwort under the genus 1527:The Journal of the Hattori Botanical Laboratory 1441:The Journal of the Hattori Botanical Laboratory 1410:The Journal of the Hattori Botanical Laboratory 1269:The Journal of the Hattori Botanical Laboratory 908:The Journal of the Hattori Botanical Laboratory 785:The Journal of the Hattori Botanical Laboratory 754:The Journal of the Hattori Botanical Laboratory 8: 867:10.2305/iucn.uk.2000.rlts.t39189a10168338.en 821:10.2305/iucn.uk.2000.rlts.t39189a10168338.en 603:10.2305/iucn.uk.2000.rlts.t39189a10168338.en 477:10.2305/IUCN.UK.2000.RLTS.T39189A10168338.en 231:family. This species was first described by 1521:Higuchi, Masanobu; Zhang, Da-Cheng (1998). 1435:Higuchi, Masanobu; Zhang, Da-Cheng (1998). 1404:Higuchi, Masanobu; Zhang, Da-Cheng (1998). 1263:Higuchi, Masanobu; Zhang, Da-Cheng (1998). 779:Higuchi, Masanobu; Zhang, Da-Cheng (1998). 748:Higuchi, Masanobu; Zhang, Da-Cheng (1998). 416:continues from the seta and into the foot. 1659: 902:Smith, David K.; Davison, Paul G. (1993). 368:The reproduction structure of the stalked 55: 28: 17: 865: 819: 601: 475: 444: 384:and contains a weak conducting strand. 360:The tapered and deeply-lobed leaves of 884: 873: 838: 827: 620: 609: 504:Γ–sterreichische Botanische Zeitschrift 267:; the elongation and twisting of the 7: 463:IUCN Red List of Threatened Species 452:Bryophyte Specialist Group (2000). 297:is terrestrial and can be found in 14: 1922:IUCN Red List vulnerable species 80: 1: 290:Geographic range and habitat 353:, which contributes to the 1948: 1613:American Journal of Botany 1558:American Journal of Botany 1472:American Journal of Botany 1355:American Journal of Botany 1300:American Journal of Botany 1214:American Journal of Botany 1159:American Journal of Botany 1104:American Journal of Botany 1049:American Journal of Botany 994:American Journal of Botany 939:American Journal of Botany 699:American Journal of Botany 644:American Journal of Botany 543:American Journal of Botany 204: 197: 182: 175: 77:Scientific classification 75: 53: 44: 36: 27: 20: 498:Grolle, Riclef (1963). " 920:10.18968/jhbl.73.0_263 883:Cite journal requires 837:Cite journal requires 619:Cite journal requires 400: 342: 207:Lepidozia ceratophylla 1539:10.18968/jhbl.84.0_57 1453:10.18968/jhbl.84.0_57 1422:10.18968/jhbl.84.0_57 1281:10.18968/jhbl.84.0_57 797:10.18968/jhbl.84.0_57 766:10.18968/jhbl.84.0_57 470:: e.T39189A10168338. 395: 337: 1703:Takakia ceratophylla 1673:Takakia ceratophylla 456:Takakia ceratophylla 428:Takakia ceratophylla 409:Takakia ceratophylla 398:Takakia ceratophylla 378:Takakia ceratophylla 362:Takakia ceratophylla 355:asexual reproduction 340:Takakia ceratophylla 295:Takakia ceratophylla 263:, tapered foot, and 248:Takakia ceratophylla 237:Takakia ceratophylla 218:Takakia ceratophylla 186:Takakia ceratophylla 168:T. ceratophylla 39:Takakia ceratophylla 22:Takakia ceratophylla 856:IUCN (2000-06-30). 810:IUCN (2000-06-30). 592:IUCN (2000-06-30). 372:and the lageniform 47:Conservation status 516:10.1007/BF01373680 401: 345:The shoots of the 343: 37:Dried specimen of 1932:Vulnerable plants 1909: 1908: 1842:Open Tree of Life 1665:Taxon identifiers 1619:(10): 1337–1350. 1564:(10): 1337–1350. 1478:(10): 1337–1350. 1361:(10): 1337–1350. 1306:(10): 1337–1350. 1220:(10): 1337–1350. 1165:(10): 1337–1350. 1110:(10): 1337–1350. 1055:(10): 1337–1350. 1000:(10): 1337–1350. 945:(10): 1337–1350. 705:(10): 1337–1350. 650:(10): 1337–1350. 549:(10): 1337–1350. 214: 213: 70: 1939: 1902: 1901: 1889: 1888: 1876: 1875: 1863: 1862: 1850: 1849: 1837: 1836: 1824: 1823: 1811: 1810: 1798: 1797: 1785: 1784: 1772: 1771: 1759: 1758: 1746: 1745: 1733: 1732: 1720: 1719: 1707: 1706: 1705: 1692: 1691: 1690: 1660: 1653: 1652: 1604: 1598: 1597: 1549: 1543: 1542: 1518: 1512: 1511: 1463: 1457: 1456: 1432: 1426: 1425: 1401: 1395: 1394: 1346: 1340: 1339: 1291: 1285: 1284: 1260: 1254: 1253: 1205: 1199: 1198: 1150: 1144: 1143: 1095: 1089: 1088: 1040: 1034: 1033: 985: 979: 978: 930: 924: 923: 899: 893: 892: 886: 881: 879: 871: 869: 853: 847: 846: 840: 835: 833: 825: 823: 807: 801: 800: 776: 770: 769: 745: 739: 738: 690: 684: 683: 635: 629: 628: 622: 617: 615: 607: 605: 589: 583: 582: 534: 528: 527: 495: 489: 488: 486: 484: 479: 449: 338:Gametophytes of 188: 85: 84: 64: 59: 58: 32: 18: 1947: 1946: 1942: 1941: 1940: 1938: 1937: 1936: 1912: 1911: 1910: 1905: 1897: 1892: 1884: 1879: 1871: 1866: 1858: 1853: 1845: 1840: 1832: 1827: 1819: 1814: 1806: 1801: 1793: 1788: 1780: 1775: 1767: 1762: 1754: 1749: 1741: 1736: 1728: 1723: 1715: 1710: 1701: 1700: 1695: 1686: 1685: 1680: 1667: 1657: 1656: 1625:10.2307/2446132 1606: 1605: 1601: 1570:10.2307/2446132 1551: 1550: 1546: 1520: 1519: 1515: 1484:10.2307/2446132 1465: 1464: 1460: 1434: 1433: 1429: 1403: 1402: 1398: 1367:10.2307/2446132 1348: 1347: 1343: 1312:10.2307/2446132 1293: 1292: 1288: 1262: 1261: 1257: 1226:10.2307/2446132 1207: 1206: 1202: 1171:10.2307/2446132 1152: 1151: 1147: 1116:10.2307/2446132 1097: 1096: 1092: 1061:10.2307/2446132 1042: 1041: 1037: 1006:10.2307/2446132 987: 986: 982: 951:10.2307/2446132 932: 931: 927: 901: 900: 896: 882: 872: 855: 854: 850: 836: 826: 809: 808: 804: 778: 777: 773: 747: 746: 742: 711:10.2307/2446132 692: 691: 687: 656:10.2307/2446132 637: 636: 632: 618: 608: 591: 590: 586: 555:10.2307/2446132 536: 535: 531: 497: 496: 492: 482: 480: 451: 450: 446: 441: 396:Sporophytes of 390: 380:is composed of 332: 327: 292: 287: 286:Species of moss 279:in sporangium, 245: 193: 190: 184: 171: 79: 71: 60: 56: 49: 12: 11: 5: 1945: 1943: 1935: 1934: 1929: 1924: 1914: 1913: 1907: 1906: 1904: 1903: 1899:wfo-0001195981 1890: 1877: 1864: 1851: 1838: 1825: 1812: 1799: 1786: 1773: 1760: 1747: 1734: 1721: 1708: 1693: 1677: 1675: 1669: 1668: 1663: 1655: 1654: 1599: 1544: 1513: 1458: 1427: 1396: 1341: 1286: 1255: 1200: 1145: 1090: 1035: 980: 925: 894: 885:|journal= 848: 839:|journal= 802: 771: 740: 685: 630: 621:|journal= 584: 529: 510:(4): 444–447. 502:im Himalaya". 490: 443: 442: 440: 437: 389: 386: 331: 328: 326: 323: 313:. The natural 291: 288: 285: 244: 241: 233:William Mitten 212: 211: 202: 201: 195: 194: 192:(Mitt.) Grolle 191: 180: 179: 173: 172: 165: 163: 159: 158: 151: 147: 146: 141: 137: 136: 131: 127: 126: 121: 117: 116: 114:Takakiophytina 111: 107: 106: 101: 97: 96: 91: 87: 86: 73: 72: 54: 51: 50: 45: 42: 41: 34: 33: 25: 24: 13: 10: 9: 6: 4: 3: 2: 1944: 1933: 1930: 1928: 1925: 1923: 1920: 1919: 1917: 1900: 1895: 1891: 1887: 1882: 1878: 1874: 1869: 1865: 1861: 1856: 1852: 1848: 1843: 1839: 1835: 1830: 1826: 1822: 1817: 1813: 1809: 1804: 1800: 1796: 1791: 1787: 1783: 1778: 1774: 1770: 1765: 1761: 1757: 1752: 1748: 1744: 1739: 1735: 1731: 1726: 1722: 1718: 1713: 1709: 1704: 1698: 1694: 1689: 1683: 1679: 1678: 1676: 1674: 1670: 1666: 1661: 1650: 1646: 1642: 1638: 1634: 1630: 1626: 1622: 1618: 1614: 1610: 1603: 1600: 1595: 1591: 1587: 1583: 1579: 1575: 1571: 1567: 1563: 1559: 1555: 1548: 1545: 1540: 1536: 1532: 1528: 1524: 1517: 1514: 1509: 1505: 1501: 1497: 1493: 1489: 1485: 1481: 1477: 1473: 1469: 1462: 1459: 1454: 1450: 1446: 1442: 1438: 1431: 1428: 1423: 1419: 1415: 1411: 1407: 1400: 1397: 1392: 1388: 1384: 1380: 1376: 1372: 1368: 1364: 1360: 1356: 1352: 1345: 1342: 1337: 1333: 1329: 1325: 1321: 1317: 1313: 1309: 1305: 1301: 1297: 1290: 1287: 1282: 1278: 1274: 1270: 1266: 1259: 1256: 1251: 1247: 1243: 1239: 1235: 1231: 1227: 1223: 1219: 1215: 1211: 1204: 1201: 1196: 1192: 1188: 1184: 1180: 1176: 1172: 1168: 1164: 1160: 1156: 1149: 1146: 1141: 1137: 1133: 1129: 1125: 1121: 1117: 1113: 1109: 1105: 1101: 1094: 1091: 1086: 1082: 1078: 1074: 1070: 1066: 1062: 1058: 1054: 1050: 1046: 1039: 1036: 1031: 1027: 1023: 1019: 1015: 1011: 1007: 1003: 999: 995: 991: 984: 981: 976: 972: 968: 964: 960: 956: 952: 948: 944: 940: 936: 929: 926: 921: 917: 913: 909: 905: 898: 895: 890: 877: 868: 863: 859: 852: 849: 844: 831: 822: 817: 813: 806: 803: 798: 794: 790: 786: 782: 775: 772: 767: 763: 759: 755: 751: 744: 741: 736: 732: 728: 724: 720: 716: 712: 708: 704: 700: 696: 689: 686: 681: 677: 673: 669: 665: 661: 657: 653: 649: 645: 641: 634: 631: 626: 613: 604: 599: 595: 588: 585: 580: 576: 572: 568: 564: 560: 556: 552: 548: 544: 540: 533: 530: 525: 521: 517: 513: 509: 505: 501: 494: 491: 478: 473: 469: 465: 464: 459: 457: 448: 445: 438: 436: 433: 429: 425: 423: 417: 415: 410: 406: 399: 394: 387: 385: 383: 379: 375: 371: 366: 363: 358: 356: 352: 348: 341: 336: 329: 324: 322: 320: 316: 312: 311:United States 308: 304: 300: 296: 289: 284: 282: 278: 274: 270: 266: 262: 257: 255: 254: 249: 242: 240: 238: 234: 230: 226: 225: 220: 219: 210: 208: 203: 200: 196: 189: 187: 181: 178: 177:Binomial name 174: 170: 169: 164: 161: 160: 157: 156: 152: 149: 148: 145: 142: 139: 138: 135: 132: 129: 128: 125: 122: 119: 118: 115: 112: 110:Subdivision: 109: 108: 105: 102: 99: 98: 95: 92: 89: 88: 83: 78: 74: 68: 63: 52: 48: 43: 40: 35: 31: 26: 23: 19: 16: 1927:Takakiopsida 1860:tro-35184413 1672: 1616: 1612: 1602: 1561: 1557: 1547: 1530: 1526: 1516: 1475: 1471: 1461: 1444: 1440: 1430: 1413: 1409: 1399: 1358: 1354: 1344: 1303: 1299: 1289: 1272: 1268: 1258: 1217: 1213: 1203: 1162: 1158: 1148: 1107: 1103: 1093: 1052: 1048: 1038: 997: 993: 983: 942: 938: 928: 911: 907: 897: 876:cite journal 851: 830:cite journal 805: 788: 784: 774: 757: 753: 743: 702: 698: 688: 647: 643: 633: 612:cite journal 587: 546: 542: 532: 507: 503: 499: 493: 481:. Retrieved 467: 461: 455: 447: 427: 426: 422:amphithecium 418: 408: 402: 397: 377: 367: 361: 359: 344: 339: 330:Gametophytes 294: 293: 258: 251: 247: 246: 236: 227:, under the 222: 217: 216: 215: 206: 205: 185: 183: 167: 166: 154: 124:Takakiopsida 38: 21: 15: 1816:NatureServe 1764:iNaturalist 1697:Wikispecies 914:: 263–271. 483:16 November 432:hygroscopic 388:Sporophytes 347:gametophyte 243:Description 229:Takakiaceae 144:Takakiaceae 1916:Categories 1855:Plant List 439:References 420:base. The 405:sporangium 374:archegonia 370:antheridia 325:Morphology 309:, and the 134:Takakiales 100:Division: 62:Vulnerable 1743:200000001 1633:0002-9122 1578:0002-9122 1533:: 57–69. 1492:0002-9122 1447:: 57–69. 1416:: 57–69. 1375:0002-9122 1320:0002-9122 1275:: 57–69. 1234:0002-9122 1179:0002-9122 1124:0002-9122 1069:0002-9122 1014:0002-9122 959:0002-9122 791:: 57–69. 760:: 57–69. 719:0002-9122 664:0002-9122 563:0002-9122 281:operculum 261:columella 253:Lepidozia 235:in 1861. 162:Species: 104:Bryophyta 90:Kingdom: 1886:35184413 1881:Tropicos 1821:2.121615 1782:10953575 1682:Wikidata 1649:21708543 1594:21708543 1508:21708543 1391:21708543 1336:21708543 1250:21708543 1195:21708543 1140:21708543 1085:21708543 1030:21708543 975:21708543 735:21708543 680:21708543 579:21708543 524:42977298 414:hydroids 382:cuticles 351:caducous 315:habitats 273:rhizoids 265:calyptra 199:Synonyms 140:Family: 67:IUCN 2.3 1756:5710305 1641:2446132 1586:2446132 1500:2446132 1383:2446132 1328:2446132 1242:2446132 1187:2446132 1132:2446132 1077:2446132 1022:2446132 967:2446132 727:2446132 672:2446132 571:2446132 500:Takakia 277:stomata 224:Takakia 155:Takakia 150:Genus: 130:Order: 120:Class: 94:Plantae 65: ( 1868:PLANTS 1834:253518 1769:155950 1717:420550 1688:Q80262 1647:  1639:  1631:  1592:  1584:  1576:  1506:  1498:  1490:  1389:  1381:  1373:  1334:  1326:  1318:  1248:  1240:  1232:  1193:  1185:  1177:  1138:  1130:  1122:  1083:  1075:  1067:  1028:  1020:  1012:  973:  965:  957:  733:  725:  717:  678:  670:  662:  577:  569:  561:  522:  319:desert 209:Mitt. 1873:TACE2 1808:39189 1795:14204 1777:IRMNG 1730:TAKCE 1637:JSTOR 1582:JSTOR 1496:JSTOR 1379:JSTOR 1324:JSTOR 1238:JSTOR 1183:JSTOR 1128:JSTOR 1073:JSTOR 1018:JSTOR 963:JSTOR 723:JSTOR 668:JSTOR 567:JSTOR 520:S2CID 307:Nepal 303:India 299:China 1847:3228 1829:NCBI 1803:IUCN 1790:ITIS 1751:GBIF 1725:EPPO 1645:PMID 1629:ISSN 1590:PMID 1574:ISSN 1504:PMID 1488:ISSN 1387:PMID 1371:ISSN 1332:PMID 1316:ISSN 1246:PMID 1230:ISSN 1191:PMID 1175:ISSN 1136:PMID 1120:ISSN 1081:PMID 1065:ISSN 1026:PMID 1010:ISSN 971:PMID 955:ISSN 889:help 843:help 731:PMID 715:ISSN 676:PMID 660:ISSN 625:help 575:PMID 559:ISSN 485:2021 468:2000 403:The 269:seta 1894:WFO 1738:FNA 1712:EoL 1621:doi 1566:doi 1535:doi 1480:doi 1449:doi 1418:doi 1363:doi 1308:doi 1277:doi 1222:doi 1167:doi 1112:doi 1057:doi 1002:doi 947:doi 916:doi 862:doi 816:doi 793:doi 762:doi 707:doi 652:doi 598:doi 551:doi 512:doi 508:110 472:doi 430:is 407:of 1918:: 1896:: 1883:: 1870:: 1857:: 1844:: 1831:: 1818:: 1805:: 1792:: 1779:: 1766:: 1753:: 1740:: 1727:: 1714:: 1699:: 1684:: 1643:. 1635:. 1627:. 1617:84 1615:. 1611:. 1588:. 1580:. 1572:. 1562:84 1560:. 1556:. 1531:84 1529:. 1525:. 1502:. 1494:. 1486:. 1476:84 1474:. 1470:. 1445:84 1443:. 1439:. 1414:84 1412:. 1408:. 1385:. 1377:. 1369:. 1359:84 1357:. 1353:. 1330:. 1322:. 1314:. 1304:84 1302:. 1298:. 1273:84 1271:. 1267:. 1244:. 1236:. 1228:. 1218:84 1216:. 1212:. 1189:. 1181:. 1173:. 1163:84 1161:. 1157:. 1134:. 1126:. 1118:. 1108:84 1106:. 1102:. 1079:. 1071:. 1063:. 1053:84 1051:. 1047:. 1024:. 1016:. 1008:. 998:84 996:. 992:. 969:. 961:. 953:. 943:84 941:. 937:. 912:73 910:. 906:. 880:: 878:}} 874:{{ 860:. 834:: 832:}} 828:{{ 814:. 789:84 787:. 783:. 758:84 756:. 752:. 729:. 721:. 713:. 703:84 701:. 697:. 674:. 666:. 658:. 648:84 646:. 642:. 616:: 614:}} 610:{{ 596:. 573:. 565:. 557:. 547:84 545:. 541:. 518:. 506:. 466:. 460:. 305:, 301:, 275:, 1651:. 1623:: 1596:. 1568:: 1541:. 1537:: 1510:. 1482:: 1455:. 1451:: 1424:. 1420:: 1393:. 1365:: 1338:. 1310:: 1283:. 1279:: 1252:. 1224:: 1197:. 1169:: 1142:. 1114:: 1087:. 1059:: 1032:. 1004:: 977:. 949:: 922:. 918:: 891:) 887:( 870:. 864:: 845:) 841:( 824:. 818:: 799:. 795:: 768:. 764:: 737:. 709:: 682:. 654:: 627:) 623:( 606:. 600:: 581:. 553:: 526:. 514:: 487:. 474:: 458:" 454:" 69:)

Index


Conservation status
Vulnerable
IUCN 2.3
Scientific classification
Edit this classification
Plantae
Bryophyta
Takakiophytina
Takakiopsida
Takakiales
Takakiaceae
Takakia
Binomial name
Synonyms
Takakia
Takakiaceae
William Mitten
Lepidozia
columella
calyptra
seta
rhizoids
stomata
operculum
China
India
Nepal
United States
habitats

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