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Teleology in biology

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824:' for apparently goal-directed biological phenomena. For Pittendrigh, the notion of 'adaptation' in biology, however it is defined, necessarily "connote that aura of design, purpose, or end-directedness, which has, since the time of Aristotle, seemed to characterize the living thing" This association with Aristotle, however, is problematic, because it meant that the study of adaptation would inevitably be bound up with teleological explanations. Pittendrigh sought to preserve the aspect of design and purpose in biological systems, while denying that this design can be understood as a causal principle. The confusion, he says, would be removed if we described these systems "by some other term, like 'teleonomic,' in order to emphasize that the recognition and description of end-directedness does not carry a commitment to Aristotelian teleology as an efficient causal principle." 749:
of their own action; for example the behavior of picking up a pen can be explained by reference to the agent's desire to write. Ayala extends this type of teleological explanation to non-human animals by noting that A deer running away from a mountain lion. . . has at least the appearance of purposeful behavior." Second, teleological explanations are useful for systems that have a mechanism for self-regulation despite fluctuations in environment; for example, the self-regulation of body temperature in animals. Finally, they are appropriate "in reference to structures anatomically and physiologically designed to perform a certain function. "
756:, also rejects the idea that teleological arguments are inadmissible because they cannot be causal. For Nagel, teleological arguments must be consistent because they can always be reformulated as non-teleological arguments. The difference between the two is, for Ayala, merely one of emphasis. Nagel writes that while teleological arguments focus on "the consequences for a given system of a constituent part or process," the equivalent non-teleological arguments focus on ""some of the conditions ... under which the system persists in its characteristic organization and activities." However, 696:
biology courses have incorporated exercises requiring students to rephrase such sentences so that they do not read teleologically. Nevertheless, biologists still frequently write in a way which can be read as implying teleology, even though that is not their intention. John Reiss argues that evolutionary biology can be purged of apparent teleology by rejecting the pre-Darwinian watchmaker analogy for natural selection; other arguments against this analogy have also been promoted by writers such as the evolutionary biologist
1446:"With the integration of Mendelian genetics and population genetics into evolutionary theory in the 1930s a new generation of biologists applied mathematical techniques to investigate how changes in the frequency of genes in populations combined with natural selection could produce species change. This demonstrated that Darwinian natural selection was the primary mechanism for evolution and that other models of evolution, such as neo-Lamarckism and orthogenesis, were invalid." 608: 450: 225: 2876: 33: 651: 674:
Fourthly, the teleological explanation of adaptation is uncomfortable because it seems to require backward causation, in which existing traits are explained by future outcomes; because it seems to attribute the action of a conscious mind when none is assumed to be present in an organism; and because,
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Other philosophers of biology argue instead that biological teleology is irreducible, and cannot be removed by any simple process of rewording. Francisco Ayala specified three separate situations in which teleological explanations are appropriate. First, if the agent consciously anticipates the goal
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for describing any function which offers an evolutionary advantage through natural selection. For example, the zoologist S. H. P. Madrell wrote that "the proper but cumbersome way of describing change by evolutionary adaptation substituted by shorter overtly teleological statements" for the sake of
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described this claim as a myth promoted by misinterpretations of his discussions, and emphasized the distinction between using teleological metaphors and actually being teleological. Michael Heads, on the other hand, describes a change in Darwin's thinking about evolution that can be traced from the
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to later volumes. For Heads, Darwin was originally a far more teleological thinker, but over time, "learned to avoid teleology." Heads cites a letter Darwin wrote in 1872, in which he downplayed the role of natural selection as a causal force on its own in explaining biological adaptation, and
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affirmed that the teleologicality of the language of biology and other fields derives from the logical structure of their background theories, and not merely from the use of teleological locutions such as "function" and "in order to". He stated that "To replace talk about function by talk about
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Statements which imply that nature has goals, for example where a species is said to do something "in order to" achieve survival, appear teleological, and therefore invalid to evolutionary biologists. It is however usually possible to rewrite such sentences to avoid the apparent teleology. Some
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that this is "the best of all possible worlds", in other words that every trait is perfectly suited to its functions. However, all that evolutionary biology requires is the weaker claim that one trait is at least slightly better in a certain context than another, and hence is selected for.
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has a function by analysing the process of selection that led to it. Therefore, Neander argues, any talk of functions must be posterior to natural selection, function must be defined by reference to the history of a species, and teleology cannot be avoided. The evolutionary biologist
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A trait which persists in a population is often assumed by biologists to have been selected for in the course of evolution, raising the question of how the trait achieves this. Biologists call any such mechanism the function of the trait, using phrases like "A
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or argument from design, namely that organs functioned well for their apparent purpose, so they were well-designed, so they must have been designed by a benevolent creator. For example, the eye had the function of seeing, and contained features like the
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A fifth reason concerns students rather than researchers: Gonzalez Galli argues that since people naturally imagine that evolution has a purpose or direction, then the use of teleological language by scientists may act as an obstacle to students when
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Phrases used by biologists like "a function of ... is to ..." or "is designed for" are teleological at least in language. The presence of real or apparent teleology in explanations of natural selection is a controversial aspect of the
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where animals in particular have been supposed to influence their own evolution through their intentions, though Lamarck himself spoke rather of habits of use, and the belief that his thinking was teleological has been challenged.
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or evolutionary progress. Such goal-directedness implies a long-term teleological force; some supporters of orthogenesis considered it to be a spiritual force, while others held that it was purely biological. For example, the
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saving space, but that this "should not be taken to imply that evolution proceeds by anything other than from mutations arising by chance, with those that impart an advantage being retained by natural selection."
832:, arguing that evolution only involved the material and formal but not the efficient cause. Mayr proposed to use the term only for "systems operating on the basis of a program of coded information." 2076:
Bednarczyk, A (2009), "[Jean-Baptiste de Lamarck (1744-1829). A dispute on the mechanism of evolution. On the bicentenary of the publication of Philosophie Zoologique (1809)]",
772:. So, for example, it is not possible to say that anything that simply winks into existence, without going through a process of selection, actually has functions. We decide whether an 578:, much to the consternation of some writers, and as an explanatory style it remains controversial. There are various reasons for discomfort with teleology among biologists. 540:. However, there is also a historical question, namely, did the trait arise at the same time as bird flight? Unfortunately for the hypothesis, this seems not to be so: 1578:
The hands and feet of all primates, except for humans, are designed for grasping. Humans have hands designed for grasping, but not feet! Humans have opposable thumbs.
532:. That would require three things: that the trait of having feathers is heritable; that the trait does serve the function of flight; and that the trait increases the 2798: 265:
stated that his intention was "to illustrate the glory of God in the knowledge of the works of nature or creation". Natural theology presented forms of the
1180:(2006). "Review of 'The Blasphemy of Intelligent Design: Creationism's Trojan Horse. The Wedge of Intelligent Design' by Barbara Forrest; Paul R. Gross". 814:
perspective on living beings." In her view of Kant, teleology implies something that cannot be explained by science, but only understood through analogy.
300:, and have often supposed that it had some kind of goal or direction (towards which the life force was striving, if they also believed in that), known as 2601:
Angel, J. Lawrence (June 1961). "Behavior and evolution. By Anne Roe and George G. Simpson, eds. vii + 557 pp. Yale University Press, New Haven, 1958".
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Heads, Michael (June 2009). "Darwin's changing views on evolution: from centres of origin and teleology to vicariance and incomplete lineage sorting".
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rather than actual goals, whether conscious or not. Some biologists and religious thinkers held that evolution itself was somehow goal-directed (
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observed that "Teleology is like a mistress to a biologist: he cannot live without her but he's unwilling to be seen with her in public."
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Neander, Karen (1998). "Functions as Selected Effects: The Conceptual Analyst's Defense," in C. Allen, M. Bekoff & G. Lauder (Eds.),
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selection is not to eliminate teleology but to rephrase it". However, Wimsatt argues that this thought does not mean an appeal to
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both assumed the existence of God and used the appearance of function in nature to argue for the existence of God. The English
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mechanisms of evolution were discovered, the hypothesis of orthogenesis was largely abandoned by biologists, especially with
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Religious thinkers and biologists have supposed that evolution was driven by some kind of life force, a philosophy known as
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Firstly, the concept of adaptation is itself controversial, as it can be taken to imply, as the evolutionary biologists
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Natural Theology: Or, Evidences of the Existence and Attributes of the Deity, Collected from the Appearances of Nature
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Natural Theology, or Evidences of the Existence and Attributes of the Deity collected from the Appearances of Nature
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in biology has attracted criticism, and attempts have been made to teach students to avoid teleological language.
2979: 2951: 789: 339: 331: 313: 1617:"Gradual assembly of avian body plan culminated in rapid rates of evolution across the dinosaur-bird transition" 3024: 1881:(1979). "The spandrels of San Marco and the Panglossian paradigm: a critique of the adaptationist programme". 1682:
Xu, X.; Wang, K.; Zhang, K.; Ma, Q.; Xing, L.; Sullivan, C.; Hu, D.; Cheng, S.; Wang, S.; et al. (2012).
663:, along with a vitalist life-force and directed orthogenetic evolution, has been rejected by most biologists. 1049:
Philosophia Rationalis Sive Logica: Methodo Scientifica Pertractata Et Ad Usum Scientiarum Atque Vitae Aptata
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argues that the presence of a complex mechanism like a watch implies the existence of a conscious designer.
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Both Pittendrigh and Mayr endorsed teleology in biology as an inherent part of evolutionary thinking.
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Various commentators view the teleological phrases used in modern evolutionary biology as a type of
3069: 3009: 2821: 2519:(1998). "Teleological explanations in evolutionary biology". In Allen, Colin; Bekoff, Marc (eds.). 840: 607: 343: 309: 186: 555:. Biologists may describe both the co-option and the earlier adaptation in teleological language. 3262: 3257: 3236: 3231: 3205: 3125: 3109: 3104: 3014: 2934: 2927: 2712: 2655: 2583: 2516: 2490: 2474: 2430: 2281: 2243: 2009: 1914: 1807: 1722: 1189: 1177: 1118: 1088: 995: 835: 783: 765: 757: 684: 676: 656: 624: 612: 552: 544: 491: 484: 472: 198: 178: 133: 94: 44: 17: 449: 278:
that assisted with seeing; therefore, ran the argument, it had been designed for that purpose.
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Nevertheless, biologists still often write about evolution as if organisms had goals, and some
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Ayala, Francisco J. (March 1970). "Teleological Explanations in Evolutionary Biology".
2264:(1994). "Darwin's language may seem teleological, but his thinking is another matter". 2053: 2028: 1486: 1013: 949: 876: 637: 628: 423: 369: 236: 78: 1305:
The Eclipse of Darwinism: Anti-Darwinian Evolution Theories in the Decades around 1900
991: 732:, which he asserts is "wholly teleological", Darwinian evolution is not teleological. 536:
of the organisms that have it. Feathers clearly meet these three conditions in living
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The Blind Watchmaker: Why the Evidence of Evolution Reveals a Universe Without Design
1947: 1752: 1510: 1241: 802: 797: 761: 725: 642: 426:, is the differential survival and reproduction of individuals due to differences in 388: 325: 316: 167: 2434: 2196: 1851: 1834: 1726: 999: 760:
argued that teleological statements are more explanatory and cannot be disposed of.
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argued that Du Noüy and Sinnott were promoting religious versions of evolution. The
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Thirdly, attributing purposes to adaptations risks confusion with popular forms of
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often use similar teleological formulations that invoke purpose, but these imply
3190: 3064: 2917: 2689: 2112:"The Role of Teleological Thinking in Learning the Darwinian Model of Evolution" 2044: 1615:
Brusatte, Stephen L.; Lloyd, Graeme T.; Wang, Steve C.; Norell, Mark A. (2014).
1506: 829: 660: 529: 454: 376: 224: 194: 86: 687:. Such language, he argues, should be removed to make teaching more effective. 3215: 3180: 3039: 3029: 2999: 2907: 2875: 2768: 2684: 2128: 2111: 1989: 1633: 1616: 825: 778: 667: 548: 521: 513: 480: 458: 438: 275: 271: 62: 32: 27:
Use of language of goal-directedness in the context of evolutionary adaptation
2771:(1974) Boston Studies in the Philosophy of Science, Volume XIV, pages 91–117. 2622: 2579: 2460: 2426: 2320: 2089: 1282:
From Cosmology to Ecology: The Monist World-view in Germany from 1770 to 1930
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is an observable structure or other feature of an organism (for example, an
462: 427: 419: 410: 206: 190: 163: 158: 122: 74: 66: 59: 37: 2097: 2062: 1902: 1860: 1718: 1642: 1465:. Harvard University Press. Chapter 7, section "Synthesis as Restriction". 810:
in the search for causal explanations of nature and ... an inevitable
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The structure of science: Problems in the logic of scientific explanation
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and beneficent intentions of a creator, as in the writings of John Ray.
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respectively. Their views were heavily criticized as non-scientific; the
320: 297: 291: 262: 239:, used teleological arguments to illustrate the glory of God from nature. 232: 110: 40: 2776: 1910: 1710: 1193: 2277: 2239: 2163:"Is Evolutionary Biology Infected with Invalid Teleological Reasoning?" 2013: 900: 811: 599: 525: 503: 113:
versions, driven by a purposeful life force. With evolution working by
2715:(1972). "Teleology and the logical structure of function statements". 2367:
Madrell, S.H.P. (1998). "Why are there no insects in the open sea?".
517: 379:" in what he called "directed additivity". With the emergence of the 366: 354: 2005: 2418: 1968: 1951: 1684:"A gigantic feathered dinosaur from the Lower Cretaceous of China" 606: 537: 448: 223: 189:
in 1728. The concept derives from the ancient Greek philosophy of
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Gonzalez Galli, Leonardo Martin; Meinardi, Elsa N. (March 2011).
2029:"Lamarck, Evolution, and the Inheritance of Acquired Characters" 1324:
Koch, Leo Francis (1957). "Vitalistic-Mechanistic Controversy".
547:, but many of them did not fly. Feathers can be described as an 506: 51:, helping the springbok to survive and allowing it to reproduce. 2780: 58:
is the use of the language of goal-directedness in accounts of
899:, to explain traits such as the colourful "tail" train of the 82: 2545:
Nature's Purposes: Analyses of Function and Design in Biology
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Nature's purposes: Analyses of Function and Design in Biology
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instead gave more weight to "laws of growth," that operate .
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have argued that Darwin was a teleologist, while others like
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Biosemiotics: Information, Codes and Signs in Living Systems
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Secondly, teleology is linked to the pre-Darwinian idea of
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argued that evolution was aiming for a supposed spiritual "
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International Journal of Emerging Technologies and Society
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Origins: A Short Etymological Dictionary of Modern English
1758:. The Belknap Press of Harvard University Press. p.  461:
for this task, having evolved for an earlier purpose in
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believed in a teleological force in nature, whereas the
185:, "a branch of learning", was coined by the philosopher 1594:"Understanding Evolution: Qualifying as an adaptation" 1394:
De Chardin, Pierre Teilhard. (2003, reprint edition).
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continued Ray's tradition with books such as his 1713
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developed vitalist evolutionary philosophies known as
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consider that teleological language is unavoidable in
2650:. New Haven: Yale University Press. pp. 390–416. 2336:"The Misnomer of Transhumanism as Directed Evolution" 1566:"Primates – marmosets, monkeys, apes, lemurs, humans" 323:, arguing for a creative force in evolution known as 1833:
Ribeiro, Manuel Gustavo Leitao; et al. (2015).
1658:"Largest feathered dinosaur yet discovered in China" 93:
to have been made to enable them to carry out their
3224: 3118: 2970: 2883: 2814: 2559:"Teleology in Biology : A Kantian Perspective" 1307:. The Johns Hopkins University Press. pp. 116–117. 2751: 2688: 2195: 1751: 1544:(4th ed.). Jones and Bartlett. pp. 4–6. 1434:Montgomery, Georgina M.; Largent, Mark A. (2015). 828:criticised Pittendrigh's confusion of Aristotle's 509:is designed (by natural selection) for grasping." 1372:The Phenomenon of Teilhard: Prophet for a New Age 781:likewise stated that "adaptedness ... is an 675:as a result, adaptation looks impossible to test 418:, introduced in 1859 as the central mechanism of 2717:Studies in the History and Philosophy of Science 1789:Teleology: The explanation that bedevils biology 1517:(1st ed.). Roberts and Company Publishers. 221:, not least for its echoes of natural theology. 89:them; their features such as eyes were taken by 1538:Hall, Brian K.; Hallgrímsson, Benedikt (2008). 1358:This View of Life: The World of an Evolutionist 1144: 1142: 752:Ayala, relying on work done by the philosopher 623:, that the natural world gives evidence of the 2634: 2632: 1436:A Companion to the History of American Science 2792: 2400: 2398: 1792:. Oxford University Press. pp. 143–155. 806:that teleology in biology is important as "a 8: 1883:Proceedings of the Royal Society of London B 1781: 1779: 1250:. Gould, Kendall, and Lincoln. pp. 18–. 764:similarly argued that the modern concept of 43:. A biologist might argue that this has the 2648:Adaptation, natural selection, and behavior 2148:Not by Design: Retiring Darwin's Watchmaker 1984: 1982: 1786:Hanke, David (2004). Cornwell, John (ed.). 1588: 1586: 1374:. Mercer University Press. pp. 60–64. 1182:History and Philosophy of the Life Sciences 895:Darwin also introduced a second mechanism, 574:Apparent teleology is a recurring issue in 2799: 2785: 2777: 1416:Genetics, Paleontology, and Macroevolution 1360:. Harcourt, Brace & World. pp. 213–233 1094:The Lagoon: How Aristotle Invented Science 1052:. Prostat in officina libraria Rengeriana. 944: 942: 940: 938: 936: 934: 932: 930: 928: 81:, organisms were seen as existing because 2603:American Journal of Physical Anthropology 2178: 2127: 2052: 1850: 1632: 1418:. Cambridge University Press. pp. 14–16. 1125:. Sinauer Associates. pp. 341, 342. 1064:"Presuppositions of Aristotle's Politics" 394:The Genetical Theory of Natural Selection 2141: 2139: 1934:Nineteenth-Century Science: An Anthology 1354:Evolutionary Theology: The New Mysticism 1205: 1203: 2679: 2677: 1973:The Oxford Handbook of Natural Theology 1754:Evolution: The First Four Billion Years 1172: 1170: 924: 888: 728:, an ideology that aims to improve the 502:that they have been detected", or "The 2663: 2653: 2498: 2488: 1815: 1805: 1596:. University of California at Berkeley 1969:"Nineteenth Century Natural Theology" 703:Some philosophers of biology such as 457:, but they were co-opted rather than 453:Feathers today serve the function of 7: 2226:(1993). "Darwin was a Teleologist". 2078:Kwartalnik Historii Nauki I Techniki 1839:História, Ciências, Saúde-Manguinhos 1462:The Structure of Evolutionary Theory 36:"Behaviour with a purpose": a young 2898:Alternatives to Darwinian evolution 2369:The Journal of Experimental Biology 1068:Stanford Encyclopedia of Philosophy 954:Stanford Encyclopedia of Philosophy 589:argued, that biologists agree with 1956:. Oxford: Oxford University Press. 722:Sandra Day O'Connor College of Law 25: 2697:. New York: Free Press. pp.  2485:. W.H. Freeman. pp. 497–504. 2150:. University of California Press. 2116:Evolution: Education and Outreach 1992:(1992). "The idea of teleology". 1352:Simpson, George Gaylord. (1964). 1264:. Nova Science Publishers. p. 7. 950:"Teleological Notions in Biology" 2874: 2754:Philosophy of Biological Science 2313:10.1111/j.1365-2699.2009.02127.x 2167:Philosophy and Theory in Biology 1656:Whitfield, John (4 April 2012). 1398:. Sussex Academic Press. p. 65. 1097:. Bloomsbury. pp. 272–275. 691:Removable teleological shorthand 685:learning about natural selection 197:(the purpose) of a thing is its 18:Teleological language in biology 2455:. Harcourt, Brace & World. 1994:Journal of the History of Ideas 1852:10.1590/S0104-59702015005000003 1515:Evolution: Making Sense of Life 77:has also been proposed. Before 2027:Burkhardt, Richard W. (2013). 1936:. Broadview Press. p. 18. 559:Status in evolutionary biology 1: 2690:"Cause and effect in biology" 2547:(pp. 313–333). The MIT Press. 1967:Eddy, Matthew Daniel (2013). 1414:Levinton, Jeffrey S. (2001). 1280:Jacobsen, Eric Paul. (2005). 1151:The English Parson-Naturalist 992:10.1016/S0003-3472(86)80052-5 391:'s argument in his 1930 book 381:modern evolutionary synthesis 2737:10.1016/0039-3681(72)90014-3 2557:Breitenbach, Angela (2009). 2180:10.3998/ptb.6959004.0002.005 1260:Barbieri, Marcello. (2013). 2045:10.1534/genetics.113.151852 1750:; Travis, J., eds. (2009). 1495:(1st ed.). p. 89. 1149:Armstrong, Patrick (2000). 1022:. Routledge. p. 4187. 796:Angela Breitenbach, from a 73:find problematic. The term 3289: 2349:(1): 71–78. Archived from 1212:"The Argument from Design" 641:. They in turn influenced 565:Tinbergen's four questions 562: 478: 404: 373:Pierre Teilhard de Chardin 285: 247: 156: 2952:Evolutionary epistemology 2872: 2483:Teleological explanations 2194:Dawkins, Richard (1987). 2129:10.1007/s12052-010-0272-7 1634:10.1016/j.cub.2014.08.034 1153:. Gracewing. p. 46. 820:coined the similar term ' 720:Andrew Askland, from the 468:Sinornithosaurus millenii 319:linked orthogenesis with 2580:10.1515/9783110196672.31 2334:Askland, Andrew (2011). 2161:Depew, David J. (2010). 1541:Strickberger's Evolution 1492:On the Origin of Species 1244:; Paxton, James (1837). 714:On the Origin of Species 500:communicate to predators 314:spiritualist philosopher 2972:Philosophers of biology 2693:. In Lerner, D. (ed.). 2644:Simpson, George Gaylord 2615:10.1002/ajpa.1330190215 2381:10.1242/jeb.201.17.2461 2301:Journal of Biogeography 2146:Reiss, John O. (2009). 1370:Lane, David H. (1996). 800:perspective, argues in 282:Goal-directed evolution 136:and biologists such as 130:philosophers of biology 99:Evolutionary biologists 71:philosophers of science 49:signalling to predators 2266:Biology and Philosophy 2228:Biology and Philosophy 1903:10.1098/rspb.1979.0086 1327:The Scientific Monthly 1284:. p. 100. Peter Lang. 787:result rather than an 616: 570:Reasons for discomfort 545:dinosaurs had feathers 476: 363:George Gaylord Simpson 340:Pierre Lecomte du Noüy 240: 52: 3273:Philosophy of biology 2866:Evolutionary taxonomy 2808:Philosophy of biology 2640:Pittendrigh, Colin S. 2407:Philosophy of Science 1932:Weber, A. S. (2000). 766:biological 'function' 744:Irreducible teleology 647:teleological argument 645:who wrote a detailed 610: 563:Further information: 452: 267:teleological argument 227: 219:philosophy of biology 35: 3268:Evolutionary biology 2262:Ghiselin, Michael T. 1396:The Human Phenomenon 1216:Princeton University 1123:Evolutionary Biology 1044:Wolff, Christian von 655:, starting with the 576:evolutionary biology 528:are adaptations for 142:evolutionary biology 87:designed and created 56:Teleology in biology 3010:Peter Godfrey-Smith 2729:1972SHPSA...3....1. 2713:Wimsatt, William C. 2475:Ayala, Francisco J. 1895:1979RSPSB.205..581G 1711:10.1038/nature10906 1703:2012Natur.484...92X 1340:1957SciMo..85..245K 1178:Ayala, Francisco J. 1119:Futuyma, Douglas J. 1089:Leroi, Armand Marie 841:backwards causation 724:claims that unlike 597:in his 1759 satire 498:by antelopes is to 344:Edmund Ware Sinnott 310:Karl Ernst von Baer 203:Aristotle's biology 187:Christian von Wolff 177:"end, purpose" and 119:inherited variation 3237:History of biology 3232:Philosophy of mind 3206:John Maynard Smith 3126:Francisco J. Ayala 3110:William C. Wimsatt 3105:Gerard Verschuuren 3015:James R. Griesemer 2278:10.1007/BF00850377 2240:10.1007/bf00857687 1457:Gould, Stephen Jay 836:William C. Wimsatt 657:watchmaker analogy 617: 613:watchmaker analogy 485:Function (biology) 477: 332:Creative Evolution 241: 230:natural theologian 205:does not envisage 97:, such as seeing. 53: 3245: 3244: 3176:Humberto Maturana 3161:Stephen Jay Gould 3055:Roberta Millstein 3020:Paul E. Griffiths 2750:Hull, D. (1973). 2642:(1958). Roe, A.; 2523:. The MIT Press. 2375:(17): 2461–2464. 2356:on 22 April 2017. 1889:(1161): 581–598. 1879:Lewontin, Richard 1875:Gould, Stephen J. 1799:978-0-19-860778-6 1627:(20): 2386–2392. 1551:978-1-4496-4722-3 1524:978-1-936221-17-2 1511:Emlen, Douglas J. 1459:(21 March 2002). 1444:978-1-4051-5625-7 1438:. Wiley. p. 218. 1313:978-0-8018-4391-4 1290:978-0-8204-7231-7 1270:978-1-60021-612-1 1160:978-0-85244-516-7 1132:978-0-87893-189-7 1104:978-1-4088-3622-4 872:Fitness landscape 853:anti-reductionist 818:Colin Pittendrigh 770:natural selection 649:for God in 1802, 416:Natural selection 407:Natural selection 401:Natural selection 342:and the botanist 260:parson-naturalist 211:natural selection 115:natural selection 103:natural selection 16:(Redirected from 3280: 3196:Joan Roughgarden 3166:Richard Lewontin 3151:Michael Ghiselin 3100:Francisco Varela 3095:Alfred I. Tauber 3050:Jane Maienschein 2878: 2801: 2794: 2787: 2778: 2772: 2766: 2760: 2759: 2758:. Prentice-Hall. 2757: 2747: 2741: 2740: 2709: 2703: 2702: 2695:Cause and effect 2692: 2681: 2672: 2671: 2665: 2661: 2659: 2651: 2636: 2627: 2626: 2598: 2592: 2591: 2563: 2554: 2548: 2541: 2535: 2534: 2517:Ayala, Francisco 2513: 2507: 2506: 2500: 2496: 2494: 2486: 2471: 2465: 2464: 2445: 2439: 2438: 2402: 2393: 2392: 2364: 2358: 2357: 2355: 2340: 2331: 2325: 2324: 2307:(6): 1018–1026. 2296: 2290: 2289: 2258: 2252: 2251: 2224:Lennox, James G. 2220: 2214: 2213: 2201: 2191: 2185: 2184: 2182: 2158: 2152: 2151: 2143: 2134: 2133: 2131: 2107: 2101: 2100: 2073: 2067: 2066: 2056: 2024: 2018: 2017: 1986: 1977: 1976: 1964: 1958: 1957: 1953:Natural Theology 1944: 1938: 1937: 1929: 1923: 1922: 1871: 1865: 1864: 1854: 1845:(4): 1321–1333. 1830: 1824: 1823: 1817: 1813: 1811: 1803: 1783: 1774: 1773: 1757: 1744: 1738: 1737: 1735: 1729:. Archived from 1688: 1679: 1673: 1672: 1670: 1668: 1662:Nature News Blog 1653: 1647: 1646: 1636: 1612: 1606: 1605: 1603: 1601: 1590: 1581: 1580: 1575: 1573: 1562: 1556: 1555: 1535: 1529: 1528: 1503: 1497: 1496: 1483: 1477: 1476: 1453: 1447: 1432: 1426: 1412: 1406: 1392: 1386: 1385: 1367: 1361: 1350: 1344: 1343: 1321: 1315: 1298: 1292: 1278: 1272: 1258: 1252: 1251: 1238: 1232: 1231: 1229: 1227: 1218:. Archived from 1207: 1198: 1197: 1174: 1165: 1164: 1146: 1137: 1136: 1115: 1109: 1108: 1085: 1079: 1078: 1076: 1074: 1060: 1054: 1053: 1040: 1034: 1033: 1010: 1004: 1003: 980:Animal Behaviour 972: 966: 965: 963: 961: 946: 913: 910: 904: 897:sexual selection 893: 860:J. B. S. Haldane 712:first volume of 709:Michael Ghiselin 633:Physico-Theology 625:conscious design 621:natural theology 587:Richard Lewontin 583:Stephen J. Gould 256:natural theology 250:Natural theology 244:Natural theology 138:J. B. S. Haldane 91:natural theology 21: 3288: 3287: 3283: 3282: 3281: 3279: 3278: 3277: 3248: 3247: 3246: 3241: 3220: 3141:Richard Dawkins 3131:Patrick Bateson 3114: 3060:Sandra Mitchell 2966: 2879: 2870: 2810: 2805: 2775: 2767: 2763: 2749: 2748: 2744: 2711: 2710: 2706: 2683: 2682: 2675: 2662: 2652: 2638: 2637: 2630: 2600: 2599: 2595: 2561: 2556: 2555: 2551: 2542: 2538: 2531: 2515: 2514: 2510: 2497: 2487: 2473: 2472: 2468: 2447: 2446: 2442: 2404: 2403: 2396: 2366: 2365: 2361: 2353: 2338: 2333: 2332: 2328: 2298: 2297: 2293: 2260: 2259: 2255: 2222: 2221: 2217: 2210: 2193: 2192: 2188: 2160: 2159: 2155: 2145: 2144: 2137: 2109: 2108: 2104: 2075: 2074: 2070: 2026: 2025: 2021: 2006:10.2307/2709913 1988: 1987: 1980: 1966: 1965: 1961: 1946: 1945: 1941: 1931: 1930: 1926: 1873: 1872: 1868: 1832: 1831: 1827: 1814: 1804: 1800: 1785: 1784: 1777: 1770: 1746: 1745: 1741: 1733: 1697:(7392): 92–95. 1686: 1681: 1680: 1676: 1666: 1664: 1655: 1654: 1650: 1621:Current Biology 1614: 1613: 1609: 1599: 1597: 1592: 1591: 1584: 1571: 1569: 1564: 1563: 1559: 1552: 1537: 1536: 1532: 1525: 1505: 1504: 1500: 1487:Darwin, Charles 1485: 1484: 1480: 1473: 1455: 1454: 1450: 1433: 1429: 1413: 1409: 1393: 1389: 1382: 1369: 1368: 1364: 1351: 1347: 1323: 1322: 1318: 1301:Bowler, Peter J 1299: 1295: 1279: 1275: 1259: 1255: 1240: 1239: 1235: 1225: 1223: 1222:on 16 July 2019 1210:Rosen, Gideon. 1209: 1208: 1201: 1176: 1175: 1168: 1161: 1148: 1147: 1140: 1133: 1117: 1116: 1112: 1105: 1087: 1086: 1082: 1072: 1070: 1062: 1061: 1057: 1042: 1041: 1037: 1030: 1014:Partridge, Eric 1012: 1011: 1007: 974: 973: 969: 959: 957: 948: 947: 926: 922: 917: 916: 911: 907: 894: 890: 885: 868: 793:goal-seeking." 758:Francisco Ayala 746: 730:human condition 705:James G. Lennox 698:Richard Dawkins 693: 595:Doctor Pangloss 572: 567: 561: 487: 479:Main articles: 447: 413: 405:Main articles: 403: 383:, in which the 360:palaeontologist 294: 286:Main articles: 284: 254:Before Darwin, 252: 246: 161: 155: 150: 134:Francisco Ayala 28: 23: 22: 15: 12: 11: 5: 3286: 3284: 3276: 3275: 3270: 3265: 3260: 3250: 3249: 3243: 3242: 3240: 3239: 3234: 3228: 3226: 3222: 3221: 3219: 3218: 3213: 3208: 3203: 3198: 3193: 3188: 3183: 3178: 3173: 3168: 3163: 3158: 3156:François Jacob 3153: 3148: 3143: 3138: 3136:Charles Darwin 3133: 3128: 3122: 3120: 3116: 3115: 3113: 3112: 3107: 3102: 3097: 3092: 3087: 3082: 3080:Sahotra Sarkar 3077: 3072: 3070:Alex Rosenberg 3067: 3062: 3057: 3052: 3047: 3042: 3037: 3035:Philip Kitcher 3032: 3027: 3022: 3017: 3012: 3007: 3005:Marjorie Grene 3002: 2997: 2992: 2990:Daniel Dennett 2987: 2985:Lindley Darden 2982: 2976: 2974: 2968: 2967: 2965: 2964: 2959: 2954: 2949: 2944: 2943: 2942: 2937: 2932: 2931: 2930: 2920: 2915: 2910: 2905: 2895: 2889: 2887: 2881: 2880: 2873: 2871: 2869: 2868: 2863: 2862: 2861: 2851: 2846: 2841: 2840: 2839: 2834: 2829: 2818: 2816: 2812: 2811: 2806: 2804: 2803: 2796: 2789: 2781: 2774: 2773: 2761: 2742: 2704: 2673: 2628: 2609:(2): 218–219. 2593: 2549: 2536: 2530:978-0262510974 2529: 2508: 2479:Dobzhansky, T. 2466: 2440: 2419:10.1086/288276 2394: 2359: 2326: 2291: 2272:(4): 489–492. 2253: 2234:(4): 409–421. 2215: 2208: 2202:. W W Norton. 2186: 2153: 2135: 2122:(1): 145–152. 2102: 2084:(3–4): 31–98, 2068: 2039:(4): 793–805. 2019: 2000:(1): 117–135. 1990:Mayr, Ernst W. 1978: 1959: 1948:Paley, William 1939: 1924: 1866: 1825: 1798: 1775: 1768: 1739: 1736:on 2012-04-17. 1674: 1648: 1607: 1582: 1557: 1550: 1530: 1523: 1498: 1478: 1472:978-0674006133 1471: 1448: 1427: 1407: 1387: 1380: 1362: 1345: 1334:(5): 245–255. 1316: 1293: 1273: 1253: 1242:Paley, William 1233: 1199: 1188:(3): 409–421. 1166: 1159: 1138: 1131: 1110: 1103: 1080: 1055: 1035: 1028: 1005: 986:(3): 663–684. 967: 923: 921: 918: 915: 914: 905: 887: 886: 884: 881: 880: 879: 877:Teleomechanism 874: 867: 864: 858:The biologist 745: 742: 692: 689: 638:Astro-Theology 629:William Derham 571: 568: 560: 557: 446: 443: 424:Charles Darwin 402: 399: 370:paleontologist 283: 280: 248:Main article: 245: 242: 237:William Derham 157:Main article: 154: 151: 149: 146: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 3285: 3274: 3271: 3269: 3266: 3264: 3261: 3259: 3256: 3255: 3253: 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2867: 2864: 2860: 2857: 2856: 2855: 2852: 2850: 2847: 2845: 2842: 2838: 2837:Kin selection 2835: 2833: 2830: 2828: 2825: 2824: 2823: 2820: 2819: 2817: 2813: 2809: 2802: 2797: 2795: 2790: 2788: 2783: 2782: 2779: 2770: 2765: 2762: 2756: 2755: 2746: 2743: 2738: 2734: 2730: 2726: 2722: 2718: 2714: 2708: 2705: 2700: 2696: 2691: 2686: 2680: 2678: 2674: 2669: 2657: 2649: 2645: 2641: 2635: 2633: 2629: 2624: 2620: 2616: 2612: 2608: 2604: 2597: 2594: 2589: 2585: 2581: 2577: 2573: 2569: 2568: 2567:Kant Yearbook 2560: 2553: 2550: 2546: 2540: 2537: 2532: 2526: 2522: 2518: 2512: 2509: 2504: 2492: 2484: 2480: 2476: 2470: 2467: 2462: 2458: 2454: 2450: 2449:Nagel, Ernest 2444: 2441: 2436: 2432: 2428: 2424: 2420: 2416: 2412: 2408: 2401: 2399: 2395: 2390: 2386: 2382: 2378: 2374: 2370: 2363: 2360: 2352: 2348: 2344: 2337: 2330: 2327: 2322: 2318: 2314: 2310: 2306: 2302: 2295: 2292: 2287: 2283: 2279: 2275: 2271: 2267: 2263: 2257: 2254: 2249: 2245: 2241: 2237: 2233: 2229: 2225: 2219: 2216: 2211: 2209:9780393022162 2205: 2200: 2199: 2190: 2187: 2181: 2176: 2172: 2168: 2164: 2157: 2154: 2149: 2142: 2140: 2136: 2130: 2125: 2121: 2117: 2113: 2106: 2103: 2099: 2095: 2091: 2087: 2083: 2080:(in Polish), 2079: 2072: 2069: 2064: 2060: 2055: 2050: 2046: 2042: 2038: 2034: 2030: 2023: 2020: 2015: 2011: 2007: 2003: 1999: 1995: 1991: 1985: 1983: 1979: 1974: 1970: 1963: 1960: 1955: 1954: 1949: 1943: 1940: 1935: 1928: 1925: 1920: 1916: 1912: 1908: 1904: 1900: 1896: 1892: 1888: 1884: 1880: 1876: 1870: 1867: 1862: 1858: 1853: 1848: 1844: 1840: 1836: 1829: 1826: 1821: 1809: 1801: 1795: 1791: 1790: 1782: 1780: 1776: 1771: 1769:9780674031753 1765: 1761: 1756: 1755: 1749: 1748:Ruse, Michael 1743: 1740: 1732: 1728: 1724: 1720: 1716: 1712: 1708: 1704: 1700: 1696: 1692: 1685: 1678: 1675: 1663: 1659: 1652: 1649: 1644: 1640: 1635: 1630: 1626: 1622: 1618: 1611: 1608: 1595: 1589: 1587: 1583: 1579: 1567: 1561: 1558: 1553: 1547: 1543: 1542: 1534: 1531: 1526: 1520: 1516: 1512: 1508: 1502: 1499: 1494: 1493: 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815: 813: 809: 805: 804: 803:Kant Yearbook 799: 794: 792: 791: 786: 785: 780: 775: 771: 767: 763: 762:Karen Neander 759: 755: 750: 743: 741: 738: 733: 731: 727: 726:transhumanism 723: 718: 715: 710: 706: 701: 699: 690: 688: 686: 680: 678: 672: 669: 664: 662: 658: 654: 653: 648: 644: 643:William Paley 640: 639: 635:and his 1714 634: 630: 626: 622: 614: 609: 605: 602: 601: 596: 592: 588: 584: 579: 577: 569: 566: 558: 556: 554: 550: 546: 543: 539: 535: 531: 527: 523: 519: 515: 510: 508: 505: 501: 497: 493: 486: 482: 474: 470: 469: 464: 460: 456: 451: 444: 442: 440: 436: 433: 429: 425: 421: 417: 412: 408: 400: 398: 396: 395: 390: 389:Ronald Fisher 386: 382: 378: 374: 371: 368: 364: 361: 357: 356: 351: 350: 345: 341: 338: 334: 333: 328: 327: 322: 318: 317:Henri Bergson 315: 311: 308: 303: 299: 293: 289: 281: 279: 277: 273: 268: 264: 261: 257: 251: 243: 238: 234: 231: 226: 222: 220: 214: 212: 208: 204: 200: 196: 192: 188: 184: 180: 176: 172: 169: 165: 160: 152: 147: 145: 143: 139: 135: 131: 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O. Wilson 3201:Rolf Sattler 3090:Kim Sterelny 3075:Michael Ruse 2962:Tree of life 2956: 2913:Orthogenesis 2849:Reductionism 2764: 2753: 2745: 2720: 2716: 2707: 2694: 2647: 2606: 2602: 2596: 2574:(1): 31–56. 2571: 2565: 2552: 2544: 2539: 2520: 2511: 2482: 2469: 2452: 2443: 2410: 2406: 2372: 2368: 2362: 2351:the original 2346: 2342: 2329: 2304: 2300: 2294: 2269: 2265: 2256: 2231: 2227: 2218: 2197: 2189: 2173:(20160629). 2170: 2166: 2156: 2147: 2119: 2115: 2105: 2081: 2077: 2071: 2036: 2032: 2022: 1997: 1993: 1972: 1962: 1952: 1942: 1933: 1927: 1886: 1882: 1869: 1842: 1838: 1828: 1788: 1753: 1742: 1731:the original 1694: 1690: 1677: 1665:. Retrieved 1661: 1651: 1624: 1620: 1610: 1598:. Retrieved 1577: 1570:. Retrieved 1560: 1540: 1533: 1514: 1507:Zimmer, Carl 1501: 1491: 1481: 1460: 1451: 1435: 1430: 1415: 1410: 1395: 1390: 1371: 1365: 1357: 1353: 1348: 1331: 1325: 1319: 1304: 1296: 1281: 1276: 1261: 1256: 1246: 1236: 1224:. Retrieved 1220:the original 1185: 1181: 1150: 1122: 1113: 1093: 1083: 1071:. Retrieved 1058: 1048: 1038: 1018: 1008: 983: 979: 976:Caro, Tim M. 970: 958:. Retrieved 953: 908: 891: 857: 855:sentiments. 834: 816: 801: 795: 788: 784:a posteriori 782: 754:Ernest Nagel 751: 747: 734: 719: 713: 702: 694: 681: 673: 665: 650: 636: 632: 618: 598: 580: 573: 511: 488: 466: 414: 392: 353: 349:telefinalism 347: 337:biophysicist 335:(1907). The 330: 329:in his book 324: 307:embryologist 302:orthogenesis 295: 288:Orthogenesis 253: 235:, and later 228:The English 215: 193:, where the 182: 174: 162: 127: 107:orthogenesis 60:evolutionary 55: 54: 29: 3191:Denis Noble 3065:Susan Oyama 2980:John Beatty 2918:Mutationism 2769:Mayr, Ernst 2723:(1): 1–80. 2685:Mayr, Ernst 2664:|work= 2499:|work= 2413:(1): 1–15. 1816:|work= 830:four causes 768:depends on 677:empirically 661:creationism 530:bird flight 441:over time. 377:Omega Point 201:. However, 195:final cause 3252:Categories 3216:Jonas Salk 3181:Ernst Mayr 3119:Biologists 3040:Tim Lewens 3030:Hans Jonas 3025:David Hull 3000:Carla Fehr 2995:John Dupré 2908:Lamarckism 2844:Naturalism 1303:. (1992). 920:References 826:Ernst Mayr 812:analogical 779:Ernst Mayr 668:Lamarckism 553:insulation 549:exaptation 522:hypothesis 514:adaptation 481:Adaptation 473:insulation 471:, perhaps 445:Adaptation 439:population 326:élan vital 117:acting on 109:), and in 67:biologists 63:adaptation 3263:Causality 3258:Teleology 2957:Teleology 2947:Darwinism 2885:Evolution 2859:Emergence 2832:Dysgenics 2666:ignored ( 2656:cite book 2623:0002-9483 2588:171216322 2501:ignored ( 2491:cite book 2461:874878031 2427:0031-8248 2321:0305-0270 2286:170997321 2248:170767015 2090:0023-589X 1950:(2006) . 1818:ignored ( 1808:cite book 849:entelechy 822:teleonomy 808:heuristic 774:appendage 737:shorthand 463:theropods 432:heritable 428:phenotype 420:evolution 411:Evolution 207:evolution 191:Aristotle 164:Teleology 159:Teleology 153:Teleology 123:teleology 95:functions 75:teleonomy 38:springbok 2940:Vitalism 2935:Theistic 2928:Spandrel 2827:Eugenics 2687:(1965). 2646:(eds.). 2477:(1977). 2451:(1961). 2435:84638701 2098:20481104 2063:23908372 2033:Genetics 1861:25650703 1727:29689629 1719:22481363 1643:25264248 1513:(2013). 1489:(1859). 1226:10 April 1194:23334140 1121:(1998). 1091:(2014). 1046:(1732). 1016:(1977). 1000:53155678 866:See also 845:vitalism 790:a priori 591:Voltaire 542:theropod 526:feathers 496:stotting 492:function 321:vitalism 298:vitalism 292:Vitalism 263:John Ray 233:John Ray 199:function 166:, from 132:such as 111:vitalist 45:function 41:stotting 3225:Related 2725:Bibcode 2481:(ed.). 2389:9698580 2054:3730912 2014:2709913 1919:2129408 1891:Bibcode 1699:Bibcode 1667:4 April 1600:29 July 1572:28 July 1568:. NHPTV 1336:Bibcode 1073:28 July 960:28 July 901:peacock 798:Kantian 659:. Such 600:Candide 534:fitness 504:primate 459:adapted 385:genetic 148:Context 2854:Holism 2822:Ethics 2815:Themes 2621:  2586:  2527:  2459:  2433:  2425:  2387:  2319:  2284:  2246:  2206:  2096:  2088:  2061:  2051:  2012:  1917:  1909:  1859:  1796:  1766:  1725:  1717:  1691:Nature 1641:  1548:  1521:  1469:  1442:  1422:  1402:  1378:  1311:  1288:  1268:  1192:  1157:  1129:  1101:  1026:  998:  518:enzyme 455:flight 435:traits 367:Jesuit 355:telism 179:-λογία 79:Darwin 2699:33–50 2584:S2CID 2562:(PDF) 2431:S2CID 2354:(PDF) 2339:(PDF) 2282:S2CID 2244:S2CID 2010:JSTOR 1915:S2CID 1911:42062 1734:(PDF) 1723:S2CID 1687:(PDF) 1356:. 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Index

Teleological language in biology

springbok
stotting
function
signalling to predators
evolutionary
adaptation
biologists
philosophers of science
teleonomy
Darwin
God
designed and created
natural theology
functions
Evolutionary biologists
natural selection
orthogenesis
vitalist
natural selection
inherited variation
teleology
philosophers of biology
Francisco Ayala
J. B. S. Haldane
evolutionary biology
Teleology
Teleology
Greek

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