522:
31:
573:
589:
544:
75:
50:
497:
and
Madagascar to the northern Red Sea. A single record was reported in the Mediterranean Sea off Israel in 1986. This species occurs near coral reefs on sand or rubble substrates at depths between 1 and 15 m (3 ft 3 in and 49 ft 3 in).
521:
543:
480:
for this species is 70 cm (28 in), although 50 cm (20 in) is more typical. The body is mottled brownish or greenish dorsally, whitish ventrally. There are 3 or 4 dark bands on the
30:
468:. The supraorbital is smooth with a tiny terminal spine, there is one preocular and two parietal spines. The eyes have notable papillae on their upper surfaces. The first
1039:
1091:
464:
are smaller dorsally than they are ventrally. There is a spine on the rear of the suborbital ridge which ends with a small spine. There are small spines on the
1171:
572:
698:
737:
1013:
1065:
800:"Order Perciformes (Part 11): Suborder Platycephaloidei: Families Bembridae, Parabembridae, Hoplichthyidae, Platycephalidae and Plectrogeniidae"
1196:
1117:
782:
588:
892:
667:
444:, which means "eye", a reference to the small and inconspicuous flattened papilla on the upper surface of the eyeball this species with
460:
The tentacled flathead has an elongate body with a depressed head with 5 prominent nuchal spines. The ridges on the preoperculum and
1143:
1191:
1186:
747:
712:
1070:
74:
354:
1181:
1052:
974:
1096:
1176:
554:
449:
855:
929:
362:
246:
979:
636:
180:
1148:
961:
733:
694:
320:
406:
342:
39:
966:
452:
means "long head" and may refer to the head of this fish being similar to that of a crocodile's.
401:
206:
69:
1083:
1044:
1130:
987:
867:
778:
742:
707:
370:
227:
153:
1135:
768:
631:
461:
414:
305:
1104:
617:
559:
507:
421:
418:
308:
301:
136:
126:
377:
350:
197:
157:
116:
1165:
1078:
888:
663:
622:
494:
374:
316:
59:
54:
992:
531:
477:
465:
389:
1026:
952:
482:
469:
324:
917:
1057:
563:
311:, the flatheads. This species is in the western Indian Ocean, including the
86:
493:
The tentacled flathead is found in the western Indian Ocean from northern
404:
1109:
1000:
946:
901:
830:"Two Red Sea Flatheads (Platycephalidae) Immigrants in the Mediterranean"
772:
676:
473:
411:
106:
856:
https://ciesm.org/atlas/fishes_2nd_edition/Papilloculiceps_longiceps.pdf
1018:
1005:
579:
535:
358:
312:
297:
1031:
829:
799:
798:
Christopher
Scharpf & Kenneth J. Lazara, eds. (7 December 2021).
96:
923:
1122:
927:
823:
821:
854:). 2nd Edition. 2021. 366p. CIESM Publishers, Paris, Monaco.
485:
and the other fins are marked with large, dark blotches.
396:
is now considered to be a junior synonym of Cuvier's
936:
637:
10.2305/IUCN.UK.2018-2.RLTS.T114173325A114173342.en
369:from Madagascar and in 1956 the American zoologist
736:; Fricke, Ron & van der Laan, Richard (eds.).
697:; Fricke, Ron & van der Laan, Richard (eds.).
616:Motomura, H.; Matsuura, K. & Khan, M. (2018).
850:Atlas of Exotic Fishes in the Mediterranean Sea (
804:The ETYFish Project Fish Name Etymology Database
327:. It is the only species in the monotypic genus
472:has 9 spines and the second dorsal fin and the
506:The tentacled flathead is a well camouflaged,
476:each have 11 soft rays. The maximum published
8:
828:Daniel Golani & Adam Ben-Tuvia (1990).
806:. Christopher Scharpf and Kenneth J. Lazara
924:
658:
656:
654:
341:The tentacled flathead was first formally
48:
29:
20:
777:(5th ed.). Wiley. pp. 467–495.
635:
361:in Eritrea. In 1873 the French zoologist
771:; T. C. Grande; M. V. H. Wilson (2016).
605:
517:
738:"Genera in the family Platycephalidae"
689:
687:
611:
609:
7:
353:in 1829 from specimens collected by
1172:IUCN Red List least concern species
623:IUCN Red List of Threatened Species
558:are the two species of the family
448:, which means "head" suffixed The
14:
380:proposed a new monotypic genus,
587:
571:
542:
520:
73:
748:California Academy of Sciences
713:California Academy of Sciences
1:
1197:Taxa named by Georges Cuvier
868:"Papilloculiceps longiceps"
618:"Papilloculiceps longiceps"
254:Papilloculiceps grandidieri
1213:
630:: e.T114173325A114173342.
286:Indian Ocean crocodilefish
262:Platycephalus papilloculus
224:Platycephalus tentaculatus
967:Papilloculiceps_longiceps
938:Papilloculiceps longiceps
895:Papilloculiceps longiceps
852:Papilloculiceps longiceps
670:Papilloculiceps longiceps
550:Papilloculiceps longiceps
510:of fish and crustaceans.
386:Platycephalus grandidieri
367:Platycephalus grandidieri
281:Papilloculiceps longiceps
243:Platycephalus grandidieri
212:
205:
190:Papilloculiceps longiceps
186:
179:
70:Scientific classification
68:
46:
37:
28:
23:
891:; Pauly, Daniel (eds.).
680:. February 2022 version.
666:; Pauly, Daniel (eds.).
489:Distribution and habitat
440:, meaning "nipple", and
365:described a new species
349:by the French zoologist
555:Cymbacephalus beauforti
347:Platycephalus longiceps
216:Platycephalus longiceps
1192:Fish described in 1829
699:"Species in the genus
410:classifies within the
319:, having invaded as a
1187:Monotypic fish genera
920:on Sealife Collection
905:. April 2007 version.
734:Eschmeyer, William N.
695:Eschmeyer, William N.
284:), also known as the
235:Rogadius tentaculatus
436:is a combination of
1182:Fish of the Red Sea
774:Fishes of the World
407:Fishes of the World
363:Henri Émile Sauvage
290:Madagascar flathead
40:Conservation status
24:Tentacled flathead
918:Tentacled flathead
321:Lessepsian migrant
276:tentacled flathead
1159:
1158:
1131:Open Tree of Life
930:Taxon identifiers
784:978-1-118-34233-6
743:Catalog of Fishes
708:Catalog of Fishes
384:, with Sauvage's
371:Henry Weed Fowler
304:belonging to the
294:longhead flathead
272:
271:
266:
258:
250:
239:
231:
220:
172:P. longiceps
161:
63:
1204:
1152:
1151:
1139:
1138:
1126:
1125:
1113:
1112:
1100:
1099:
1087:
1086:
1074:
1073:
1061:
1060:
1048:
1047:
1035:
1034:
1022:
1021:
1009:
1008:
996:
995:
983:
982:
970:
969:
957:
956:
955:
925:
906:
880:
879:
877:
875:
864:
858:
848:
842:
841:
825:
816:
815:
813:
811:
795:
789:
788:
765:
759:
758:
756:
754:
730:
724:
723:
721:
719:
691:
682:
681:
660:
649:
648:
646:
644:
639:
613:
594:True colouration
591:
575:
546:
524:
415:Platycephaloidei
400:. This genus is
373:and the Israeli
264:
256:
245:
237:
226:
218:
192:
152:
78:
77:
57:
52:
51:
33:
21:
1212:
1211:
1207:
1206:
1205:
1203:
1202:
1201:
1177:Platycephalidae
1162:
1161:
1160:
1155:
1147:
1142:
1134:
1129:
1121:
1116:
1108:
1105:Observation.org
1103:
1095:
1090:
1082:
1077:
1069:
1064:
1056:
1051:
1043:
1038:
1030:
1025:
1017:
1012:
1004:
999:
991:
986:
978:
973:
965:
960:
951:
950:
945:
932:
913:
887:
884:
883:
873:
871:
866:
865:
861:
849:
845:
827:
826:
819:
809:
807:
797:
796:
792:
785:
767:
766:
762:
752:
750:
732:
731:
727:
717:
715:
701:Papilloculiceps
693:
692:
685:
662:
661:
652:
642:
640:
615:
614:
607:
602:
595:
592:
583:
576:
567:
562:which resemble
560:Platycephalidae
547:
538:
525:
516:
508:ambush predator
504:
491:
458:
434:Papilloculiceps
432:The genus name
430:
422:Scorpaeniformes
382:Papilloculiceps
339:
330:Papilloculiceps
309:Platycephalidae
302:ray-finned fish
257:(Sauvage, 1873)
238:(Rüppell, 1838)
201:
194:
188:
175:
151:
148:Papilloculiceps
137:Platycephalidae
127:Scorpaeniformes
72:
64:
53:
49:
42:
17:
16:Species of fish
12:
11:
5:
1210:
1208:
1200:
1199:
1194:
1189:
1184:
1179:
1174:
1164:
1163:
1157:
1156:
1154:
1153:
1140:
1127:
1114:
1101:
1088:
1075:
1062:
1049:
1036:
1023:
1010:
997:
984:
971:
958:
942:
940:
934:
933:
928:
922:
921:
912:
911:External links
909:
908:
907:
889:Froese, Rainer
882:
881:
859:
843:
817:
790:
783:
760:
725:
683:
664:Froese, Rainer
650:
604:
603:
601:
598:
597:
596:
593:
586:
584:
577:
570:
568:
548:
541:
539:
528:Crocodile fish
526:
519:
515:
512:
503:
500:
490:
487:
457:
454:
429:
426:
394:P. grandidieri
378:Heinz Steinitz
351:Georges Cuvier
338:
335:
270:
269:
268:
267:
259:
251:
240:
232:
221:
210:
209:
203:
202:
195:
184:
183:
177:
176:
169:
167:
163:
162:
144:
140:
139:
134:
130:
129:
124:
120:
119:
117:Actinopterygii
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
66:
65:
47:
44:
43:
38:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
1209:
1198:
1195:
1193:
1190:
1188:
1185:
1183:
1180:
1178:
1175:
1173:
1170:
1169:
1167:
1150:
1145:
1141:
1137:
1132:
1128:
1124:
1119:
1115:
1111:
1106:
1102:
1098:
1093:
1089:
1085:
1080:
1076:
1072:
1067:
1063:
1059:
1054:
1050:
1046:
1041:
1037:
1033:
1028:
1024:
1020:
1015:
1011:
1007:
1002:
998:
994:
989:
985:
981:
976:
972:
968:
963:
959:
954:
948:
944:
943:
941:
939:
935:
931:
926:
919:
915:
914:
910:
904:
903:
898:
896:
890:
886:
885:
870:. Summit Post
869:
863:
860:
857:
853:
847:
844:
839:
835:
831:
824:
822:
818:
805:
801:
794:
791:
786:
780:
776:
775:
770:
764:
761:
749:
745:
744:
739:
735:
729:
726:
714:
710:
709:
704:
702:
696:
690:
688:
684:
679:
678:
673:
671:
665:
659:
657:
655:
651:
638:
633:
629:
625:
624:
619:
612:
610:
606:
599:
590:
585:
581:
574:
569:
565:
561:
557:
556:
551:
545:
540:
537:
533:
529:
523:
518:
513:
511:
509:
501:
499:
496:
495:KwaZulu-Natal
488:
486:
484:
479:
475:
471:
467:
463:
455:
453:
451:
450:specific name
447:
443:
439:
435:
427:
425:
423:
420:
416:
413:
409:
408:
403:
399:
395:
391:
387:
383:
379:
376:
375:ichthyologist
372:
368:
364:
360:
356:
352:
348:
344:
336:
334:
332:
331:
326:
322:
318:
317:Mediterranean
314:
310:
307:
303:
299:
295:
291:
287:
283:
282:
277:
263:
260:
255:
252:
248:
244:
241:
236:
233:
229:
225:
222:
217:
214:
213:
211:
208:
204:
199:
193:
191:
185:
182:
181:Binomial name
178:
174:
173:
168:
165:
164:
159:
155:
150:
149:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
76:
71:
67:
61:
56:
55:Least Concern
45:
41:
36:
32:
27:
22:
19:
937:
900:
894:
872:. Retrieved
862:
851:
846:
837:
833:
808:. Retrieved
803:
793:
773:
769:J. S. Nelson
763:
751:. Retrieved
741:
728:
716:. Retrieved
706:
700:
675:
669:
641:. Retrieved
627:
621:
553:
549:
532:Ras Muhammad
527:
505:
492:
478:total length
466:preoperculum
459:
445:
441:
437:
433:
431:
405:
398:P. longiceps
397:
393:
390:type species
385:
381:
366:
346:
340:
329:
328:
323:through the
293:
289:
285:
280:
279:
275:
273:
265:Fowler, 1935
261:
253:
242:
234:
223:
219:Cuvier, 1829
215:
189:
187:
171:
170:
147:
146:
18:
1027:iNaturalist
840:(1): 57–61.
456:Description
1166:Categories
916:Photos of
600:References
578:Detail of
564:crocodiles
483:caudal fin
470:dorsal fin
402:classified
325:Suez Canal
300:of marine
1084:114173325
582:over eyes
462:operculum
428:Etymology
355:Ehrenberg
343:described
198:G. Cuvier
166:Species:
93:Kingdom:
87:Eukaryota
1045:10150287
1001:FishBase
953:Q2124700
947:Wikidata
902:FishBase
677:FishBase
474:anal fin
412:suborder
337:Taxonomy
315:and the
207:Synonyms
158:Steinitz
133:Family:
107:Chordata
103:Phylum:
97:Animalia
83:Domain:
60:IUCN 3.1
1019:2335818
874:13 July
810:12 July
753:12 July
718:12 July
643:12 July
580:lappets
536:Red Sea
530:at 20m
514:Gallery
502:Biology
438:papilla
417:in the
388:as its
359:Massawa
313:Red Sea
298:species
296:, is a
247:Sauvage
228:Rüppell
200:, 1829)
143:Genus:
123:Order:
113:Class:
58: (
1149:218145
1136:677396
1123:218145
1097:990995
1071:643694
1058:116118
1032:372161
834:Cybium
781:
442:oculus
306:family
249:, 1873
230:, 1838
160:, 1956
156:&
154:Fowler
1144:WoRMS
1110:79491
1040:IRMNG
993:4CKXY
980:10815
419:order
274:The
1118:OBIS
1092:NCBI
1079:IUCN
1066:ITIS
1014:GBIF
1006:7896
975:BOLD
876:2022
812:2022
779:ISBN
755:2022
720:2022
645:2022
628:2018
566:most
552:and
446:ceps
1053:ISC
988:CoL
962:ADW
632:doi
424:.
357:at
345:as
292:or
1168::
1146::
1133::
1120::
1107::
1094::
1081::
1068::
1055::
1042::
1029::
1016::
1003::
990::
977::
964::
949::
899:.
838:14
836:.
832:.
820:^
802:.
746:.
740:.
711:.
705:.
686:^
674:.
653:^
626:.
620:.
608:^
534:,
392:.
333:.
288:,
897:"
893:"
878:.
814:.
787:.
757:.
722:.
703:"
672:"
668:"
647:.
634::
278:(
196:(
62:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.