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Theria

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stage). However, the age estimates of the site are disputed based on the geological complexity and the geographically widespread nature of the Tiaojishan Formations. Further, King and Beck in 2020 argue for an Early Cretaceous age for
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Cornelis, G.; Vernochet, C.; Carradec, Q.; Souquere, S.; Mulot, B.; Catzeflis, F.; Nilsson, M. A.; Menzies, B. R.; Renfree, M. B.; Pierron, G.; Zeller, U.; Heidmann, O.; Dupressoir, A.; Heidmann, T. (2015).
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Molecular data suggests that therians may have originated even earlier, during the Early Jurassic. Therian mammals began to diversify 10-20 million years before the dinosaur extinction.
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The flexible nose in therian mammals is not found in any other vertebrates, and is the product of modified cells involved in the development of the upper jaw in other tetrapods.
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In the above system, Theria is a subclass. Alternatively, in the system proposed by McKenna and Bell (1997) it is ranked as a supercohort under the subclass Theriiformes:
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mammals arose in the Southern Hemisphere during the Early Jurassic, around 50 million years prior to the clade's earliest undisputed appearance in the Northern Hemisphere.
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The rank of "Theria" may vary depending on the classification system used. The textbook classification system by Vaughan et al. (2000) gives the following:
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Luo, Z.-X., Z. Kielan-Jaworowska, and R. L. Cifelli. 2002. In quest for a phylogeny of Mesozoic mammals. Acta Palaeontologica Polonica, 47:1-78.
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The marsupials and the placental mammals evolved from a common therian ancestor that gave live birth by suppressing the mother's
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Another classification proposed by Luo et al. (2002) does not assign any rank to the taxonomic levels, but uses a purely
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Flannery, Timothy F.; Rich, Thomas H.; Vickers-Rich, Patricia; Veatch, E. Grace; Helgen, Kristofer M. (2022-10-02).
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Bi, Shundong; Zheng, Xiaoting; Wang, Xiaoli; Cignetti, Natalie E.; Yang, Shiling; Wible, John R. (June 2018).
487:(external ears) are also a distinctive trait that is a therian exclusivity, though some therians, such as the 4310: 1263: 836:
SUMIYAMA K; MIYAKE T; GRIMWOOD J; STUART A; DICKSON M; SCHMUTZ J; RUDDLE FH; MYERS RM; AMEMIYA CT (2012).
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and various extinct mammals evolving prior to the common ancestor of placentals and marsupials.
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such as in marsupials. Genetic studies have suggested a viral origin of syncytins through the
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Higashiyama, H.; Koyabu, D.; Hirasawa, T.; Werneburg, I.; Kuratani, S.; Kurihara, H. (2021).
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A recent review of the Southern Hemisphere Mesozoic mammal fossil record has argued that
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Proceedings of the National Academy of Sciences of the United States of America
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Proceedings of the National Academy of Sciences of the United States of America
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Myers, P.; R. Espinosa; C. S. Parr; T. Jones; G. S. Hammond & T. A. Dewey.
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Journal of Experimental Zoology Part B: Molecular and Developmental Evolution
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Subclass Theriiformes: live-bearing mammals and their prehistoric relatives
478: 448: 416: 412: 176: 112: 4281:(Lorises, lemurs, tarsiers, monkeys, apes (including humans) and relatives) 1091: 1065: 993: 938: 879: 800: 4327: 4201: 4057: 3885: 3575: 3477: 3267: 3206: 2833: 1961: 1760: 1619: 1382: 1330: 1303: 1033: 617: 571: 504: 474: 444: 400: 258: 196: 102: 97: 82: 77: 67: 31: 861: 17: 4278: 4155: 3987: 3897: 3834: 3749: 3370: 3166: 2486: 2001: 1782: 1708: 815:"Ancient "genomic parasites" spurred evolution of pregnancy in mammals" 117: 92: 4388: 4270: 4246: 4094: 3879: 3866: 3447: 1450: 1409: 1297: 1279: 1173:"Mammals began their takeover long before the death of the dinosaurs" 954:"An Early Cretaceous eutherian and the placental-marsupial dichotomy" 593: 550: 443:
which allow exchanges between the mother and its offspring through a
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Vaughan, Terry A., James M. Ryan, and Nicholas J. Czaplewski. 2000.
462:. While the marsupials continued to give birth to an underdeveloped 4139: 463: 4375: 30:"Therians" redirects here. For the contemporary subculture, see 4308: 3838: 3672: 2763: 1919: 1448: 1290: 1252: 4193: 607:: modern live-bearing mammals and their prehistoric relatives 352: 326: 349: 323: 128: 1248: 358: 355: 332: 329: 1054:
Proceedings of the Royal Society B: Biological Sciences
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Alcheringa: An Australasian Journal of Palaeontology
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(2020-06-10). 132: 1217:McKenna, Malcolm C., and Bell, Susan K. 1997. 3850: 1264: 517:, in line with similar early mammaliaformes. 27:Subclass of mammals in the clade Theriiformes 8: 502:The earliest known therian mammal fossil is 895:"Mammalian face as an evolutionary novelty" 4305: 4173: 4074: 3967: 3946: 3912: 3857: 3843: 3835: 3680: 3669: 3553: 3396: 3195: 2990: 2789: 2777: 2773: 2760: 2516: 1940: 1929: 1916: 1697: 1587: 1468: 1458: 1445: 1387: 1360: 1351: 1287: 1271: 1257: 1249: 755:exaptation for placentation in marsupials" 48: 1130: 1081: 1032: 928: 918: 869: 790: 780: 770: 4474:Extant Middle Jurassic first appearances 4188:(Hedgehogs, shrews, moles and relatives) 4484:Taxa named by William Aitcheson Haswell 1201:. Saunders College Publishing, 565 pp. 751:"Retroviral envelope gene captures and 710: 7: 32:Otherkin § Terms and identities 1179:from the original on 8 January 2022 1175:. Field Museum of Natural History. 473:Therian mammals no longer have the 1171:Golembiewski, Kate (2 June 2016). 825:from the original on Jan 14, 2024. 25: 718:ITIS Standard Report Page: Theria 3655: 3649: 2746: 2740: 1901: 1894: 435:to live young without a shelled 342: 316: 163: 508:, from China's Late Jurassic ( 491:, have lost them secondarily. 415:) but excludes the egg-laying 1: 1132:10.1080/03115518.2022.2132288 477:bone, unlike their cousins, 428: 427:Therian mammals give birth 4500: 4257:(Rabbits, hares and pikas) 4212:(Dogs, cats and relatives) 4118:(Tenrecs and golden moles) 534: 373: 36: 29: 4043:(Kangaroos and relatives) 3874: 3679: 3668: 3646: 3556: 3399: 3198: 2993: 2792: 2776: 2772: 2759: 2737: 2519: 1943: 1928: 1915: 1891: 1700: 1471: 1457: 1444: 1390: 1363: 1350: 1286: 1199:Mammalogy: Fourth Edition 978:10.1038/s41586-018-0210-3 387: 'wild beast') is a 251: 246: 160:Scientific classification 158: 127: 51: 4035:(Bilbies and bandicoots) 37:For the moth genus, see 3927:(Platypus and echidnas) 1021:National Science Review 920:10.1073/pnas.2111876118 782:10.1073/pnas.1417000112 560:: live-bearing mammals 58:Early Cretaceous–Recent 4479:Taxa described in 1897 4158:(Dugongs and manatees) 4097:(Anteaters and sloths) 1066:10.1098/rspb.2020.0943 399:. Theria includes the 153: 4423:Paleobiology Database 4027:(Quolls and dunnarts) 537:Mammal classification 152: 3745:Saint Bathans Mammal 821:. January 29, 2015. 735:Animal Diversity Web 1123:2022Alch...46..277F 970:2018Natur.558..390B 911:2021PNAS..11811876H 905:(44): e2111876118. 862:10.1002/jez.b.22469 854:2012JEZB..318..639S 772:2015PNAS..112E.487C 4011:(Monito del monte) 3190:Prototribosphenida 1693:Ornithorhynchoidea 1543:Ausktribosphenidae 1060:(1928): 20200943. 1034:10.1093/nsr/nwu070 1015:Meng, Jin (2014). 699:Patagomaia chainko 631:: therian mammals 629:Supercohort Theria 154: 4451: 4450: 4311:Taxon identifiers 4302: 4301: 4298: 4297: 4294: 4293: 4290: 4289: 4282: 4274: 4266: 4258: 4250: 4229: 4221: 4213: 4205: 4197: 4189: 4159: 4151: 4143: 4135: 4127: 4126:(Elephant shrews) 4119: 4098: 4090: 4071: 4052: 4051: 4044: 4036: 4028: 4020: 4019:(Marsupial moles) 4017:Notoryctemorphia 4012: 3991: 3983: 3980:Paucituberculata 3964: 3936: 3935: 3928: 3832: 3831: 3828: 3827: 3824: 3823: 3664: 3663: 3644: 3643: 3640: 3639: 3636: 3635: 3632: 3631: 3183: 3182: 3135: 3134: 2977: 2976: 2755: 2754: 2735: 2734: 2731: 2730: 2727: 2726: 2684:Symmetrodontoides 2540:Zhangheotheriidae 1911: 1910: 1889: 1888: 1859:multituberculates 1832: 1831: 1828: 1827: 1720:Ornithorhynchidae 1577: 1576: 1440: 1439: 1436: 1435: 1432: 1431: 1428: 1427: 964:(7710): 390–395. 819:UChicago Medicine 731:"Subclass Theria" 694:Placental mammals 663: 662: 582: 581: 515:Juramaia sinensis 405:placental mammals 309: 308: 242: 16:(Redirected from 4491: 4444: 4443: 4431: 4430: 4418: 4417: 4405: 4404: 4392: 4391: 4379: 4378: 4366: 4365: 4353: 4352: 4351: 4338: 4337: 4336: 4306: 4280: 4272: 4264: 4256: 4248: 4239:Euarchontoglires 4227: 4219: 4211: 4203: 4195: 4187: 4174: 4157: 4149: 4141: 4133: 4125: 4117: 4096: 4088: 4075: 4070: 4069: 4061: 4042: 4034: 4033:Peramelemorphia 4026: 4018: 4010: 3989: 3988:Didelphimorphia 3982:(Shrew opossums) 3981: 3968: 3963: 3962: 3954: 3947: 3926: 3917:Australosphenida 3913: 3859: 3852: 3845: 3836: 3736: 3726: 3716: 3706: 3696: 3681: 3670: 3659: 3654: 3653: 3562: 3554: 3535: 3525: 3515: 3505: 3495: 3485: 3475: 3465: 3455: 3445: 3435: 3425: 3415: 3405: 3397: 3378:Tendagurutherium 3338: 3315: 3305: 3295: 3285: 3275: 3265: 3255: 3244: 3234: 3224: 3214: 3204: 3196: 3142: 3006:Austrotriconodon 2991: 2984: 2961:Portopinheirodon 2813:Drescheratherium 2790: 2783: 2778: 2774: 2761: 2750: 2745: 2744: 2677:Spalacotheroides 2663:Spalacotheridium 2598:Spalacotheriidae 2596: 2538: 2517: 2468: 2423: 2322: 2291: 2261: 2197:Gobiconodontidae 2195: 2171: 2161: 2151: 2141: 2131: 2121: 2111: 2101: 2091: 2081: 2071: 2061: 2051: 2041: 2030: 2019: 2009: 1999: 1989: 1979: 1969: 1959: 1949: 1941: 1930: 1917: 1906: 1905: 1899: 1898: 1839: 1802: 1792: 1768: 1751: 1740: 1729: 1706: 1698: 1663: 1632: 1593: 1588: 1552:Ausktribosphenos 1541: 1511: 1490:Tendagurutherium 1487: 1477: 1469: 1463:Australosphenida 1459: 1446: 1388: 1361: 1352: 1345: 1344: 1288: 1273: 1266: 1259: 1250: 1238: 1235: 1229: 1215: 1209: 1195: 1189: 1188: 1186: 1184: 1168: 1162: 1151: 1145: 1144: 1134: 1117:(3–4): 277–290. 1102: 1096: 1095: 1085: 1045: 1039: 1038: 1036: 1012: 1006: 1005: 949: 943: 942: 932: 922: 890: 884: 883: 873: 833: 827: 826: 811: 805: 804: 794: 784: 774: 765:(5): E487–E496. 745: 739: 738: 726: 720: 715: 669:system instead. 587: 586: 544: 543: 434: 384: 377: 365: 364: 361: 360: 357: 354: 351: 348: 339: 338: 335: 334: 331: 328: 325: 322: 233: 168: 167: 131: 122: 59: 55:Temporal range: 49: 21: 4499: 4498: 4494: 4493: 4492: 4490: 4489: 4488: 4469:Mammal taxonomy 4454: 4453: 4452: 4447: 4439: 4434: 4426: 4421: 4413: 4408: 4400: 4395: 4387: 4382: 4374: 4369: 4361: 4356: 4347: 4346: 4341: 4332: 4331: 4326: 4313: 4303: 4286: 4233: 4218:Perissodactyla 4163: 4102: 4063: 4062: 4060: 4048: 4025:Dasyuromorphia 4009:Microbiotheria 4001:Australidelphia 3995: 3956: 3955: 3953: 3932: 3902: 3870: 3863: 3833: 3820: 3754: 3675: 3660: 3648: 3628: 3599: 3570: 3543: 3391: 3390:Tribosphenida / 3384: 3331: 3192: 3179: 3131: 3089:Mesungulatoidea 3083: 2973: 2846: 2768: 2751: 2739: 2723: 2703: 2589: 2584:Zhangheotherium 2531: 2506: 2494:Trishulotherium 2461: 2456:Volaticotherium 2416: 2403:Victoriaconodon 2324:Triconodontidae 2315: 2284: 2254: 2188: 2144:Phascolotherium 1937: 1924: 1907: 1900: 1893: 1885: 1865: 1824: 1777: 1761:Ornithorhynchus 1714: 1687: 1656: 1634:Steropodontidae 1625: 1595:Kollikodontidae 1573: 1534: 1504: 1453: 1424: 1404: 1377: 1346: 1293: 1292: 1282: 1277: 1247: 1242: 1241: 1236: 1232: 1216: 1212: 1196: 1192: 1182: 1180: 1170: 1169: 1165: 1152: 1148: 1104: 1103: 1099: 1047: 1046: 1042: 1014: 1013: 1009: 951: 950: 946: 892: 891: 887: 835: 834: 830: 813: 812: 808: 747: 746: 742: 728: 727: 723: 716: 712: 707: 675: 558:Subclass Theria 539: 533: 500: 449:rudimental ones 425: 423:Characteristics 411:(including the 403:(including the 345: 341: 319: 315: 232: 162: 151: 129: 123: 121: 120: 115: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 57: 56: 53: 45: 35: 28: 23: 22: 15: 12: 11: 5: 4497: 4495: 4487: 4486: 4481: 4476: 4471: 4466: 4456: 4455: 4449: 4448: 4446: 4445: 4432: 4419: 4406: 4393: 4380: 4367: 4354: 4339: 4323: 4321: 4315: 4314: 4309: 4300: 4299: 4296: 4295: 4292: 4291: 4288: 4287: 4285: 4284: 4276: 4268: 4260: 4252: 4243: 4241: 4235: 4234: 4232: 4231: 4223: 4215: 4207: 4199: 4191: 4182: 4180: 4178:Laurasiatheria 4171: 4165: 4164: 4162: 4161: 4153: 4145: 4137: 4132:Tubulidentata 4129: 4124:Macroscelidea 4121: 4112: 4110: 4104: 4103: 4101: 4100: 4092: 4083: 4081: 4072: 4054: 4053: 4050: 4049: 4047: 4046: 4041:Diprotodontia 4038: 4030: 4022: 4014: 4005: 4003: 3997: 3996: 3994: 3993: 3985: 3976: 3974: 3965: 3944: 3938: 3937: 3934: 3933: 3931: 3930: 3921: 3919: 3910: 3904: 3903: 3901: 3900: 3894: 3888: 3882: 3875: 3872: 3871: 3864: 3862: 3861: 3854: 3847: 3839: 3830: 3829: 3826: 3825: 3822: 3821: 3819: 3818: 3813: 3811:Amphitheriidae 3808: 3803: 3798: 3793: 3788: 3783: 3778: 3776:Eutriconodonta 3773: 3768: 3762: 3760: 3756: 3755: 3753: 3752: 3747: 3742: 3732: 3722: 3712: 3702: 3691: 3689: 3686:Incertae sedis 3677: 3676: 3673: 3666: 3665: 3662: 3661: 3647: 3645: 3642: 3641: 3638: 3637: 3634: 3633: 3630: 3629: 3627: 3626: 3625: 3624: 3618: 3609: 3607: 3601: 3600: 3598: 3597: 3596: 3595: 3589: 3580: 3578: 3572: 3571: 3569: 3568: 3557: 3551: 3545: 3544: 3542: 3541: 3531: 3521: 3511: 3501: 3491: 3481: 3471: 3461: 3451: 3441: 3431: 3421: 3418:Argaliatherium 3411: 3400: 3394: 3386: 3385: 3383: 3382: 3374: 3367: 3360: 3353: 3344: 3342: 3333: 3332: 3330: 3329: 3321: 3311: 3301: 3291: 3281: 3271: 3261: 3251: 3247:Brancatherulum 3240: 3230: 3220: 3217:Amphibetulimus 3210: 3199: 3193: 3188: 3185: 3184: 3181: 3180: 3178: 3177: 3170: 3163: 3156: 3148: 3146: 3137: 3136: 3133: 3132: 3130: 3129: 3122: 3119:Peligrotherium 3115: 3108: 3101: 3098:Coloniatherium 3093: 3091: 3085: 3084: 3082: 3081: 3074: 3071:Quirogatherium 3067: 3060: 3053: 3046: 3038: 3034:Groebertherium 3030: 3023: 3016: 3009: 3002: 2994: 2988: 2986:Meridiolestida 2979: 2978: 2975: 2974: 2972: 2971: 2964: 2957: 2950: 2943: 2936: 2928: 2921: 2914: 2907: 2903:Groebertherium 2899: 2892: 2885: 2878: 2875:Anthracolestes 2871: 2864: 2856: 2854: 2848: 2847: 2845: 2844: 2837: 2830: 2827:Henkelotherium 2823: 2816: 2809: 2802: 2798:Brancatherulum 2793: 2787: 2770: 2769: 2764: 2757: 2756: 2753: 2752: 2738: 2736: 2733: 2732: 2729: 2728: 2725: 2724: 2722: 2721: 2720: 2719: 2713: 2711: 2705: 2704: 2702: 2701: 2694: 2691:Symmetrolestes 2687: 2680: 2673: 2670:Spalacotherium 2666: 2659: 2652: 2645: 2638: 2631: 2624: 2621:Cifellitherium 2617: 2610: 2602: 2600: 2591: 2590: 2588: 2587: 2580: 2573: 2566: 2559: 2552: 2544: 2542: 2533: 2532: 2530: 2529: 2520: 2514: 2508: 2507: 2505: 2504: 2497: 2490: 2483: 2479:Gobiotheriodon 2474: 2472: 2463: 2462: 2460: 2459: 2452: 2444: 2441:Ichthyoconodon 2437: 2434:Argentoconodon 2429: 2427: 2425:Volaticotheria 2418: 2417: 2415: 2414: 2410:Volaticotheria 2406: 2399: 2392: 2385: 2378: 2371: 2364: 2357: 2350: 2343: 2340:Arundelconodon 2336: 2328: 2326: 2317: 2316: 2314: 2313: 2306: 2298: 2296: 2286: 2285: 2283: 2282: 2275: 2267: 2265: 2263:Jeholodentidae 2256: 2255: 2253: 2252: 2245: 2238: 2231: 2223: 2216: 2209: 2201: 2199: 2190: 2189: 2187: 2186: 2178: 2167: 2157: 2154:Sangarotherium 2147: 2137: 2127: 2117: 2107: 2097: 2087: 2077: 2067: 2057: 2054:Indotriconodon 2047: 2037: 2026: 2015: 2005: 1995: 1985: 1975: 1965: 1955: 1944: 1938: 1933: 1926: 1925: 1920: 1913: 1912: 1909: 1908: 1892: 1890: 1887: 1886: 1884: 1883: 1882: 1881: 1875: 1873: 1867: 1866: 1864: 1863: 1862: 1861: 1855: 1846: 1844: 1834: 1833: 1830: 1829: 1826: 1825: 1823: 1822: 1815: 1808: 1798: 1787: 1785: 1783:Tachyglossidae 1779: 1778: 1776: 1775: 1771:Patagorhynchus 1764: 1757: 1747: 1736: 1724: 1722: 1716: 1715: 1713: 1712: 1701: 1695: 1689: 1688: 1686: 1685: 1678: 1669: 1667: 1658: 1657: 1655: 1654: 1647: 1638: 1636: 1627: 1626: 1624: 1623: 1616: 1608: 1600: 1598: 1585: 1579: 1578: 1575: 1574: 1572: 1571: 1563: 1555: 1547: 1545: 1536: 1535: 1533: 1532: 1525: 1517: 1515: 1506: 1505: 1503: 1502: 1494: 1483: 1472: 1466: 1455: 1454: 1449: 1442: 1441: 1438: 1437: 1434: 1433: 1430: 1429: 1426: 1425: 1423: 1422: 1421: 1420: 1414: 1412: 1406: 1405: 1403: 1402: 1401: 1400: 1391: 1385: 1379: 1378: 1376: 1375: 1374: 1373: 1364: 1358: 1348: 1347: 1343: 1342: 1340:Mammaliaformes 1333: 1324: 1315: 1306: 1300: 1291: 1284: 1283: 1278: 1276: 1275: 1268: 1261: 1253: 1246: 1245:External links 1243: 1240: 1239: 1230: 1210: 1190: 1163: 1146: 1097: 1040: 1027:(4): 521–542. 1007: 944: 885: 848:(8): 639–650. 828: 806: 740: 721: 709: 708: 706: 703: 702: 701: 696: 691: 686: 681: 674: 671: 661: 660: 659: 658: 657: 656: 655: 654: 653: 652: 651: 650: 649: 648: 647: 646: 639: 580: 579: 578: 577: 576: 575: 568: 535:Main article: 532: 529: 499: 496: 466:after a short 453:endogenization 424: 421: 307: 306: 305: 304: 303: 302: 290:Incertae sedis 286: 283:Spelaeomolitor 278: 270: 261: 249: 248: 244: 243: 227: 223: 222: 217: 210: 209: 204: 200: 199: 194: 190: 189: 184: 180: 179: 174: 170: 169: 156: 155: 125: 124: 116: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 60: 54: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 4496: 4485: 4482: 4480: 4477: 4475: 4472: 4470: 4467: 4465: 4462: 4461: 4459: 4442: 4437: 4433: 4429: 4424: 4420: 4416: 4411: 4407: 4403: 4398: 4394: 4390: 4385: 4381: 4377: 4372: 4368: 4364: 4359: 4355: 4350: 4344: 4340: 4335: 4329: 4325: 4324: 4322: 4320: 4316: 4312: 4307: 4283: 4277: 4275: 4269: 4267: 4261: 4259: 4253: 4251: 4245: 4244: 4242: 4240: 4236: 4230: 4226:Artiodactyla 4224: 4222: 4216: 4214: 4208: 4206: 4200: 4198: 4192: 4190: 4186:Eulipotyphla 4184: 4183: 4181: 4179: 4175: 4172: 4170: 4169:Boreoeutheria 4166: 4160: 4154: 4152: 4146: 4144: 4138: 4136: 4130: 4128: 4122: 4120: 4116:Afrosoricida 4114: 4113: 4111: 4109: 4105: 4099: 4093: 4091: 4085: 4084: 4082: 4080: 4076: 4073: 4067: 4059: 4055: 4045: 4039: 4037: 4031: 4029: 4023: 4021: 4015: 4013: 4007: 4006: 4004: 4002: 3998: 3992: 3986: 3984: 3978: 3977: 3975: 3973: 3969: 3966: 3960: 3952: 3948: 3945: 3943: 3939: 3929: 3923: 3922: 3920: 3918: 3914: 3911: 3909: 3905: 3899: 3895: 3893: 3889: 3887: 3883: 3881: 3877: 3876: 3873: 3868: 3860: 3855: 3853: 3848: 3846: 3841: 3840: 3837: 3817: 3814: 3812: 3809: 3807: 3806:Paurodontidae 3804: 3802: 3801:Eupantotheria 3799: 3797: 3796:Symmetrodonta 3794: 3792: 3789: 3787: 3784: 3782: 3781:Amphilestidae 3779: 3777: 3774: 3772: 3769: 3767: 3764: 3763: 3761: 3757: 3751: 3748: 3746: 3743: 3741: 3740: 3733: 3731: 3730: 3723: 3721: 3720: 3713: 3711: 3710: 3709:Chronoperates 3703: 3701: 3700: 3693: 3692: 3690: 3688: 3687: 3682: 3678: 3671: 3667: 3658: 3652: 3623: 3619: 3617: 3613: 3612: 3611: 3610: 3608: 3606: 3602: 3594: 3590: 3588: 3584: 3583: 3582: 3581: 3579: 3577: 3573: 3567: 3566: 3559: 3558: 3555: 3552: 3550: 3546: 3540: 3539: 3538:Tribactonodon 3532: 3530: 3529: 3522: 3520: 3519: 3512: 3510: 3509: 3502: 3500: 3499: 3498:Kielantherium 3492: 3490: 3489: 3482: 3480: 3479: 3472: 3470: 3469: 3462: 3460: 3459: 3458:Holoclemensia 3452: 3450: 3449: 3442: 3440: 3439: 3432: 3430: 3429: 3422: 3420: 3419: 3412: 3410: 3409: 3402: 3401: 3398: 3395: 3393: 3392:Boreosphenida 3387: 3380: 3379: 3375: 3373: 3372: 3368: 3366: 3365: 3361: 3359: 3358: 3354: 3351: 3350: 3346: 3345: 3343: 3341: 3334: 3327: 3326: 3322: 3320: 3319: 3312: 3310: 3309: 3308:Palaeoxonodon 3302: 3300: 3299: 3292: 3290: 3289: 3282: 3280: 3279: 3272: 3270: 3269: 3262: 3260: 3259: 3252: 3249: 3248: 3241: 3239: 3238: 3231: 3229: 3228: 3221: 3219: 3218: 3211: 3209: 3208: 3201: 3200: 3197: 3194: 3191: 3186: 3176: 3175: 3171: 3169: 3168: 3164: 3162: 3161: 3157: 3155: 3154: 3150: 3149: 3147: 3145: 3138: 3128: 3127: 3123: 3121: 3120: 3116: 3114: 3113: 3109: 3107: 3106: 3102: 3100: 3099: 3095: 3094: 3092: 3090: 3086: 3080: 3079: 3078:Solanutherium 3075: 3073: 3072: 3068: 3066: 3065: 3064:Paraungulatum 3061: 3059: 3058: 3054: 3052: 3051: 3047: 3044: 3043: 3039: 3036: 3035: 3031: 3029: 3028: 3024: 3022: 3021: 3017: 3015: 3014: 3010: 3008: 3007: 3003: 3001: 3000: 2996: 2995: 2992: 2989: 2987: 2980: 2970: 2969: 2968:Treslagosodon 2965: 2963: 2962: 2958: 2956: 2955: 2951: 2949: 2948: 2944: 2942: 2941: 2937: 2934: 2933: 2929: 2927: 2926: 2925:Krebsotherium 2922: 2920: 2919: 2915: 2913: 2912: 2908: 2905: 2904: 2900: 2898: 2897: 2893: 2891: 2890: 2886: 2884: 2883: 2879: 2877: 2876: 2872: 2870: 2869: 2865: 2863: 2862: 2858: 2857: 2855: 2853: 2849: 2843: 2842: 2838: 2836: 2835: 2831: 2829: 2828: 2824: 2822: 2821: 2817: 2815: 2814: 2810: 2808: 2807: 2803: 2800: 2799: 2795: 2794: 2791: 2788: 2786: 2779: 2775: 2771: 2767: 2762: 2758: 2749: 2743: 2717: 2716: 2715: 2714: 2712: 2710: 2706: 2700: 2699: 2695: 2693: 2692: 2688: 2686: 2685: 2681: 2679: 2678: 2674: 2672: 2671: 2667: 2665: 2664: 2660: 2658: 2657: 2656:Spalacolestes 2653: 2651: 2650: 2646: 2644: 2643: 2639: 2637: 2636: 2635:Infernolestes 2632: 2630: 2629: 2628:Heishanlestes 2625: 2623: 2622: 2618: 2616: 2615: 2611: 2609: 2608: 2604: 2603: 2601: 2599: 2592: 2586: 2585: 2581: 2579: 2578: 2574: 2572: 2571: 2570:Ningchengodon 2567: 2565: 2564: 2560: 2558: 2557: 2553: 2551: 2550: 2546: 2545: 2543: 2541: 2534: 2527: 2526: 2522: 2521: 2518: 2515: 2513: 2512:Trechnotheria 2509: 2503: 2502: 2498: 2496: 2495: 2491: 2489: 2488: 2484: 2481: 2480: 2476: 2475: 2473: 2471: 2464: 2458: 2457: 2453: 2450: 2449: 2448:Triconolestes 2445: 2443: 2442: 2438: 2436: 2435: 2431: 2430: 2428: 2426: 2419: 2412: 2411: 2407: 2405: 2404: 2400: 2398: 2397: 2393: 2391: 2390: 2386: 2384: 2383: 2379: 2377: 2376: 2372: 2370: 2369: 2365: 2363: 2362: 2358: 2356: 2355: 2351: 2349: 2348: 2344: 2342: 2341: 2337: 2335: 2334: 2330: 2329: 2327: 2325: 2318: 2312: 2311: 2307: 2305: 2304: 2300: 2299: 2297: 2294: 2287: 2281: 2280: 2276: 2274: 2273: 2269: 2268: 2266: 2264: 2257: 2251: 2250: 2246: 2244: 2243: 2239: 2237: 2236: 2232: 2229: 2228: 2227:Huasteconodon 2224: 2222: 2221: 2217: 2215: 2214: 2210: 2208: 2207: 2206:Fuxinoconodon 2203: 2202: 2200: 2198: 2191: 2184: 2183: 2179: 2176: 2175: 2168: 2166: 2165: 2158: 2156: 2155: 2148: 2146: 2145: 2138: 2136: 2135: 2128: 2126: 2125: 2118: 2116: 2115: 2108: 2106: 2105: 2098: 2096: 2095: 2094:Kryptotherium 2088: 2086: 2085: 2078: 2076: 2075: 2068: 2066: 2065: 2058: 2056: 2055: 2048: 2046: 2045: 2038: 2035: 2034: 2027: 2024: 2023: 2016: 2014: 2013: 2006: 2004: 2003: 1996: 1994: 1993: 1992:Chaoyangodens 1986: 1984: 1983: 1976: 1974: 1973: 1966: 1964: 1963: 1956: 1954: 1953: 1946: 1945: 1942: 1939: 1936: 1931: 1927: 1923: 1918: 1914: 1904: 1897: 1879: 1878: 1877: 1876: 1874: 1872: 1868: 1860: 1856: 1854: 1850: 1849: 1848: 1847: 1845: 1842: 1835: 1821: 1820: 1816: 1814: 1813: 1809: 1807: 1806: 1805:Murrayglossus 1799: 1797: 1796: 1795:Megalibgwilia 1789: 1788: 1786: 1784: 1780: 1773: 1772: 1765: 1763: 1762: 1758: 1756: 1755: 1748: 1745: 1744: 1737: 1734: 1733: 1726: 1725: 1723: 1721: 1717: 1711: 1710: 1703: 1702: 1699: 1696: 1694: 1690: 1684: 1683: 1679: 1676: 1675: 1671: 1670: 1668: 1666: 1665:Teinolophidae 1659: 1653: 1652: 1648: 1645: 1644: 1640: 1639: 1637: 1635: 1628: 1622: 1621: 1617: 1614: 1613: 1609: 1607: 1606: 1602: 1601: 1599: 1596: 1589: 1586: 1584: 1580: 1569: 1568: 1564: 1561: 1560: 1556: 1554: 1553: 1549: 1548: 1546: 1544: 1537: 1531: 1530: 1526: 1524: 1523: 1519: 1518: 1516: 1514: 1507: 1500: 1499: 1495: 1492: 1491: 1484: 1482: 1481: 1474: 1473: 1470: 1467: 1464: 1460: 1456: 1452: 1447: 1443: 1418: 1417: 1416: 1415: 1413: 1411: 1407: 1399: 1395: 1394: 1393: 1392: 1389: 1386: 1384: 1380: 1372: 1368: 1367: 1366: 1365: 1362: 1359: 1357: 1353: 1349: 1341: 1337: 1334: 1332: 1328: 1325: 1323: 1319: 1316: 1314: 1310: 1307: 1305: 1301: 1299: 1295: 1294: 1289: 1285: 1281: 1274: 1269: 1267: 1262: 1260: 1255: 1254: 1251: 1244: 1234: 1231: 1228: 1227:0-231-11013-8 1224: 1220: 1214: 1211: 1208: 1207:0-03-025034-X 1204: 1200: 1194: 1191: 1178: 1174: 1167: 1164: 1161:56(4):543-63. 1160: 1156: 1153:Hugall, A.F. 1150: 1147: 1142: 1138: 1133: 1128: 1124: 1120: 1116: 1112: 1108: 1101: 1098: 1093: 1089: 1084: 1079: 1075: 1071: 1067: 1063: 1059: 1055: 1051: 1044: 1041: 1035: 1030: 1026: 1022: 1018: 1011: 1008: 1003: 999: 995: 991: 987: 983: 979: 975: 971: 967: 963: 959: 955: 948: 945: 940: 936: 931: 926: 921: 916: 912: 908: 904: 900: 896: 889: 886: 881: 877: 872: 867: 863: 859: 855: 851: 847: 843: 839: 832: 829: 824: 820: 816: 810: 807: 802: 798: 793: 788: 783: 778: 773: 768: 764: 760: 756: 754: 744: 741: 736: 732: 725: 722: 719: 714: 711: 704: 700: 697: 695: 692: 690: 687: 685: 682: 680: 677: 676: 672: 670: 668: 644: 640: 637: 633: 632: 630: 627: 626: 625: 624:Tribosphenida 621: 620: 619: 615: 614: 613: 609: 608: 606: 602: 601: 599: 598: 597: 596: 595: 589: 588: 585: 573: 569: 566: 562: 561: 559: 556: 555: 554: 553: 552: 546: 545: 542: 538: 530: 528: 525: 523: 518: 516: 511: 507: 506: 497: 495: 492: 490: 489:earless seals 486: 482: 480: 476: 471: 469: 465: 461: 460:immune system 456: 454: 450: 446: 442: 438: 432: 422: 420: 418: 414: 410: 406: 402: 398: 394: 390: 386: 383: 376: 372: 369: 368:Ancient Greek 363: 337: 313: 301: 300: 295: 294: 292: 291: 287: 285: 284: 279: 277: 276: 271: 269: 265: 262: 260: 256: 253: 252: 250: 245: 240: 236: 231: 228: 225: 224: 221: 220:Tribosphenida 218: 215: 212: 211: 208: 205: 202: 201: 198: 195: 192: 191: 188: 185: 182: 181: 178: 175: 172: 171: 166: 161: 157: 126: 119: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 50: 47: 43: 41: 33: 19: 4318: 4265:(Treeshrews) 4148:Proboscidea 4089:(Armadillos) 3972:Ameridelphia 3941: 3925:Monotremata 3786:Theriiformes 3737: 3727: 3717: 3707: 3697: 3684: 3563: 3548: 3536: 3526: 3516: 3508:Pappotherium 3506: 3496: 3486: 3476: 3466: 3456: 3446: 3438:Carinalestes 3436: 3426: 3416: 3406: 3376: 3369: 3364:Peramuroides 3362: 3355: 3347: 3323: 3316: 3306: 3296: 3286: 3276: 3266: 3256: 3245: 3235: 3227:Amphitherium 3225: 3215: 3205: 3172: 3165: 3158: 3151: 3144:Donodontidae 3126:Reigitherium 3124: 3117: 3110: 3105:Mesungulatum 3103: 3096: 3076: 3069: 3062: 3055: 3048: 3042:Lakotalestes 3040: 3032: 3025: 3020:Casamiquelia 3018: 3011: 3004: 2997: 2966: 2959: 2954:Phascolestes 2952: 2947:Minutolestes 2945: 2938: 2932:Lakotalestes 2930: 2923: 2916: 2911:Guimarotodus 2909: 2901: 2894: 2887: 2880: 2873: 2868:Amblotherium 2866: 2859: 2852:Dryolestidae 2839: 2832: 2825: 2818: 2811: 2804: 2796: 2696: 2689: 2682: 2675: 2668: 2661: 2654: 2647: 2640: 2633: 2626: 2619: 2612: 2605: 2582: 2575: 2568: 2561: 2554: 2547: 2523: 2499: 2492: 2485: 2477: 2470:Tinodontidae 2454: 2446: 2439: 2432: 2408: 2401: 2394: 2387: 2380: 2373: 2366: 2361:Eotriconodon 2359: 2354:Corviconodon 2352: 2347:Astroconodon 2345: 2338: 2331: 2308: 2301: 2277: 2270: 2247: 2240: 2233: 2225: 2218: 2211: 2204: 2180: 2172: 2164:Tendagurodon 2162: 2152: 2142: 2132: 2122: 2112: 2102: 2092: 2082: 2072: 2062: 2052: 2042: 2033:Fruitafossor 2031: 2020: 2010: 2000: 1990: 1980: 1970: 1960: 1950: 1935:Theriimorpha 1922:Theriimorpha 1871:Theriimorpha 1817: 1812:Tachyglossus 1810: 1803: 1793: 1769: 1759: 1752: 1743:Monotrematum 1741: 1730: 1707: 1680: 1672: 1649: 1641: 1618: 1610: 1603: 1565: 1557: 1550: 1527: 1522:Asfaltomylos 1520: 1513:Henosferidae 1496: 1488: 1478: 1335: 1326: 1317: 1308: 1233: 1218: 1213: 1198: 1193: 1181:. Retrieved 1166: 1158: 1154: 1149: 1114: 1110: 1100: 1057: 1053: 1043: 1024: 1020: 1010: 961: 957: 947: 902: 898: 888: 845: 841: 831: 818: 809: 762: 758: 752: 743: 734: 724: 713: 664: 645:: placentals 638:: marsupials 628: 622:Infralegion 591: 590: 583: 574:: placentals 567:: marsupials 557: 548: 547: 540: 526: 519: 514: 503: 501: 493: 483: 472: 457: 426: 409:metatherians 397:Theriiformes 395:amongst the 381: 378: 371: 311: 310: 297: 288: 281: 273: 263: 254: 229: 213: 46: 39: 4384:iNaturalist 4343:Wikispecies 4271:Dermoptera 4263:Scandentia 4255:Lagomorpha 4204:(Pangolins) 4194:Chiroptera 4150:(Elephants) 4140:Hyracoidea 4134:(Aardvarks) 3896:(unranked) 3816:Marsupionta 3791:Pantotheria 3766:Prototheria 3719:Microderson 3528:Tirotherium 3428:Brasilestes 3357:Kouriogenys 3349:Afriquiamus 3340:Peramuridae 3325:Monotremata 3318:Vincelestes 3258:Chunnelodon 3160:Anoualestes 3153:Amazighodon 3112:Orretherium 3057:Necrolestes 2999:Amarillodon 2889:Crusafontia 2785:Dryolestida 2766:Cladotheria 2709:Cladotheria 2698:Yaverlestes 2607:Akidolestes 2577:Origolestes 2556:Kiyatherium 2525:Monotremata 2396:Trioracodon 2333:Alticonodon 2293:Klameliidae 2279:Yanoconodon 2249:Spinolestes 2242:Repenomamus 2235:Meemannodon 2213:Gobiconodon 2182:Monotremata 2124:Manchurodon 2114:Liaotherium 2104:Liaoconodon 2064:Juchilestes 2044:Hakusanodon 2022:Dyskritodon 1982:Aploconodon 1972:Amphilestes 1732:Dharragarra 1682:Teinolophos 1583:Monotremata 1498:Monotremata 684:Marsupionta 643:Placentalia 636:Marsupialia 612:Cladotheria 603:Infraclass 570:Infraclass 563:Infraclass 522:triosphenic 366:; from 4458:Categories 4210:Carnivora 4202:Pholidota 4108:Afrotheria 4087:Cingulata 4068:inclusive) 3990:(Opossums) 3961:inclusive) 3951:Metatheria 3908:Yinotheria 3892:Vertebrata 3890:Subphylum 3771:Yinotheria 3759:Other taxa 3739:Tingamarra 3729:Thereuodon 3674:Other taxa 3622:marsupials 3616:Metatheria 3605:Metatheria 3593:placentals 3565:Patagomaia 3408:Aegialodon 3298:Nanolestes 2918:Hercynodon 2896:Dryolestes 2820:Euthlastus 2806:Dorsetodon 2718:see below↓ 2649:Shalbaatar 2563:Maotherium 2389:Triconodon 2375:Meiconodon 2303:Ferganodon 2272:Jeholodens 2174:Allotheria 2074:Jueconodon 2012:Condorodon 1952:Acinacodus 1880:see below↓ 1853:Allotheria 1841:Allotheria 1651:Steropodon 1643:Parvopalus 1612:Kryoryctes 1605:Kollikodon 1567:Kryoparvus 1529:Henosferus 1419:see below↓ 1398:Cynodontia 1383:Cynodontia 1331:Cynodontia 1322:Therapsida 1159:Syst Biol. 705:References 689:Monotremes 679:Marsupials 616:Sublegion 605:Holotheria 565:Metatheria 479:monotremes 431:Viviparity 417:monotremes 413:marsupials 407:) and the 401:eutherians 299:Tingamarra 275:Patagomaia 268:Metatheria 247:Subgroups 226:Subclass: 4279:Primates 4273:(Colugos) 4247:Rodentia 4142:(Hyraxes) 4079:Xenarthra 4066:Placental 3959:Marsupial 3699:Atlasodon 3620:includes 3591:includes 3488:Kermackia 3468:Hypomylos 3174:Stylodens 3050:Leonardus 3013:Bondesius 2940:Laolestes 2861:Achyrodon 2841:Tathiodon 2642:Lactodens 2382:Priacodon 2368:Jugulator 2220:Hangjinia 2134:Nakunodon 2084:Kemchugia 1857:includes 1819:Zaglossus 1754:Obdurodon 1674:Stirtodon 1371:Synapsida 1356:Synapsida 1313:Synapsida 1296:Kingdom: 1183:8 January 1141:0311-5518 1074:0962-8452 986:1476-4687 667:cladistic 510:Oxfordian 498:Evolution 468:pregnancy 455:process. 441:syncytins 183:Kingdom: 177:Eukaryota 4376:47048248 4328:Wikidata 4156:Sirenia 4058:Eutheria 3886:Chordata 3880:Animalia 3878:Kingdom 3587:Eutheria 3576:Eutheria 3518:Picopsis 3478:Juramaia 3268:Magnimus 3237:Arguimus 3207:Abelodon 3027:Cronopio 2882:Beckumia 2834:Paurodon 2549:Anebodon 2501:Yermakia 2310:Klamelia 1962:Amphidon 1620:Sundrius 1480:Ambondro 1451:Mammalia 1410:Mammalia 1304:Chordata 1302:Phylum: 1298:Animalia 1280:Mammalia 1177:Archived 1092:32517606 1002:49183466 994:29899454 939:34716275 880:22951979 823:Archived 801:25605903 753:syncytin 673:See also 618:Zatheria 594:Mammalia 572:Eutheria 551:Mammalia 531:Taxonomy 505:Juramaia 475:coracoid 445:placenta 389:subclass 296:Genus: † 280:Genus: † 272:Genus: † 259:Eutheria 207:Mammalia 197:Chordata 193:Phylum: 187:Animalia 173:Domain: 18:Therians 4334:Q130942 4095:Pilosa 3898:Amniota 3884:Phylum 3865:Extant 3750:UA 8699 3371:Peramus 3288:Mozomus 3278:Minimus 3167:Donodon 2487:Tinodon 2002:Comodon 1709:Opalios 1559:Bishops 1119:Bibcode 1083:7341916 966:Bibcode 930:8673075 907:Bibcode 871:3651898 850:Bibcode 792:4321253 767:Bibcode 641:Cohort 634:Cohort 610:Legion 447:, even 393:mammals 382:thēríon 239:Haswell 203:Class: 4464:Theria 4441:380416 4402:179916 4389:333419 4363:Theria 4349:Theria 4319:Theria 4196:(Bats) 3942:Theria 3869:orders 3867:mammal 3549:Theria 3448:Eomaia 2614:Aliaga 1225:  1205:  1155:et al. 1139:  1090:  1080:  1072:  1000:  992:  984:  958:Nature 937:  927:  878:  868:  799:  789:  592:Class 549:Class 485:Pinnae 375:θηρίον 312:Theria 241:, 1897 237:& 235:Parker 230:Theria 52:Theria 42:(moth) 40:Theria 4436:WoRMS 4428:39860 4415:32525 1336:Clade 1327:Clade 1318:Clade 1309:Clade 998:S2CID 464:fetus 429:(see 370: 264:Clade 255:Clade 214:Clade 4410:NCBI 4397:ITIS 3614:see 3585:see 1851:see 1396:see 1369:see 1223:ISBN 1203:ISBN 1185:2022 1137:ISSN 1088:PMID 1070:ISSN 990:PMID 982:ISSN 935:PMID 876:PMID 797:PMID 63:PreꞒ 4371:EoL 4358:ADW 1127:doi 1078:PMC 1062:doi 1058:287 1029:doi 974:doi 962:558 925:PMC 915:doi 903:118 866:PMC 858:doi 846:318 787:PMC 777:doi 763:112 437:egg 391:of 340:or 327:ɪər 4460:: 4438:: 4425:: 4412:: 4399:: 4386:: 4373:: 4360:: 4345:: 4330:: 1338:: 1329:: 1320:: 1311:: 1135:. 1125:. 1115:46 1113:. 1109:. 1086:. 1076:. 1068:. 1056:. 1052:. 1023:. 1019:. 996:. 988:. 980:. 972:. 960:. 956:. 933:. 923:. 913:. 901:. 897:. 874:. 864:. 856:. 844:. 840:. 817:. 795:. 785:. 775:. 761:. 757:. 733:. 481:. 353:ɛr 293:: 266:: 257:: 216:: 113:Pg 4064:( 3957:( 3858:e 3851:t 3844:v 3735:† 3725:† 3715:† 3705:† 3695:† 3561:† 3534:† 3524:† 3514:† 3504:† 3494:† 3484:† 3474:† 3464:† 3454:† 3444:† 3434:† 3424:† 3414:† 3404:† 3381:? 3352:? 3337:† 3328:? 3314:† 3304:† 3294:† 3284:† 3274:† 3264:† 3254:† 3250:? 3243:† 3233:† 3223:† 3213:† 3203:† 3141:† 3045:? 3037:? 2983:† 2935:? 2906:? 2801:? 2782:† 2595:† 2537:† 2528:? 2482:? 2467:† 2451:? 2422:† 2413:? 2321:† 2295:? 2290:† 2260:† 2230:? 2194:† 2185:? 2177:? 2170:† 2160:† 2150:† 2140:† 2130:† 2120:† 2110:† 2100:† 2090:† 2080:† 2070:† 2060:† 2050:† 2040:† 2036:? 2029:† 2025:? 2018:† 2008:† 1998:† 1988:† 1978:† 1968:† 1958:† 1948:† 1843:? 1838:† 1801:† 1791:† 1774:? 1767:† 1750:† 1746:? 1739:† 1735:? 1728:† 1705:† 1677:? 1662:† 1646:? 1631:† 1615:? 1597:? 1592:† 1570:? 1562:? 1540:† 1510:† 1501:? 1493:? 1486:† 1476:† 1465:? 1272:e 1265:t 1258:v 1187:. 1143:. 1129:: 1121:: 1094:. 1064:: 1037:. 1031:: 1025:1 1004:. 976:: 968:: 941:. 917:: 909:: 882:. 860:: 852:: 803:. 779:: 769:: 737:. 433:) 385:) 379:( 362:/ 359:ə 356:i 350:θ 347:ˈ 344:/ 336:/ 333:ə 330:i 324:θ 321:ˈ 318:/ 314:( 118:N 108:K 103:J 98:T 93:P 88:C 83:D 78:S 73:O 68:Ꞓ 44:. 34:. 20:)

Index

Therians
Otherkin § Terms and identities
Theria (moth)
PreꞒ

O
S
D
C
P
T
J
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Pg
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Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Mammalia
Tribosphenida
Theria
Parker
Haswell
Eutheria
Metatheria
Patagomaia
Spelaeomolitor
Incertae sedis

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