33:
576:
593:
330:
415:
360:. A smaller tail is located slightly more forward on the hindwing. The upperwings of these butterflies are largely or entirely colored brilliant blue. But these traits – as opposed to the androconia pouches discussed below – are also found in several other Eumaeini, which have been included in this genus but do not really appear to belong here.
1221:
396:
has golden-green hindwing undersides dusted with black, with a black band running roughly parallel to the distal wing margin, longer tails, and lacks sexual dichromatism. The
384:. The hindwing underside is brownish in these butterflies, with a subbasal line running between the subcosta and the anal vein branch of the cubital vein. This genus has some
377:
tucked away in a flat deep pouch in space 1b between the wing veins. Some species also have androconia on the forewings, but it needs to be clarified whether these are indeed
676:
1281:
1106:
1084:
1307:
672:
1135:
1255:
1363:
1353:
352:
but altogether still smallish butterflies. They typically have a pennant-like hindwing "tail" which projects outwards at an approximate
1294:
951:
as well as some species of the disputed assemblage are known from Santa
Catarina or its surroundings, while the rest of the
1104:
from the Andes and its spectroscopic characterization with some notes on the genus (Lepidoptera: Lycaenidae: Eumaeini).
439:, with about 30 species. A more recent proposal restricts the genus to 6-8 valid species, which all possess the typical
323:
1206:
1358:
1312:
802:
357:
867:
1149:
747:
1299:
897:
767:
460:
1211:
757:
597:
560:
530:
397:
777:
689:
520:
419:
907:
887:
540:
335:
1242:
919:
877:
827:
812:
787:
699:
633:
510:
490:
402:
185:
46:
1028:
857:
842:
737:
581:
475:
714:
615:
385:
627:
432:
214:
65:
675:, because an appropriate genus description was not given; the matter has been submitted to the
1348:
1320:
1286:
1229:
365:
310:
1325:
278:
289:
32:
435:
schemes for this genus in the late 20th century. The traditional view delimits the genus
400:
have a long, straight and tubular genital duct with a plain anal end in the type species
168:
592:
329:
1187:
654:
575:
414:
373:(scent scales) on the hindwing upperside near the base of the cubital vein; these are
1342:
939:
918:
In addition, there are at least three more undescribed species placed here: one from
645:
380:
653:
proper. All in all, it is not unlikely that some of the genera nowadays considered
722:
610:
444:
177:
57:
51:
1234:
1092:
1268:
1181:
1114:
667:
665:, and thus perhaps least likely to be valid. One of these has been argued to be
389:
374:
353:
243:
112:
1172:
1076:
Bálint, Zs.; Boyer, P.; Dahners, H.W.; Salazar-Escobar, J.A. & Kerstész, K.
956:
436:
370:
285:
281:
124:
662:
299:
296:
136:
88:
1198:
679:
for discussion. That all nonwithstanding, species provisionally placed in
1166:
639:
440:
349:
292:
148:
1260:
621:
304:
197:
1273:
1134:(2011): Markku Savela's Lepidoptera and some other life forms –
923:
100:
76:
1143:
1123:
609:, but apparently will have to be split off if the genus is to be a
927:
591:
574:
424:
413:
339:
328:
274:
41:
1049:(2006, 2007), Brower (2007), and see references in Savela (2011)
931:
1147:
1247:
1082:, a genus of rare Andean eumaeine Lycaenidae (Lepidoptera).
406:, but variation within the genus is insufficiently studied.
1078:(2006): Comments on the systematics and natural history of
1098:
Bálint, Zs.; Wojtusiak, J.; Kerstész, K. & Biró, L.P.
661:
turn out to be valid. Several of the proposed genera are
1140:. Version of December 24, 2011. Retrieved April 1, 2012.
1032:, whose female genitals markedly differ from those of
302:, it is usually placed in the group around the genus
1156:
579:Male imagines, upperside (left) and underwing, of
308:. In particular, it seems most closely related to
1128:. Version of 2008-MAR-24. Retrieved 2009-FEB-18.
637:and some allies could just as well be closer to
443:and can be divided into three species groups or
525:(Bálint, Wojtusiak, Kerstész & Biró, 2007)
258:
601:male imagines, upperside (left) and underwing
8:
585:, which has been proposed for separation in
392:of the blue upperwing color. By comparison,
1062:(2007), and see references in Savela (2011)
1144:
605:A number of species are usually placed in
31:
20:
1122:(2008): Tree of Life Web Project –
356:from the squared-off edge of the tapered
983:
981:
934:. It is unknown whether they are indeed
968:
7:
631:or even belong in the latter, while
241:D'Abrera & Bálint, 2001 (may be
16:Butterfly genus in family Lycaenidae
955:group but also many other related
431:There have been 2 major competing
14:
322:For a key to the terms used, see
427:, upperside (left) and underwing
277:of gossamer-winged butterflies (
975:See references in Savela (2011)
619:for example might be closer to
545:(C.Felder & R.Felder, 1865)
772:(Bálint & Wojtusiak, 2006)
1:
388:in that the males show green
227:Salazar & K.Johnson, 1997
348:species are large among the
324:Glossary of entomology terms
206:
1354:Lycaenidae of South America
1107:Acta Zool. Acad. Sci. Hung.
1100:(2007): The description of
1085:Acta Zool. Acad. Sci. Hung.
1380:
1364:Taxa named by Jacob HĂĽbner
321:
1113:(Supplement 1): 211–224.
571:Provisionally placed here
220:
213:
203:
196:
183:
176:
66:Scientific classification
64:
39:
30:
23:
959:are found in the Andes.
671:per Article 13.1 of the
482:(sometimes included in
467:(sometimes included in
803:Theritas curitibaensis
602:
589:
428:
342:
991:(2007), Brower (2007)
868:Theritas margaritacea
817:(C. & R.Felder, )
595:
578:
417:
332:
920:Santa Catarina state
748:Theritas augustinula
60:, underwing pattern
1120:Brower, Andrew V.Z.
898:Theritas theocritus
768:Theritas boliboyeri
461:Theritas kalikimaka
386:sexual dichromatism
205:6 to about 30, see
758:Theritas augustula
603:
590:
561:Theritas triquetra
531:Theritas harrietta
429:
343:
1359:Lycaenidae genera
1336:
1335:
1321:Open Tree of Life
1150:Taxon identifiers
1102:Theritas gozmanyi
926:and two from the
913:
903:
902:(Fabricius, 1793)
893:
883:
873:
863:
848:
834:(type species of
833:
819:(type species of
818:
808:
794:(type species of
793:
783:
778:Theritas chalumna
773:
763:
753:
743:
720:
705:
695:
690:Theritas acontius
566:
546:
536:
526:
521:Theritas gozmanyi
516:
496:
481:
466:
375:autapomorphically
266:
265:
255:
248:
235:
228:
1371:
1329:
1328:
1316:
1315:
1303:
1302:
1290:
1289:
1277:
1276:
1264:
1263:
1251:
1250:
1238:
1237:
1225:
1224:
1215:
1214:
1202:
1201:
1192:
1191:
1190:
1177:
1176:
1175:
1145:
1133:
1121:
1099:
1077:
1063:
1056:
1050:
1043:
1037:
1024:
1018:
1011:
1005:
998:
992:
985:
976:
973:
911:
908:Theritas viresco
901:
892:(Hewitson, 1865)
891:
888:Theritas phegeus
882:(Hewitson, 1867)
881:
871:
861:
846:
832:(Hewitson, 1874)
831:
816:
806:
791:
781:
771:
761:
751:
741:
718:
703:
693:
565:(Hewitson, 1865)
564:
544:
541:Theritas paupera
534:
524:
514:
494:
479:
464:
253:
240:
233:
226:
35:
21:
1379:
1378:
1374:
1373:
1372:
1370:
1369:
1368:
1339:
1338:
1337:
1332:
1324:
1319:
1311:
1306:
1298:
1293:
1285:
1280:
1272:
1267:
1259:
1254:
1246:
1241:
1233:
1228:
1220:
1218:
1210:
1205:
1197:
1195:
1186:
1185:
1180:
1171:
1170:
1165:
1152:
1131:
1119:
1097:
1075:
1072:
1067:
1066:
1057:
1053:
1044:
1040:
1025:
1021:
1012:
1008:
999:
995:
986:
979:
974:
970:
965:
916:
878:Theritas monica
828:Theritas deniva
813:Theritas danaus
807:(Johnson, 1992)
788:Theritas crines
752:(Goodson, 1845)
742:(Goodson, 1945)
700:Theritas adamsi
694:(Goodson, 1945)
655:junior synonyms
573:
511:Theritas drucei
491:Theritas mavors
480:(Goodson, 1945)
412:
398:female genitals
327:
320:
284:) found in the
256:
249:
236:
234:K.Johnson, 1992
229:
222:
192:
189:
186:Theritas mavors
172:
166:
165:
151:
139:
127:
115:
103:
91:
79:
55:
17:
12:
11:
5:
1377:
1375:
1367:
1366:
1361:
1356:
1351:
1341:
1340:
1334:
1333:
1331:
1330:
1317:
1304:
1291:
1278:
1265:
1252:
1239:
1226:
1216:
1203:
1193:
1178:
1162:
1160:
1154:
1153:
1148:
1142:
1141:
1132:Savela, Markku
1129:
1117:
1095:
1091:(3): 331–352.
1071:
1068:
1065:
1064:
1051:
1038:
1019:
1006:
993:
977:
967:
966:
964:
961:
915:
914:
904:
894:
884:
874:
872:(Draudt, 1919)
864:
858:Theritas lisus
854:
843:Theritas hemon
839:
824:
821:Margaritheclus
809:
799:
784:
782:(Schaus, 1902)
774:
764:
754:
744:
738:Theritas arene
734:
711:
696:
685:
587:Margaritheclus
572:
569:
568:
567:
548:
547:
537:
527:
517:
498:
497:
487:
476:Theritas lotis
472:
465:(Clench, 1944)
411:
408:
319:
316:
264:
263:
251:Margaritheclus
218:
217:
211:
210:
201:
200:
194:
193:
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181:
180:
174:
173:
167:
159:
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133:
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128:
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111:
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105:
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99:
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81:
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73:
69:
68:
62:
61:
37:
36:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
1376:
1365:
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1350:
1347:
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1344:
1327:
1322:
1318:
1314:
1309:
1305:
1301:
1296:
1292:
1288:
1283:
1279:
1275:
1270:
1266:
1262:
1257:
1253:
1249:
1244:
1240:
1236:
1231:
1227:
1223:
1217:
1213:
1208:
1204:
1200:
1194:
1189:
1183:
1179:
1174:
1168:
1164:
1163:
1161:
1159:
1155:
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1146:
1139:
1138:
1130:
1127:
1126:
1118:
1116:
1112:
1109:
1108:
1103:
1096:
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1090:
1087:
1086:
1081:
1074:
1073:
1069:
1061:
1055:
1052:
1048:
1042:
1039:
1035:
1031:
1030:
1023:
1020:
1016:
1010:
1007:
1003:
997:
994:
990:
984:
982:
978:
972:
969:
962:
960:
958:
954:
950:
946:
942:
941:
940:sensu stricto
937:
933:
929:
925:
921:
912:(Druce, 1907)
910:
909:
905:
900:
899:
895:
890:
889:
885:
880:
879:
875:
870:
869:
865:
862:(Stoll, 1790)
860:
859:
855:
852:
845:
844:
840:
837:
830:
829:
825:
822:
815:
814:
810:
805:
804:
800:
797:
792:(Druce, 1907)
790:
789:
785:
780:
779:
775:
770:
769:
765:
762:(Kirby, 1877)
760:
759:
755:
750:
749:
745:
740:
739:
735:
732:
728:
724:
719:(Druce, 1907)
717:
716:
715:Theritas anna
712:
709:
704:(Druce, 1909)
702:
701:
697:
692:
691:
687:
686:
684:
682:
678:
674:
670:
669:
664:
660:
656:
652:
648:
647:
646:Penaincisalia
642:
641:
636:
635:
630:
629:
624:
623:
618:
617:
612:
608:
600:
599:
598:T. theocritus
594:
588:
584:
583:
577:
570:
563:
562:
558:
557:
556:
555:
553:
543:
542:
538:
535:(Weeks, 1901)
533:
532:
528:
523:
522:
518:
515:(Lathy, 1926)
513:
512:
508:
507:
506:
505:
503:
493:
492:
488:
485:
478:
477:
473:
470:
463:
462:
458:
457:
456:
455:
453:
448:
446:
442:
438:
434:
426:
422:
421:
416:
409:
407:
405:
404:
399:
395:
391:
387:
383:
382:
381:sensu stricto
376:
372:
368:
367:
361:
359:
355:
351:
347:
341:
338:
337:
333:Underwing of
331:
325:
317:
315:
313:
312:
307:
306:
301:
298:
294:
291:
287:
283:
280:
276:
272:
271:
262:
260:
252:
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209:
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142:
138:
134:
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130:
126:
122:
119:
118:
114:
110:
107:
106:
102:
98:
95:
94:
90:
86:
83:
82:
78:
74:
71:
70:
67:
63:
59:
53:
49:
48:
43:
38:
34:
29:
26:
22:
19:
1157:
1136:
1124:
1115:PDF fulltext
1110:
1105:
1101:
1093:PDF fulltext
1088:
1083:
1080:Aveexcrenota
1079:
1059:
1054:
1046:
1041:
1033:
1027:
1022:
1014:
1009:
1004:(2006, 2007)
1001:
996:
988:
971:
952:
949:T. triquetra
948:
944:
938:
935:
917:
906:
896:
886:
876:
866:
856:
850:
841:
835:
826:
820:
811:
801:
795:
786:
776:
766:
756:
746:
736:
730:
729:; including
727:Aveexcrenota
726:
723:type species
713:
707:
698:
688:
680:
666:
658:
650:
644:
638:
632:
626:
620:
614:
611:monophyletic
606:
604:
596:
586:
580:
559:
551:
550:
549:
539:
529:
519:
509:
501:
500:
499:
495:HĂĽbner, 1818
489:
483:
474:
468:
459:
451:
450:
449:
445:superspecies
430:
420:T. augustula
418:
401:
393:
378:
369:, they have
364:
362:
345:
344:
334:
309:
303:
288:. Among the
269:
268:
267:
254:Bálint, 2002
250:
242:
237:
230:
224:Aveexcrenota
223:
221:
204:
191:HĂĽbner, 1818
184:
178:Type species
161:
160:
52:type species
45:
24:
18:
1269:iNaturalist
1182:Wikispecies
851:"T." deniva
731:A. mimianna
708:"T." deniva
668:nomen nudum
634:"T." deniva
441:apomorphies
390:iridescence
358:tornal area
354:right angle
318:Description
244:nomen nudum
132:Subfamily:
113:Lepidoptera
1343:Categories
1070:References
1029:"T." hemon
957:Lycaenidae
849:(close to
847:(Cramer, )
706:(close to
371:androconia
336:T. paupera
286:Neotropics
282:Lycaenidae
125:Lycaenidae
89:Arthropoda
1219:ButMoth:
1034:T. mavors
963:Footnotes
945:T. drucei
796:"Lucilda"
673:ICZN Code
663:monotypic
616:"T." anna
582:T. danaus
552:triquetra
484:T. mavors
469:T. mavors
433:taxonomic
403:T. mavors
379:Theritas
300:Theclinae
297:subfamily
257:(but see
137:Theclinae
72:Kingdom:
47:T. mavors
1349:Eumaeini
1295:LepIndex
1196:BioLib:
1188:Theritas
1173:Q7783003
1167:Wikidata
1158:Theritas
1137:Theritas
1125:Theritas
936:Theritas
681:Theritas
659:Theritas
651:Theritas
649:than to
640:Micandra
607:Theritas
425:imagines
350:Eumaeini
346:Theritas
293:Eumaeini
270:Theritas
215:Synonyms
162:Theritas
149:Eumaeini
120:Family:
84:Phylum:
77:Animalia
58:Amazonia
25:Theritas
1287:1308180
1261:1920921
1222:28961.0
1058:Bálint
1045:Bálint
1013:Bálint
1000:Bálint
987:Bálint
953:paupera
836:Denivia
622:Brangas
613:group.
502:paupera
437:loosely
410:Species
305:Atlides
295:of its
238:Lucilda
231:Denivia
198:Species
156:Genus:
144:Tribe:
108:Order:
101:Insecta
96:Class:
40:Female
1326:200521
1313:705835
1300:200092
1274:133422
1199:687512
1060:et al.
1047:et al.
1017:(2007)
1015:et al.
1002:et al.
989:et al.
924:Brazil
628:Evenus
452:mavors
279:family
171:, 1818
169:HĂĽbner
1282:IRMNG
1248:33334
1212:58850
1026:E.g.
928:Andes
683:are:
554:group
504:group
454:group
423:male
394:Arcas
366:Arcas
363:Like
340:imago
311:Arcas
290:tribe
275:genus
273:is a
56:from
42:imago
1308:NCBI
1256:GBIF
1235:7VR4
1207:BOLD
947:and
932:Peru
677:ICZN
643:and
625:and
259:text
207:text
1243:EoL
1230:CoL
930:of
922:in
725:of
657:of
44:of
1345::
1323::
1310::
1297::
1284::
1271::
1258::
1245::
1232::
1209::
1184::
1169::
1111:53
1089:52
980:^
943:.
853:?)
710:?)
447::
314:.
261:)
1036::
838:)
823:)
798:)
733:)
721:(
486:)
471:)
326:.
247:)
54:)
50:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
