Knowledge (XXG)

Tomicus piniperda

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the tree. There is one generation per year, with most adults dying after breeding many times, though a few survive to breed again a year later. In late winter to early spring, when daily high temperatures exceed 10–13 °C (50–55 °F), adults initiate flight from their overwintering sites and seek breeding material as a host, including recently cut pine trees, logs, branches, and stumps.
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thrown and less resistance pines in which the beetle almost exclusively breeds. Their mass aggregation in these pines is due to their ability to respond swiftly to monoterpenes that are released from injuries to the trees. This allows these beetles to locate mates quickly, even when these beetles are not able to utilise pheromones.
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After 2000, all infested states have surveyed only with the baited traps in spring and early summer. In Canada, techniques included using a combination of baited traps, trap logs, and visual inspections. In recent years, Canada has also started to use monoterpene baited traps. At United States ports of entry,
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infested tips. Another method of controlling this beetle is to use permethrin or carbaryl before adult mating flight. This usually occurs on the first warm day in spring. Throughout the summer, continue to treat shoots with permethrin. One of the best methods used to control these insects is to use a summer
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During September and October of 1992, seven states (Florida, Georgia, Kansas, Louisiana, North Caroline, Oregon, and West Viriginia) had their own state-wide quarantines on various pine articles from the other infested states. This included allowing importation of regulated articles only after being
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in strong apical shoots of healthy young trees, killing the bored-out shoots. This does not kill the tree, but causes damage to the growth form, lowering the economic value of the timber by reducing growth rates and stem straightness. They are also capable of damaging trees by infesting the trunk of
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is black or dark brown, 3.5–4.8 mm long, with a cylindrical body, rounded at the head and abdomen ends. It breeds in recently dead and dying trees, most often in windblown trees lying on the ground but also in fire-killed standing trees. The adults tunnel a breeding gallery in the spring, up to
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beetles did not influence the attraction of these beetles to trees. Both sexes respond similarly to a concentration range of each monoterpene, although myrcene was not as attractive to the beetles as the other monoterpenes. In addition to Scots pines trees, these beetles are also able to respond to
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From 1993 to 2000, US states within and near the quarantine zone used a combination of trap logs and monoterpene baited funnel traps to capture parent adults during their initial spring flight. Visual surveys were administered in late summer and autumn to locate the beetles' shoot-feeding damage.
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distributes itself among the tree to avoid conflict. They are found with the highest density near the top of tree trunks and towards the ground. Rarely, they may fully occupy the entirety of the trunk surface. They are most likely to attack the tree at increasing heights and lay their eggs there.
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has two main patterns of trunk attack. One method of attack that adults employ is to aggregate in the tree crowns first and then infest the trunk for breeding. A second method of attack is when adults directly attack without aggregating within the shoot. For either method of attack, colonisation
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physically is to place freshly cut pine logs or other trees in a pine field so that the beetle will attack the killed logs when finding a place to breed. After the end of their breeding period, remove and destroy all infested logs. During the summer and into early autumn, remove and destroy all
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This behaviour is especially important in Northern Europe and Scandinavia, where it is the most important insect pest of pines. The beetle typically swarms in the spring before other bark beetle species that infest pines, and as a result is successful in competing for the limited number of wind
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The federal quarantine disallowed movement of host material from infested areas to non-infested areas within the United States, including logs and lumber with bark from all species of pine. In 1993, stumps from recently cut pine trees were added as a regulated article. These pine stumps were
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make it a good candidate for local adaptive structure. These include mate location that takes place on a host plant, larval development being completed on one individual host, and selection pressures being highly variable between hosts, which magnify the strength of the insect-host plant
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success depends on the degree of tree resistance that the beetles encounter. Aggregation occurs towards the end of the shoot-feeding period, which intersects with the weakening of the tree. This explains the successful mass attacks that take place in the bole of trees and kills them.
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is regularly found and intercepted. During a 16 year period, this beetle was intercepted 151 times on cargo arriving from at least 18 different countries. Interception data helps to narrow the screening process when working with this beetle that occurs over a wide geographic range.
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volatiles released from logs and other types of trees, including the Norway spruce. However, the concentrations of monoterpenes found in Norway spruces are lower than in Scots pines, which in part explains why these beetles are more likely to select Scots pines as a host.
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from the Mediterranean that were sequenced are structured into two haplotypic groups. One group corresponds to most insects collected in Northern Europe (France) whereas the second group corresponds more similarly to Mediterranean insect populations. This distribution of
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adult found during inspection can make it impossible for producers of Christmas trees to fulfill their contracts. As a result, scientists and regulators developed the Pine Shoot Beetle Compliance Management Program for both pine Christmas trees and nursery stock.
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scent emitted by damaged specimens of the host species, including storm-fallen Scots pines. These trees are homes to bark beetles because of an injured vascular system that can not provide adequate resin to defend against new attacks by beetles.
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causes damage to various pine species throughout Northern Europe, trees in the Mediterranean are especially vulnerable and may even be killed. Larvae in the Mediterranean have a few characteristics that separate it from other populations of
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suggests that this beetle can be split into two separate species. This is further confirmed by the distance between the two haplotypic groups being similar to the distance between
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Kirkendall & Faccoli sp. n. (Curculionidae, Scolytinae), an unusually aggressive pine shoot beetle from southern China, with a key to the species of
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Furthermore, the intra-group distances of the two haplotypic groups are compatible with intra-specific variation commonly observed in insects.
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radially from the gallery for several months, emerging as new adults in late summer. The adults then feed through the autumn and winter on the
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in 1993 by the Canadian authorities. Surveys during 1993 and 1994 found several new infested counties but no new infested states. On average,
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influencing pupal survival. In addition, both benefit mutually from each other if tree resistance is high. Similar interactions occur between
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including Indiana, Pennsylvania, Michigan, New York, and Illinois. The first known occurrence in North America was found in July 1992 at a
944: 759: 668:. In addition, pine bark, pine Christmas trees, pine wreaths, and pine nursery stock were also considered regulated articles in 1993. 1796: 1561: 1380: 287:. Scots pine is the most important forest tree species in East-central Europe, with Scots pine occupying 68% of total forest area in 805: 1613: 869: 166: 320:
25 cm (9.8 in) long, parallel to the wood grain where they lay their eggs. On hatching, the larvae chew through the
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As with all bark beetle species, this species is known to associate with a wide range of fungal taxa, including the genera
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In Europe, the population of these beetles living in Mediterranean areas differ from the populations of other areas. While
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Vasconcelos, T., Nazare, N., Branco, M., Kerdelhue, C., Sauvard, D., & Lieutier, F. (2003). Host Preference of
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is able to recognize smells while still in flight by means of olfaction of several different plant
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Proceedings: JUFRO Kanazava 2003 “Forest Insect Population Dynamics and Host Influences"
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in the Mediterranean region, which differs in details of ecology, infesting primarily
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Horn, Agnès; Kerdelhué, Carole; Lieutier, François; Rossi, Jean-Pierre (2012-05-21).
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Byers, J. A.; Lanne, B. S.; Löfqvist, J.; Schlyter, F.; Bergström, G. (June 1985).
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and are forced to take parts of the trees that haven't been attacked yet.
1722: 1522: 1501: 1469: 1220:"Olfactory recognition of host-tree susceptibility by pine shoot beetles" 446: 256: 1527: 1219: 1605: 1443:. Lists general information and resources for Common Pine Shoot Beetle. 1243: 899:
in Poland and assessment of their virulence using Scots pine seedlings"
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for pre-beeding association and pairing, but instead hones in on the
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was found in more than 30 new countries per year from 1992 to 2000.
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coexists with other species in the trunks of pines. As a result,
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in the United States. As a precautionary step to help protect
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alongside the destruction of brood logs and trees during May.
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The beetle has been introduced accidentally to northeastern
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evaporating from wound resin. These monoterpenes include
1284:"Predicting the distribution of the two bark beetles 1459: 1069:Haack, Robert A.; Poland, Therese M. (2001-09-01). 796:J M Davies; C J King (1977). "Pine Shoot Beetles". 548:as a separate species. Several characteristics of 1439:, National Invasive Species Information Center, 693:There are multiple strategies to dealing with a 1115:Kirkendall, L. R.; Faccoli, M.; Hui Ye (2008). 742:is strongly associated with moulds and yeasts. 605:, from where it has spread to 11 states in the 307:Pine shoot galleries. Plate from John Curtis's 999:(Col., Scolytidae) on the trunk of the living 621:. The beetle has been identified as a serious 853: 851: 791: 789: 787: 785: 783: 781: 8: 1433:Species Profile - Common Pine Shoot Beetle ( 770:. Crop Protection Compendium. Archived from 1441:United States National Agricultural Library 1424:Institute of Food and Agricultural Sciences 507:attacks trees that are already infested by 1447: 1394:. University of New Mexico. Archived from 31: 20: 1110: 1108: 491:attack patterns are heavily regulated by 1117:"Description of the Yunnan shoot borer, 1292:in Europe and the Mediterranean region" 751: 828: 826: 824: 822: 820: 818: 483:This is unlike other species, such as 273:to a small extent, and more rarely on 1375: 1373: 1371: 1369: 1367: 1365: 1363: 1277: 1275: 1273: 1271: 1269: 1267: 1265: 1263: 1261: 1213: 1211: 1209: 1207: 1205: 1203: 1201: 1199: 1197: 1157: 1155: 1153: 1064: 1062: 1060: 1058: 1056: 984: 982: 664:originally used to generate fuel and 527:in Europe and Mediterranean regions. 40:Adult feeding in pine shoot, Hungary 7: 1593:07d2c619-d91b-4ea4-b8ae-3848463d20a1 1054: 1052: 1050: 1048: 1046: 1044: 1042: 1040: 1038: 1036: 995:on distribution and reproduction of 980: 978: 976: 974: 972: 970: 968: 966: 964: 962: 888: 886: 884: 1328:Ministry of Agriculture and Lands: 989:Hui, Y.; Xue-Song, D. (July 1999). 738:attack them. In addition to fungi, 701:One method that is used to control 499:attacks occur 1-2 weeks later than 1296:Agricultural and Forest Entomology 893:Jankowiak, Robert (October 2006). 14: 1792:Insect vectors of plant pathogens 1787:Insect pests of temperate forests 1308:10.1111/j.1461-9563.2012.00576.x 1168:Annales des Sciences Forestières 1019:10.1046/j.1439-0418.1999.00353.x 949:Pests and Diseases Image Library 842:Global Invasive Species Database 49: 659:inspected and declared free of 470:Interactions with other species 411:Damage to a log by T. piniperda 1162:Hui, Y.; Lieutier, F. (1997). 1: 1007:Journal of Applied Entomology 798:Forestry Commission Leaflet 3 388:, which has consequences for 1802:Taxa named by Carl Linnaeus 337:Species closely related to 1818: 1342:"Common Pine Shoot Beetle" 420:Unlike most bark beetles, 227:Its primary host plant is 1797:Beetles described in 1758 1392:Pests of trees and shrubs 1144:10.11646/zootaxa.1819.1.2 585:, where it has become an 181: 174: 151: 144: 46:Scientific classification 44: 39: 30: 23: 903:Annals of Forest Science 865:for Three Pine Species. 556:Studies have found that 457:. The presence of other 206:common pine shoot beetle 1180:10.1051/forest:19970704 1091:10.1023/A:1015298114837 895:"Fungi associated with 386:reproductively isolated 923:10.1051/forest:2006063 655: 412: 313: 1744:Paleobiology Database 1420:University of Florida 1348:. Don't Move Firewood 651: 644:Management strategies 416:Olfactory recognition 410: 306: 190:Myelophilus piniperda 1588:Fauna Europaea (new) 1079:Biological Invasions 295:an important pest. 1381:"Pine shoot beetle 1236:1985NW.....72..324B 1224:Naturwissenschaften 1119:Tomicus yunnanensis 945:"Pine shoot beetle" 915:2006AnFSc..63..801J 714:Fungal associations 531:Population genetics 367:Tomicus yunnanensis 236:European black pine 234:, but it also uses 185:Dermestes piniperda 1427:Featured Creatures 1244:10.1007/bf00454776 875:2011-07-22 at the 656: 653:Damage to branches 413: 314: 310:British Entomology 250:eastern white pine 212:native throughout 1769: 1768: 1759:tomicus-piniperda 1731:Open Tree of Life 1491:Tomicus piniperda 1461:Tomicus piniperda 1453:Taxon identifiers 1435:Tomicus piniperda 1416:pine shoot beetle 1401:on 25 April 2024. 1383:Tomicus piniperda 1330:Pine Shoot Beetle 1290:Tomicus piniperda 1286:Tomicus destruens 1073:Tomicus piniperda 1001:Pinus yunnanensis 997:Tomicus piniperda 897:Tomicus piniperda 863:Tomicus destruens 859:Tomicus piniperda 836:Tomicus piniperda 768:CAB International 762:Tomicus piniperda 577:Invasive problems 422:Tomicus piniperda 378:Pinus yunnanensis 348:Tomicus destruens 339:Tomicus piniperda 201:Tomicus piniperda 197: 196: 155:Tomicus piniperda 137:T. piniperda 25:Tomicus piniperda 16:Species of beetle 1809: 1762: 1761: 1752: 1751: 1739: 1738: 1726: 1725: 1713: 1712: 1700: 1699: 1697:NBNSYS0000025550 1687: 1686: 1674: 1673: 1661: 1660: 1648: 1647: 1635: 1634: 1622: 1621: 1609: 1608: 1596: 1595: 1583: 1582: 1570: 1569: 1557: 1556: 1544: 1543: 1531: 1530: 1518: 1517: 1505: 1504: 1495: 1494: 1493: 1480: 1479: 1478: 1448: 1403: 1402: 1400: 1389: 1377: 1358: 1357: 1355: 1353: 1346:Gallery of Pests 1338: 1332: 1326: 1320: 1319: 1279: 1256: 1255: 1215: 1192: 1191: 1159: 1148: 1147: 1129: 1112: 1103: 1102: 1066: 1031: 1030: 986: 957: 956: 951:. Archived from 941: 935: 934: 890: 879: 855: 846: 845: 830: 813: 811: 800:. London: HMSO. 793: 776: 775: 756: 731:Ceratocystiopsis 587:invasive species 495:attacks because 474:In many places, 369:in southwestern 232:Pinus sylvestris 157: 54: 53: 35: 21: 1817: 1816: 1812: 1811: 1810: 1808: 1807: 1806: 1772: 1771: 1770: 1765: 1757: 1755: 1747: 1742: 1734: 1729: 1721: 1718:Observation.org 1716: 1708: 1703: 1695: 1690: 1682: 1677: 1669: 1664: 1656: 1651: 1643: 1638: 1630: 1625: 1617: 1612: 1604: 1599: 1591: 1586: 1578: 1573: 1565: 1560: 1552: 1547: 1539: 1534: 1526: 1521: 1513: 1508: 1500: 1498: 1489: 1488: 1483: 1474: 1473: 1468: 1455: 1412: 1407: 1406: 1398: 1387: 1379: 1378: 1361: 1351: 1349: 1340: 1339: 1335: 1327: 1323: 1281: 1280: 1259: 1217: 1216: 1195: 1161: 1160: 1151: 1127: 1114: 1113: 1106: 1068: 1067: 1034: 988: 987: 960: 943: 942: 938: 892: 891: 882: 877:Wayback Machine 856: 849: 832: 831: 816: 808: 795: 794: 779: 758: 757: 753: 748: 716: 691: 646: 579: 533: 503:attacks. Thus, 472: 418: 398: 384:, but they are 335: 301: 220:, and northern 216:, northwestern 170: 159: 153: 140: 48: 17: 12: 11: 5: 1815: 1813: 1805: 1804: 1799: 1794: 1789: 1784: 1774: 1773: 1767: 1766: 1764: 1763: 1753: 1740: 1727: 1714: 1701: 1688: 1675: 1662: 1649: 1636: 1623: 1610: 1597: 1584: 1575:Fauna Europaea 1571: 1558: 1545: 1532: 1519: 1506: 1496: 1481: 1465: 1463: 1457: 1456: 1451: 1445: 1444: 1430: 1411: 1410:External links 1408: 1405: 1404: 1359: 1333: 1321: 1302:(4): 358–366. 1257: 1230:(6): 324–326. 1193: 1174:(7): 635–641. 1149: 1104: 1085:(3): 307–322. 1032: 1013:(6): 329–333. 958: 955:on 2008-07-29. 936: 909:(7): 801–808. 880: 847: 814: 806: 777: 774:on 2006-10-04. 750: 749: 747: 744: 715: 712: 690: 687: 645: 642: 597:farm close to 595:Christmas tree 578: 575: 553:relationship. 532: 529: 471: 468: 417: 414: 397: 394: 334: 331: 300: 297: 195: 194: 193: 192: 187: 179: 178: 172: 171: 160: 149: 148: 142: 141: 134: 132: 128: 127: 120: 116: 115: 110: 106: 105: 100: 96: 95: 90: 86: 85: 80: 76: 75: 70: 66: 65: 60: 56: 55: 42: 41: 37: 36: 28: 27: 15: 13: 10: 9: 6: 4: 3: 2: 1814: 1803: 1800: 1798: 1795: 1793: 1790: 1788: 1785: 1783: 1780: 1779: 1777: 1760: 1754: 1750: 1745: 1741: 1737: 1732: 1728: 1724: 1719: 1715: 1711: 1706: 1702: 1698: 1693: 1689: 1685: 1680: 1676: 1672: 1667: 1663: 1659: 1654: 1650: 1646: 1641: 1637: 1633: 1628: 1624: 1620: 1615: 1611: 1607: 1602: 1598: 1594: 1589: 1585: 1581: 1576: 1572: 1568: 1563: 1559: 1555: 1550: 1546: 1542: 1537: 1533: 1529: 1524: 1520: 1516: 1511: 1507: 1503: 1497: 1492: 1486: 1482: 1477: 1471: 1467: 1466: 1464: 1462: 1458: 1454: 1449: 1442: 1438: 1436: 1431: 1428: 1425: 1421: 1417: 1414: 1413: 1409: 1397: 1393: 1386: 1384: 1376: 1374: 1372: 1370: 1368: 1366: 1364: 1360: 1347: 1343: 1337: 1334: 1331: 1325: 1322: 1317: 1313: 1309: 1305: 1301: 1297: 1293: 1291: 1287: 1278: 1276: 1274: 1272: 1270: 1268: 1266: 1264: 1262: 1258: 1253: 1249: 1245: 1241: 1237: 1233: 1229: 1225: 1221: 1214: 1212: 1210: 1208: 1206: 1204: 1202: 1200: 1198: 1194: 1189: 1185: 1181: 1177: 1173: 1169: 1165: 1158: 1156: 1154: 1150: 1145: 1141: 1137: 1133: 1126: 1124: 1120: 1111: 1109: 1105: 1100: 1096: 1092: 1088: 1084: 1080: 1076: 1075:(Coleoptera)" 1074: 1065: 1063: 1061: 1059: 1057: 1055: 1053: 1051: 1049: 1047: 1045: 1043: 1041: 1039: 1037: 1033: 1028: 1024: 1020: 1016: 1012: 1008: 1004: 1002: 998: 994: 993:Tomicus minor 985: 983: 981: 979: 977: 975: 973: 971: 969: 967: 965: 963: 959: 954: 950: 946: 940: 937: 932: 928: 924: 920: 916: 912: 908: 904: 900: 898: 889: 887: 885: 881: 878: 874: 871: 868: 864: 860: 854: 852: 848: 843: 839: 837: 829: 827: 825: 823: 821: 819: 815: 809: 807:0-11-710219-9 803: 799: 792: 790: 788: 786: 784: 782: 778: 773: 769: 765: 763: 755: 752: 745: 743: 741: 737: 733: 732: 727: 726: 722: 713: 711: 709: 704: 700: 696: 688: 686: 683: 678: 675: 669: 667: 662: 661:T. piniperda. 654: 650: 643: 641: 639: 635: 631: 628: 624: 620: 616: 612: 608: 607:United States 604: 600: 596: 592: 591:T. piniperda, 588: 584: 583:North America 576: 574: 572: 568: 564: 559: 554: 551: 547: 543: 538: 530: 528: 526: 522: 518: 514: 510: 506: 502: 498: 494: 490: 486: 481: 477: 469: 467: 463: 460: 456: 452: 448: 444: 440: 436: 431: 427: 424:does not use 423: 415: 409: 405: 402: 395: 393: 391: 387: 383: 379: 376: 372: 368: 364: 361: 360:maritime pine 357: 354: 350: 349: 344: 343:Tomicus minor 340: 332: 330: 327: 323: 318: 312: 311: 305: 298: 296: 294: 290: 286: 283: 279: 276: 272: 268: 265: 261: 258: 254: 251: 247: 244: 243:maritime pine 240: 237: 233: 230: 225: 223: 219: 215: 211: 207: 203: 202: 191: 188: 186: 183: 182: 180: 177: 173: 168: 164: 158: 156: 150: 147: 146:Binomial name 143: 139: 138: 133: 130: 129: 126: 125: 121: 118: 117: 114: 113:Curculionidae 111: 108: 107: 104: 101: 98: 97: 94: 91: 88: 87: 84: 81: 78: 77: 74: 71: 68: 67: 64: 61: 58: 57: 52: 47: 43: 38: 34: 29: 26: 22: 19: 1460: 1434: 1426: 1396:the original 1391: 1382: 1350:. Retrieved 1345: 1336: 1324: 1299: 1295: 1289: 1285: 1227: 1223: 1171: 1167: 1135: 1131: 1122: 1118: 1082: 1078: 1072: 1010: 1006: 1000: 996: 992: 991:"Impacts of 953:the original 948: 939: 906: 902: 896: 866: 862: 858: 841: 835: 797: 772:the original 767: 761: 754: 740:T. piniperda 739: 736:T. piniperda 735: 729: 721:Ceratocystis 719: 717: 708:foliar spray 703:T. piniperda 702: 698: 695:T. piniperda 694: 692: 682:T. piniperda 681: 679: 674:T. piniperda 673: 670: 660: 657: 652: 638:T. piniperda 637: 590: 580: 570: 567:T. piniperda 566: 563:T. piniperda 562: 558:T. piniperda 557: 555: 550:T. piniperda 549: 546:T. piniperda 545: 542:T. piniperda 541: 537:T. piniperda 536: 534: 525:T. piniperda 524: 521:T. destruens 520: 517:T. piniperda 516: 512: 509:T. piniperda 508: 504: 501:T. piniperda 500: 496: 493:T. piniperda 492: 488: 484: 480:T. piniperda 479: 476:T. piniperda 475: 473: 464: 459:T. piniperda 458: 443:alpha-pinene 439:monoterpenes 435:T. piniperda 434: 421: 419: 401:T. piniperda 400: 399: 390:pest control 382:T. piniperda 381: 377: 366: 362: 355: 346: 342: 338: 336: 317:T. piniperda 316: 315: 308: 293:T. piniperda 292: 284: 277: 267:P. banksiana 266: 259: 252: 245: 238: 231: 226: 205: 200: 199: 198: 189: 184: 154: 152: 136: 135: 123: 24: 18: 1679:NatureServe 1627:iNaturalist 1485:Wikispecies 697:infestation 630:plantations 451:terpinolene 375:Yunnan pine 363:P. pinaster 299:Description 260:P. resinosa 246:P. pinaster 210:bark beetle 1782:Scolytinae 1776:Categories 1352:12 October 746:References 725:Ophiostoma 666:turpentine 426:pheromones 353:stone pine 269:and other 253:P. strobus 229:Scots pine 103:Coleoptera 83:Arthropoda 1316:1461-9555 1252:0028-1042 1188:0003-4312 1138:: 25–39. 1099:1573-1464 1027:0931-2048 931:1286-4560 838:(insect)" 680:A single 599:Cleveland 571:T. minor. 396:Behaviour 291:, making 264:jack pine 131:Species: 69:Kingdom: 63:Eukaryota 1684:2.816163 1645:11614692 1523:BugGuide 1499:BioLib: 1476:Q1256962 1470:Wikidata 873:Archived 515:affects 513:T. minor 505:T. minor 497:T. minor 489:T. minor 485:T. minor 447:3-carene 356:P. pinea 341:include 333:Taxonomy 257:red pine 239:P. nigra 176:Synonyms 163:Linnaeus 109:Family: 79:Phylum: 73:Animalia 59:Domain: 1671:1085382 1606:1211257 1429:website 1418:on the 1232:Bibcode 1132:Zootaxa 1123:Tomicus 911:Bibcode 689:Control 611:Ontario 609:and to 455:myrcene 208:, is a 124:Tomicus 119:Genus: 99:Order: 93:Insecta 89:Class: 1749:302851 1736:363101 1710:143020 1632:178812 1580:250004 1567:268544 1554:BLASPI 1528:731856 1314:  1250:  1186:  1097:  1025:  1003:trees" 929:  804:  728:, and 634:Canada 619:Canada 615:Quebec 453:, and 365:; and 322:phloem 289:Poland 275:spruce 218:Africa 214:Europe 204:, the 1756:PPE: 1723:20298 1658:54154 1640:IRMNG 1562:EUNIS 1541:57D3N 1515:32272 1502:13879 1399:(PDF) 1388:(PDF) 1128:(PDF) 870:19–21 430:resin 371:China 285:Larix 282:larch 278:Picea 271:pines 1705:NCBI 1666:ITIS 1619:1200 1614:GISD 1601:GBIF 1549:EPPO 1510:BOLD 1354:2011 1312:ISSN 1288:and 1248:ISSN 1184:ISSN 1136:1819 1095:ISSN 1023:ISSN 927:ISSN 861:and 802:ISBN 627:pine 623:pest 613:and 603:Ohio 569:and 523:and 358:and 326:pith 280:and 222:Asia 167:1758 1692:NBN 1653:ISC 1536:CoL 1304:doi 1240:doi 1176:doi 1140:doi 1087:doi 1015:doi 1011:123 919:doi 617:in 373:on 1778:: 1746:: 1733:: 1720:: 1707:: 1694:: 1681:: 1668:: 1655:: 1642:: 1629:: 1616:: 1603:: 1590:: 1577:: 1564:: 1551:: 1538:: 1525:: 1512:: 1487:: 1472:: 1422:/ 1390:. 1362:^ 1344:. 1310:. 1300:14 1298:. 1294:. 1260:^ 1246:. 1238:. 1228:72 1226:. 1222:. 1196:^ 1182:. 1172:54 1170:. 1166:. 1152:^ 1134:. 1130:. 1107:^ 1093:. 1081:. 1077:. 1035:^ 1021:. 1009:. 1005:. 961:^ 947:. 925:. 917:. 907:63 905:. 901:. 883:^ 850:^ 840:. 817:^ 780:^ 766:. 723:, 601:, 449:, 445:, 392:. 262:, 255:, 248:, 241:, 165:, 1437:) 1385:" 1356:. 1318:. 1306:: 1254:. 1242:: 1234:: 1190:. 1178:: 1146:. 1142:: 1125:" 1101:. 1089:: 1083:3 1029:. 1017:: 933:. 921:: 913:: 844:. 834:" 812:. 810:. 764:" 760:" 699:. 169:) 161:(

Index


Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Coleoptera
Curculionidae
Tomicus
Binomial name
Linnaeus
1758
Synonyms
bark beetle
Europe
Africa
Asia
Scots pine
European black pine
maritime pine
eastern white pine
red pine
jack pine
pines
spruce
larch
Poland

British Entomology

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