363:
1941:
359:
species, the female occupies the burrow of their mate while she lays her clutch of eggs. Research shows that the male major claw size is also correlated with burrow width; the width of the burrow influences incubation temperature. Therefore, the female will choose a male mate whose claw size indicates the best burrow environment for her clutch of eggs. The waving display is also thought to indicate to females the overall healthiness of the male; a more vigorous display is more difficult to do and thus requires the male to be in prime health condition, which suggests that the male will help produce viable offspring.
2036:
1981:
82:
351:
1965:
302:
58:
342:, the sediment is replaced in the form of a little ball. The presence of these sediment balls near the entrance to a burrow is a good indication of its occupation. Some experts believe that the feeding habits of fiddler crabs play a vital role in the preservation of wetland environments; by sifting through the sands, they aerate the substrate and prevent anaerobic conditions.
2655:
2138:
Levinton, J. S., Judge, M. L., and
Kurdziel, J. P., 1995, Functional differences between the major and minor claws of fiddler crabs (Uca, family Ocypodidae, order Decapoda, Subphylum Crustacea): A result of selection or developmental constraint?: Journal of Experimental Marine Biology and Ecology, v.
234:
species are also found in the family
Ocypodidae. This entire group is composed of small crabs, the largest being slightly over two inches (5 cm) across. Fiddler crabs are found along sea beaches and brackish intertidal mud flats, lagoons, swamps, and various other types of brackish or salt-water
358:
Fiddler crabs live rather brief lives of no more than two years (up to three years in captivity). Male fiddler crabs use the major claw to perform a waving display as a form of female courtship. Females choose their mate based on claw size and also quality of the waving display. In many fiddler crab
2066:
Fiddler crabs are occasionally kept as pets. The fiddler crabs sold in pet stores generally come from brackish water lagoons. Because they live in lower salinity water, pet stores may call them fresh-water crabs, but they cannot survive indefinitely in fresh water. Fiddler crabs have been known to
246:
as they grow. If they have lost legs or claws during their present growth cycle, a new one will be present when they molt. If the large fiddle claw is lost, males will develop one on the same side after their next molt. Newly molted crabs are very vulnerable because of their soft shells. They are
370:
Male versus male competition also occurs as fighting with the major claws. The female fiddler carries her eggs in a mass on the underside of her body. She remains in her burrow during a two-week gestation period, after which she ventures out to release her eggs into the receding tide. The larvae
319:; used in clashes of ritualised combat of courtship over a female and signal their intentions between conspecifics. The movement of the smaller claw from ground to mouth during feeding explains the crabs' common name; it looks as if the animal were playing the larger claw like a fiddle.
2003:
2035:
2007:
2011:
2009:
2005:
2004:
2010:
2442:"Systematics of the family Ocypodidae Rafinesque, 1815 (Crustacea: Brachyura), based on phylogenetic relationships, with a reorganization of subfamily rankings and a review of the taxonomic status of Uca Leach, 1814, sensu lato and its subgenera"
2008:
384:
have been shown to bluff about their fighting ability. Upon regrowing a lost claw, a crab will occasionally regrow a weaker claw that nevertheless intimidates crabs with smaller but stronger claws. This is an example of dishonest
392:
The dual functionality of the major claw of fiddler crabs has presented an evolutionary conundrum in that the claw mechanics best suited for fighting do not match up with the mechanics best suited for a waving display.
2355:
Callander, S., Kahn, A. T., Maricic, T., Jennions, M. D., and
Backwell, P. R. Y., 2013, Weapons or mating signals? Claw shape and mate choice in a fiddler crab: Behavioral Ecology and Sociobiology, v. 67, p.
2328:
Perez, D. M., Rosenberg, M. S., and Pie, M. R., 2012, The evolution of waving displays in fiddler crabs (Uca spp., Crustacea: Ocypodidae): Biological
Journal of the Linnean Society, v. 106, p. 307-315.
2337:
Reaney, L. T., and
Backwell, P. R. Y., 2007, Temporal constraints and female preference for burrow width in the fiddler crab, Uca mjoebergi: Behavioral Ecology and Sociobiology, v. 61, p. 1515-1521.
1940:
2346:
Matsumasa, M., Murai, M., and
Christy, J. H., 2013, A low-cost sexual ornament reliably signals male condition in the fiddler crab Uca beebei: Animal Behaviour, v. 85, p. 1335-1341.
1980:
2006:
362:
2430:
Swanson, B. O., George, M. N., Anderson, S. P., and
Christy, J. H., 2013, Evolutionary variation in the mechanics of fiddler crab claws: Bmc Evolutionary Biology, v. 13.
2851:
2540:"Tubuca alcocki, a new pseudocryptic species of fiddler crab from the Indian Ocean, sister to the southeastern African T. urvillei (H. Milne Edwards, 1852)"
322:
The crab's smaller claw picks up a chunk of sediment from the ground and brings it to the mouth, where its contents are sifted through (making the crab a
2319:
Pope, D. S., 2000, Testing function of fiddler crab claw waving by manipulating social context: Behavioral
Ecology and Sociobiology, v. 47, p. 432-437.
2812:
2838:
2672:
2197:
2596:
226:
claws; the males' major claw is much larger than the minor claw, while the females' claws are both the same size. A smaller number of
2367:
2903:
2954:
2464:
2843:
2371:
2402:
2760:
2472:
2018:
1386:
81:
2226:
2856:
2095:
2118:"A fresh look at the biodiversity lexicon for fiddler crabs (Decapoda: Brachyura: Ocypodidae). Part 1: Taxonomy"
1746:
725:
685:
2921:
2263:"Effects of Fiddler Crab Burrows on Sediment Properties in the Mangrove Mudflats of Sungai Sepang, Malaysia"
1375:
1624:
1412:
1032:
572:
535:
2700:
1216:
1197:
807:
735:
1613:
1602:
1208:
705:
695:
675:
660:
2877:
1917:
1650:
1432:
1401:
1160:
1069:
1047:
888:
715:
637:
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583:
561:
251:
2765:
1897:
1484:
1318:
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1175:
1125:
1110:
1058:
899:
855:
491:
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1758:
1672:
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1304:
947:
921:
652:
502:
475:
2944:
2908:
2747:
2665:
2214:
1964:
1498:
1227:
1186:
1140:
1084:
980:
910:
603:
513:
483:
31:
2504:
sp. nov., a new species of fiddler crab (Crustacea: Brachyura: Ocypodidae) from the
Western Pacific"
1827:
1783:
1698:
1565:
1473:
1340:
866:
1888:
1882:
1816:
1774:
1719:
1556:
1539:
1530:
1392:
1329:
1260:
1038:
552:
466:
1871:
1850:
1768:
1451:
1351:
413:
were elevated to genus rank, and the fiddler crabs now occupy 11 genera making up the subfamilies
2949:
2243:
2076:
2023:
1806:
1736:
1509:
1368:
1021:
995:
844:
524:
460:
380:
76:
2604:
1860:
1839:
1271:
2882:
2440:
Shih, Hsi-Te; Ng, Peter K. L.; Davie, Peter J. F.; Schubart, Christoph D.; et al. (2016).
2150:
1794:
615:
2864:
2786:
2567:
2465:"Systema Brachyurorum: Part I. An annotated checklist of extant Brachyuran crabs of the world"
2302:
2284:
2193:
2170:
1550:
969:
829:
818:
757:
546:
386:
307:
223:
1425:
596:
258:
that mimics the ebb and flow of the tides: they turn dark during the day and light at night.
2869:
2557:
2547:
2520:
2411:
2292:
2274:
2235:
2162:
1990:
1910:
1524:
1010:
777:
746:
255:
2659:
2394:"Regenerated claws dishonestly signal performance and fighting ability in the fiddler crab
2676:
1713:
1576:
1099:
958:
877:
301:
2050:
2499:
2117:
350:
2562:
2297:
2262:
1971:
1687:
1462:
936:
792:
429:
315:
Fiddler crabs communicate by a sequence of waves and gestures; males have an oversized
279:
401:
More than 100 species of fiddler crabs make up 11 of the 13 genera in the crab family
366:
Fiddler Crabs
Fighting in Belle Hall, Mount Pleasant, South Carolina on March 24, 2023
2938:
2441:
2416:
2393:
2046:
1704:
1678:
1641:
1635:
1582:
1515:
1490:
1357:
1309:
1282:
1166:
1153:
1131:
1116:
1090:
1075:
1001:
986:
927:
835:
798:
783:
666:
643:
628:
316:
283:
231:
46:
2539:
2247:
2791:
2481:
2261:
Mokhtari, Mohammad; Abd Ghaffar, Mazlan; Usup, Gires; Che Cob, Zaidi (2016-01-19).
1661:
1293:
1249:
287:
123:
65:
2392:
Simon P. Lailvaux; Leeann T. Reaney; Patricia R. Y. Backwell (November 11, 2008).
2239:
2627:
2825:
2732:
414:
323:
275:
187:
163:
143:
2723:
298:, fiddler crabs are easily recognized by their distinctively asymmetric claws.
2552:
2524:
402:
227:
219:
173:
2738:
2288:
2174:
2799:
2752:
2600:
2166:
2067:
attack small fish in captivity, as opposed to their natural feeding habits.
2042:
1994:
1956:
113:
93:
2571:
2306:
2279:
2654:
57:
2773:
2717:
442:
372:
339:
295:
267:
236:
133:
2778:
2916:
2817:
2688:
2511:
2045:
of male fiddler crabs make them useful figures in illustrating the non-
770:
331:
291:
243:
2215:"Claw waving display changes with receiver distance in fiddler crabs,
214:
may be any of more than one hundred species of semiterrestrial marine
2830:
2192:(8 ed.). San Francisco: Pearson Benjamin Cummings. p. 144.
1444:
422:
335:
103:
2694:
2804:
2628:"The Complete Fiddler Crab Care Guide: Tank, Food, Facts and More…"
17:
2890:
2027:
1952:
361:
349:
300:
271:
2538:
Shih, H. T.; Chan, Benny K. K.; Ng, Peter K. L. (29 March 2018).
327:
215:
153:
2698:
2682:
2368:"Fiddler crabs reveal honesty is not always the best policy"
2689:
The colorful fiddler crabs in the mangrove forest of Borneo
2151:"On the Other Hand: The Myth of Fiddler Crab Claw Reversal"
2463:
Peter K. L. Ng; Danièle Guinot; Peter J. F. Davie (2008).
2683:
Info on systematics, phylogeny and morphology of fiddlers
680:(Shih, Naruse & Ng, 2010) (jocelyn's fiddler crab)
2707:
2213:M. J. How; J. M. Hemmi; J. Zeil; R. Peters (2008).
179:
75:
41:
2498:Hsi-Te Shih; Tohru Naruse; Peter K. L. Ng (2010).
2366:British Ecological Society (November 13, 2008).
247:reclusive and hide until the new shell hardens.
2111:
2109:
8:
326:). After anything edible is salvaged, be it
405:. These were formerly members of the genus
2695:
56:
38:
30:"Uca" redirects here. For other uses, see
2590:
2588:
2561:
2551:
2415:
2296:
2278:
270:, in salt marshes, and on sandy or muddy
2597:"How to Set Up a Tank for Fiddler Crabs"
2087:
2049:of certain geometric objects, like the
1936:
305:Male lemon-yellow clawed fiddler crab (
2188:Smith, Thomas; Smith, Robert (2012).
540:(Naderloo, Schubart & Shih, 2016)
7:
2922:9AE90DF7-8F95-4D5A-AC16-B8A6969C40E0
2896:646187D2-FFD3-7070-1EDC-EE34FB527293
409:. In 2016, most of the subgenera of
496:(Naderloo, Türkay & Chen, 2010)
254:, fiddler crabs exhibit a constant
1276:(Landstorfer & Schubart, 2010)
25:
354:General anatomy of a fiddler crab
311:), waving his big claw in display
2653:
2417:10.1111/j.1365-2435.2008.01501.x
2034:
2001:
1979:
1963:
1939:
1347:(Atlantic hairback fiddler crab)
1106:(Atlantic mangrove fiddler crab)
80:
2446:The Raffles Bulletin of Zoology
2096:"Uca Leach 1814 (fiddler crab)"
1314:(Pacific mangrove fiddler crab)
1204:(Pacific hairback fiddler crab)
1136:(Pacific mangrove fiddler crab)
633:(northern calling fiddler crab)
2116:Rosenberg, Michael S. (2019).
1043:(narrow-fingered fiddler crab)
671:(western calling fiddler crab)
656:(eastern calling fiddler crab)
242:Like all crabs, fiddler crabs
1:
2372:University of New South Wales
2240:10.1016/j.anbehav.2007.09.004
2149:Weis, Judith S (2019-04-01).
2122:Journal of Crustacean Biology
1657:(Asian mangrove fiddler crab)
1380:(Shih, Komai & Liu, 2013)
1362:(lesser Mexican fiddler crab)
1334:(Salmon & Atsaides, 1968)
1289:(Atlantic marsh fiddler crab)
1232:(Salmon & Atsaides, 1968)
1182:(narrow-fronted fiddler crab)
1080:(slender-legged fiddler crab)
906:(lateral-handed fiddler crab)
720:Hombron & Jacquinot, 1846
252:controlled laboratory setting
2666:Movie of two fiddler crabs (
1878:(large Mexican fiddler crab)
1779:(distinguished fiddler crab)
1620:(long-fingered fiddler crab)
1437:Ng, Shih & Christy, 2015
1382:(bonin islands fiddler crab)
1028:(spiny-wristed fiddler crab)
976:(Atlantic sand fiddler crab)
917:(thin-fingered fiddler crab)
764:(orange-clawed fiddler crab)
2473:Raffles Bulletin of Zoology
2019:Paraleptuca chlorophthalmus
1790:(intermediate fiddler crab)
1753:(American Red fiddler crab)
1609:(hairy-handed fiddler crab)
1587:(flame-backed fiddler crab)
1408:(thick-legged fiddler crab)
1387:Paraleptuca chlorophthalmus
1336:(green-banded fiddler crab)
1223:(la herradura fiddler crab)
1054:(Laguna Madre fiddler crab)
542:(East African fiddler crab)
456:(white-handed fiddler crab)
436:(West African fiddler crab)
2971:
2670:) waving the enlarged claw
1788:von Prahl & Toro, 1985
1724:(d'urville's fiddler crab)
1607:(George & Jones, 1982)
1570:(George & Jones, 1982)
1561:(dussumier's fiddler crab)
1505:(belligerent fiddler crab)
1256:(red-jointed fiddler crab)
963:(Novak & Salmon, 1974)
932:(Pacific mud fiddler crab)
851:(Deichmann's fiddler crab)
471:(ring-legged fiddler crab)
66:Red-jointed fiddler crab (
29:
2553:10.3897/zookeys.747.23468
2525:10.11646/zootaxa.2337.1.4
1596:(Adams & White, 1849)
1535:(compressed fiddler crab)
1467:Shih, Chan & Ng, 2018
1397:(green-eyed fiddler crab)
742:(tetragonal fiddler crab)
579:(triangular fiddler crab)
557:(perplexing fiddler crab)
375:for a further two weeks.
184:
77:Scientific classification
64:
55:
1834:(necklaced fiddler crab)
1823:(Brazilian fiddler crab)
1694:(signaling fiddler crab)
1546:(demanding fiddler crab)
1520:(capricorn fiddler crab)
1325:(victorian fiddler crab)
1212:(galápagos fiddler crab)
1147:(Uruguayan fiddler crab)
1017:(brilliant fiddler crab)
1006:(energetic fiddler crab)
965:(gulf sand fiddler crab)
862:(Dorothy's fiddler crab)
726:Gelasimus subeylindricus
686:Gelasimus neocultrimanus
648:(dampier's fiddler crab)
2480:: 1–286. Archived from
1946:Compressed fiddler crab
1646:(polished fiddler crab)
1469:(alcock's fiddler crab)
1419:(splendid fiddler crab)
1234:(longwave gulf fiddler)
1193:(burger's fiddler crab)
1065:(Peruvian fiddler crab)
884:(Heller's fiddler crab)
577:(A.Milne-Edwards, 1873)
222:, well known for their
2955:Arthropod common names
2280:10.3390/biology5010007
1904:(Laguna Madre fiddler)
1811:Dominguez-Alonso, 2008
1801:(greater fiddler crab)
1709:(typhoon fiddler crab)
1683:(shaking fiddler crab)
1598:(forceps fiddler crab)
1572:(elegant fiddler crab)
1376:Paraleptuca boninensis
1300:(mudflat fiddler crab)
1095:(dancing fiddler crab)
943:(musical fiddler crab)
873:(Festa's fiddler crab)
840:(heaping fiddler crab)
825:(Mexican fiddler crab)
814:(painted fiddler crab)
803:(Beebe's fiddler crab)
788:(beating fiddler crab)
753:(calling fiddler crab)
509:(iranian fiddler crab)
498:(cryptic fiddler crab)
378:Fiddler crabs such as
367:
355:
312:
2878:Paleobiology Database
2679:- University of Kyoto
2626:Robert (2019-02-22).
2607:on September 29, 2011
2167:10.1093/biosci/biz011
1893:(styled fiddler crab)
1867:(ornate fiddler crab)
1631:(spined fiddler crab)
1625:Tubuca paradussumieri
1413:Paraleptuca splendida
1402:Paraleptuca crassipes
1267:(biting fiddler crab)
1121:(matted fiddler crab)
1048:Leptuca subcylindrica
1033:Leptuca stenodactylus
895:(uneven fiddler crab)
590:(motley fiddler crab)
573:Austruca triangularis
536:Austruca occidentalis
531:(banana fiddler crab)
487:(citrus fiddler crab)
479:(bengal fiddler crab)
365:
353:
304:
2662:at Wikimedia Commons
1987:Leptuca leptodactyla
1480:(bowed fiddler crab)
1458:(acute fiddler crab)
1245:(olmec fiddler crab)
1228:Minuca longisignalis
1217:Minuca herradurensis
1198:Minuca ecuadoriensis
1141:Leptuca uruguayensis
1085:Leptuca terpsichores
991:(pygmy fiddler crab)
911:Leptuca leptodactyla
808:Leptuca coloradensis
736:Gelasimus tetragonon
568:(indus fiddler crab)
520:(milky fiddler crab)
338:, or other decaying
32:UCA (disambiguation)
2190:Elements of Ecology
1668:(rose fiddler crab)
1614:Tubuca longidigitum
1603:Tubuca hirsutimanus
1287:(S. I. Smith, 1870)
1209:Minuca galapagensis
1171:(clay fiddler crab)
1104:M. J. Rathbun, 1900
954:(aqua fiddler crab)
706:Gelasimus pugilator
696:Gelasimus palustris
676:Gelasimus jocelynae
661:Gelasimus hesperiae
2685:- Fiddlercrab.info
2675:2011-07-22 at the
2403:Functional Ecology
2077:Declawing of crabs
1918:Xeruca formosensis
1763:Martins-Neto, 2001
1651:Tubuca rhizophorae
1510:Tubuca capricornis
1433:Petruca panamensis
1278:(osa fiddler crab)
1161:Minuca argillicola
1070:Leptuca tenuipedis
1041:& Lucas, 1843)
1022:Leptuca spinicarpa
996:Leptuca saltitanta
889:Leptuca inaequalis
845:Leptuca deichmanni
823:(Lockington, 1877)
716:Gelasimus rubripes
638:Gelasimus dampieri
623:Gelasimus borealis
584:Austruca variegata
562:Austruca sindensis
525:Austruca mjoebergi
461:Austruca annulipes
397:Genera and species
381:Austruca mjoebergi
368:
356:
313:
224:sexually dimorphic
2932:
2931:
2865:Open Tree of Life
2701:Taxon identifiers
2668:Uca lactea lactea
2658:Media related to
2632:Fishkeeping World
2199:978-0-321-73607-9
2012:
1923:
1903:
1898:Uca subcylindrica
1892:
1877:
1866:
1856:
1845:
1833:
1822:
1812:
1800:
1789:
1778:
1764:
1752:
1742:
1723:
1708:
1693:
1682:
1667:
1656:
1645:
1630:
1619:
1608:
1597:
1586:
1571:
1560:
1551:Tubuca dussumieri
1545:
1534:
1519:
1504:
1494:
1485:Tubuca australiae
1479:
1468:
1457:
1438:
1418:
1407:
1396:
1381:
1361:
1346:
1335:
1324:
1323:(von Hagen, 1987)
1319:Minuca victoriana
1313:
1299:
1288:
1277:
1266:
1255:
1244:
1239:Minuca marguerita
1233:
1222:
1203:
1192:
1181:
1176:Minuca brevifrons
1170:
1146:
1135:
1126:Leptuca umbratila
1120:
1111:Leptuca tomentosa
1105:
1094:
1079:
1064:
1063:(von Hagen, 1968)
1059:Leptuca tallanica
1053:
1042:
1027:
1016:
1005:
990:
975:
970:Leptuca pugilator
964:
953:
942:
931:
916:
905:
900:Leptuca latimanus
894:
883:
872:
861:
860:(von Hagen, 1968)
856:Leptuca dorotheae
850:
839:
830:Leptuca cumulanta
824:
819:Leptuca crenulata
813:
802:
787:
763:
758:Gelasimus vomeris
752:
741:
731:
721:
711:
701:
691:
681:
670:
647:
632:
609:
589:
578:
567:
556:
547:Austruca perplexa
541:
530:
519:
508:
507:(Pretzmann, 1971)
497:
492:Austruca cryptica
470:
455:
450:Austruca albimana
435:
308:Austruca perplexa
244:shed their shells
204:
203:
198:
192:
50:
16:(Redirected from
2962:
2925:
2924:
2912:
2911:
2899:
2898:
2886:
2885:
2873:
2872:
2860:
2859:
2847:
2846:
2834:
2833:
2821:
2820:
2808:
2807:
2795:
2794:
2782:
2781:
2769:
2768:
2756:
2755:
2743:
2742:
2741:
2728:
2727:
2726:
2696:
2657:
2642:
2641:
2639:
2638:
2623:
2617:
2616:
2614:
2612:
2603:. Archived from
2592:
2583:
2582:
2580:
2578:
2565:
2555:
2535:
2529:
2528:
2508:
2495:
2489:
2488:
2486:
2469:
2460:
2454:
2453:
2437:
2431:
2428:
2422:
2421:
2419:
2389:
2383:
2382:
2380:
2378:
2363:
2357:
2353:
2347:
2344:
2338:
2335:
2329:
2326:
2320:
2317:
2311:
2310:
2300:
2282:
2258:
2252:
2251:
2234:(3): 1015–1022.
2227:Animal Behaviour
2223:
2210:
2204:
2203:
2185:
2179:
2178:
2146:
2140:
2139:193, p. 147-160.
2136:
2130:
2129:
2113:
2104:
2103:
2092:
2038:
2014:
2013:
1991:Margarita Island
1989:in El Guamache,
1983:
1967:
1949:Tubuca coarctata
1943:
1921:
1901:
1886:
1875:
1864:
1854:
1843:
1831:
1820:
1810:
1798:
1787:
1772:
1762:
1759:Uca inaciobritoi
1750:
1747:Uca heteropleura
1740:
1717:
1702:
1691:
1676:
1673:Tubuca seismella
1665:
1654:
1639:
1628:
1618:(Kingsley, 1880)
1617:
1606:
1595:
1592:Tubuca forcipata
1580:
1569:
1554:
1543:
1528:
1525:Tubuca coarctata
1513:
1502:
1488:
1477:
1466:
1456:(Stimpson, 1858)
1455:
1436:
1417:(Stimpson, 1858)
1416:
1405:
1390:
1379:
1355:
1344:
1333:
1322:
1308:
1305:Minuca umbratila
1297:
1286:
1275:
1264:
1254:(Le Conte, 1855)
1253:
1242:
1231:
1220:
1202:(Maccagno, 1928)
1201:
1191:(Holthuis, 1967)
1190:
1180:(Stimpson, 1860)
1179:
1164:
1144:
1129:
1114:
1103:
1088:
1073:
1062:
1052:(Stimpson, 1859)
1051:
1036:
1025:
1014:
1011:Leptuca speciosa
999:
984:
973:
962:
951:
948:Leptuca oerstedi
940:
925:
922:Leptuca limicola
914:
903:
892:
881:
870:
859:
848:
833:
822:
811:
796:
781:
778:Leptuca batuenta
761:
751:(Linnaeus, 1758)
750:
747:Gelasimus vocans
739:
729:
719:
709:
699:
689:
679:
664:
653:Gelasimus excisa
641:
626:
608:(Hoffmann, 1874)
607:
587:
576:
565:
550:
539:
528:
517:
506:
503:Austruca iranica
495:
476:Austruca bengali
464:
454:(Kossmann, 1877)
453:
433:
280:Western Atlantic
256:circadian rhythm
196:
190:
180:Groups included
85:
84:
60:
45:Temporal range:
44:
39:
21:
2970:
2969:
2965:
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2963:
2961:
2960:
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2935:
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2746:
2737:
2736:
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2722:
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2716:
2703:
2677:Wayback Machine
2650:
2645:
2636:
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2610:
2608:
2595:Lianne McLeod.
2594:
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2054:
2039:
2030:
2015:
2002:
1997:
1984:
1975:
1968:
1959:
1947:
1944:
1932:
1922:(Rathbun, 1921)
1889:H.Milne Edwards
1844:Desmarest, 1822
1775:H.Milne Edwards
1720:H.Milne Edwards
1714:Tubuca urvillei
1666:(Tweedie, 1937)
1655:(Tweedie, 1950)
1577:Tubuca flammula
1557:H.Milne Edwards
1544:(Ortmann, 1897)
1531:H.Milne Edwards
1499:Tubuca bellator
1478:(De Haan, 1835)
1393:H.Milne Edwards
1243:(Thurman, 1981)
1187:Minuca burgersi
1100:Leptuca thayeri
1026:(Rathbun, 1900)
981:Leptuca pygmaea
959:Leptuca panacea
952:(Rathbun, 1904)
941:(Rathbun, 1914)
915:(Rathbun, 1898)
904:(Rathbun, 1893)
893:(Rathbun, 1935)
882:(Rathbun, 1902)
878:Leptuca helleri
849:(Rathbun, 1935)
812:(Rathbun, 1893)
762:(McNeill, 1920)
604:Cranuca inversa
553:H.Milne Edwards
529:(Rathbun, 1924)
518:(De Haan, 1835)
514:Austruca lactea
484:Austruca citrus
467:H.Milne Edwards
399:
348:
284:Eastern Pacific
264:
79:
51:
35:
28:
23:
22:
15:
12:
11:
5:
2968:
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2699:
2693:
2692:
2686:
2680:
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2649:
2648:External links
2646:
2644:
2643:
2618:
2584:
2546:(747): 41–62.
2530:
2490:
2487:on 2011-06-06.
2455:
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2060:
2056:
2055:
2040:
2033:
2031:
2016:
2000:
1998:
1985:
1978:
1976:
1972:Afruca tangeri
1969:
1962:
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1945:
1938:
1935:
1934:
1933:
1931:
1928:
1927:
1926:
1925:
1924:
1907:
1906:
1905:
1902:Stimpson, 1862
1894:
1879:
1868:
1857:
1846:
1835:
1828:Uca monilifera
1824:
1821:Latreille 1803
1813:
1802:
1791:
1784:Uca intermedia
1780:
1765:
1754:
1743:
1727:
1726:
1725:
1710:
1699:Tubuca typhoni
1695:
1688:Tubuca signata
1684:
1669:
1658:
1647:
1632:
1621:
1610:
1599:
1588:
1573:
1566:Tubuca elegans
1562:
1547:
1536:
1521:
1506:
1495:
1481:
1474:Tubuca arcuata
1470:
1463:Tubuca alcocki
1459:
1441:
1440:
1439:
1422:
1421:
1420:
1409:
1398:
1383:
1365:
1364:
1363:
1348:
1345:(Herbst, 1804)
1341:Minuca vocator
1337:
1326:
1315:
1301:
1290:
1279:
1268:
1257:
1246:
1235:
1224:
1213:
1205:
1194:
1183:
1172:
1150:
1149:
1148:
1145:(Nobili, 1901)
1137:
1122:
1107:
1096:
1081:
1066:
1055:
1044:
1029:
1018:
1007:
992:
977:
966:
955:
944:
937:Leptuca musica
933:
918:
907:
896:
885:
874:
871:(Nobili, 1902)
867:Leptuca festae
863:
852:
841:
826:
815:
804:
793:Leptuca beebei
789:
767:
766:
765:
754:
743:
740:(Herbst, 1790)
732:
730:Stimpson, 1862
722:
712:
710:Stimpson, 1862
702:
700:Stimpson, 1862
692:
682:
672:
657:
649:
634:
612:
611:
610:
593:
592:
591:
588:(Heller, 1862)
580:
569:
566:(Alcock, 1900)
558:
543:
532:
521:
510:
499:
488:
480:
472:
457:
439:
438:
437:
434:(Eydoux, 1835)
430:Afruca tangeri
398:
395:
347:
344:
292:Algarve region
263:
260:
218:in the family
202:
201:
200:
199:
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182:
181:
177:
176:
171:
167:
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62:
61:
53:
52:
43:
27:Genus of crabs
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2967:
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2697:
2691:- mysabah.com
2690:
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2502:Uca jocelynae
2494:
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2396:Uca mjoebergi
2388:
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2075:
2074:
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2052:
2048:
2047:orientability
2044:
2037:
2032:
2029:
2025:
2021:
2020:
1999:
1996:
1992:
1988:
1982:
1977:
1974:
1973:
1966:
1961:
1958:
1954:
1950:
1942:
1937:
1929:
1920:
1919:
1915:
1914:
1913:
1912:
1908:
1900:
1899:
1895:
1890:
1885:
1884:
1883:Uca stylifera
1880:
1876:(Smith, 1870)
1874:
1873:
1869:
1865:(Smith, 1870)
1863:
1862:
1858:
1855:Rathbun, 1926
1853:
1852:
1847:
1842:
1841:
1836:
1832:Rathbun, 1914
1830:
1829:
1825:
1819:
1818:
1817:Uca maracoani
1814:
1809:
1808:
1803:
1797:
1796:
1792:
1786:
1785:
1781:
1776:
1771:
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1755:
1751:(Smith, 1870)
1749:
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1670:
1664:
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1659:
1653:
1652:
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1643:
1638:
1637:
1636:Tubuca polita
1633:
1627:
1626:
1622:
1616:
1615:
1611:
1605:
1604:
1600:
1594:
1593:
1589:
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1578:
1574:
1568:
1567:
1563:
1558:
1553:
1552:
1548:
1542:
1541:
1540:Tubuca demani
1537:
1532:
1527:
1526:
1522:
1517:
1512:
1511:
1507:
1503:(White, 1847)
1501:
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1414:
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1406:(White, 1847)
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1330:Minuca virens
1327:
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1320:
1316:
1311:
1307:
1306:
1302:
1298:(Smith, 1870)
1296:
1295:
1291:
1285:
1284:
1283:Minuca pugnax
1280:
1274:
1273:
1269:
1265:(Smith, 1870)
1263:
1262:
1261:Minuca mordax
1258:
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232:mangrove crab
229:
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194:
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185:
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160:Superfamily:
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71:
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63:
59:
54:
48:
47:Early Miocene
40:
37:
33:
19:
2708:
2667:
2635:. Retrieved
2631:
2621:
2609:. Retrieved
2605:the original
2575:. Retrieved
2543:
2533:
2516:
2510:
2501:
2493:
2482:the original
2477:
2471:
2458:
2449:
2445:
2435:
2426:
2407:
2401:
2395:
2387:
2377:November 19,
2375:. Retrieved
2361:
2351:
2342:
2333:
2324:
2315:
2270:
2266:
2256:
2231:
2225:
2217:Uca perplexa
2216:
2208:
2189:
2183:
2158:
2154:
2144:
2134:
2125:
2121:
2099:
2090:
2065:
2051:Möbius strip
2041:The obvious
2017:
1986:
1970:
1948:
1916:
1909:
1896:
1881:
1872:Uca princeps
1870:
1859:
1851:Uca oldroydi
1849:
1838:
1826:
1815:
1805:
1799:Herbst, 1782
1793:
1782:
1769:Uca insignis
1767:
1757:
1745:
1735:
1729:
1712:
1697:
1692:(Hess, 1865)
1686:
1671:
1662:Tubuca rosea
1660:
1649:
1634:
1629:(Bott, 1973)
1623:
1612:
1601:
1590:
1575:
1564:
1549:
1538:
1523:
1508:
1497:
1483:
1472:
1461:
1452:Tubuca acuta
1450:
1443:
1431:
1424:
1411:
1400:
1385:
1374:
1367:
1352:Minuca zacae
1350:
1339:
1328:
1317:
1303:
1294:Minuca rapax
1292:
1281:
1270:
1259:
1250:Minuca minax
1248:
1237:
1226:
1221:(Bott, 1954)
1215:
1207:
1196:
1185:
1174:
1159:
1152:
1139:
1124:
1109:
1098:
1083:
1068:
1057:
1046:
1031:
1020:
1015:(Ives, 1891)
1009:
994:
979:
974:(Bosc, 1802)
968:
957:
946:
935:
920:
909:
898:
887:
876:
865:
854:
843:
828:
817:
806:
791:
776:
769:
756:
745:
734:
724:
714:
704:
694:
690:(Bott, 1973)
684:
674:
659:
651:
636:
621:
614:
602:
595:
582:
571:
560:
545:
534:
523:
512:
501:
490:
482:
474:
459:
448:
441:
428:
421:
417:and Ucinae.
410:
406:
400:
391:
379:
377:
369:
357:
321:
314:
306:
288:Indo-Pacific
265:
249:
241:
212:calling crab
211:
208:fiddler crab
207:
205:
150:Infraorder:
124:Malacostraca
68:Minuca minax
67:
42:Fiddler crab
36:
2945:Ocypodoidea
2826:iNaturalist
2733:Wikispecies
2611:January 13,
2053:shown here.
1807:Uca marinae
1741:Brito, 1972
1737:Uca antiqua
1369:Paraleptuca
415:Gelasiminae
324:detritivore
276:West Africa
191:Miers, 1886
188:Gelasiminae
164:Ocypodoidea
144:Pleocyemata
2939:Categories
2637:2021-04-21
2410:(2): 359.
2356:1163-1167.
2155:BioScience
2083:References
1861:Uca ornata
1840:Uca nitida
1272:Minuca osa
403:Ocypodidae
387:signalling
373:planktonic
346:Life cycle
228:ghost crab
220:Ocypodidae
197:Dana, 1851
174:Ocypodidae
140:Suborder:
114:Arthropoda
2950:Asymmetry
2601:About.com
2519:: 47–62.
2289:2079-7737
2175:0006-3568
2062:Captivity
2043:asymmetry
1995:Venezuela
1957:Indonesia
1795:Uca major
616:Gelasimus
268:mangroves
266:Found in
154:Brachyura
100:Kingdom:
94:Eukaryota
2774:BugGuide
2724:Q1329937
2718:Wikidata
2673:Archived
2572:29674902
2307:26797647
2273:(1): 7.
2248:44197123
2071:See also
1951:male in
443:Austruca
340:detritus
332:microbes
296:Portugal
237:wetlands
170:Family:
134:Decapoda
110:Phylum:
104:Animalia
90:Domain:
2917:ZooBank
2818:6586700
2577:3 April
2563:5904522
2544:ZooKeys
2512:Zootaxa
2298:4810164
2267:Biology
2024:Okinawa
1930:Gallery
1891:, 1852)
1777:, 1852)
1722:, 1852)
1707:, 1975)
1681:, 1975)
1644:, 1975)
1585:, 1975)
1559:, 1852)
1533:, 1852)
1518:, 1975)
1493:, 1975)
1426:Petruca
1395:, 1837)
1360:, 1941)
1169:, 1941)
1134:, 1941)
1119:, 1941)
1093:, 1941)
1078:, 1941)
1004:, 1941)
989:, 1941)
930:, 1941)
838:, 1943)
801:, 1941)
786:, 1941)
771:Leptuca
669:, 1975)
646:, 1975)
631:, 1975)
597:Cranuca
555:, 1852)
469:, 1837)
371:remain
272:beaches
262:Ecology
195:Ucinae
130:Order:
120:Class:
49:-recent
2909:106971
2870:289211
2805:1UCASG
2779:354364
2570:
2560:
2305:
2295:
2287:
2246:
2196:
2173:
1911:Xeruca
1445:Tubuca
1312:, 1941
1154:Minuca
423:Afruca
336:fungus
282:, the
278:, the
2904:WoRMS
2891:Plazi
2883:22745
2857:53313
2844:99084
2831:51930
2792:7NXYV
2507:(PDF)
2485:(PDF)
2468:(PDF)
2244:S2CID
2222:(PDF)
2028:Japan
1953:Rinca
1705:Crane
1679:Crane
1642:Crane
1583:Crane
1516:Crane
1491:Crane
1358:Crane
1310:Crane
1167:Crane
1132:Crane
1117:Crane
1091:Crane
1076:Crane
1002:Crane
987:Crane
928:Crane
836:Crane
799:Crane
784:Crane
667:Crane
644:Crane
629:Crane
328:algae
317:chela
250:In a
216:crabs
2852:NCBI
2839:ITIS
2813:GBIF
2800:EPPO
2766:5660
2761:BOLD
2613:2010
2579:2018
2568:PMID
2517:2337
2379:2008
2303:PMID
2285:ISSN
2194:ISBN
2171:ISSN
2128:(6).
2100:PBDB
290:and
230:and
206:The
2787:CoL
2753:Uca
2748:ADW
2739:Uca
2709:Uca
2660:Uca
2558:PMC
2548:doi
2521:doi
2412:doi
2293:PMC
2275:doi
2236:doi
2163:doi
1730:Uca
411:Uca
407:Uca
294:of
274:of
239:.
210:or
18:Uca
2941::
2919::
2906::
2893::
2880::
2867::
2854::
2841::
2828::
2815::
2802::
2789::
2776::
2763::
2750::
2735::
2720::
2630:.
2599:.
2587:^
2566:.
2556:.
2542:.
2515:.
2509:.
2478:17
2476:.
2470:.
2450:64
2448:.
2444:.
2408:23
2406:.
2400:.
2370:.
2301:.
2291:.
2283:.
2269:.
2265:.
2242:.
2232:75
2230:.
2224:.
2169:.
2159:69
2157:.
2153:.
2126:39
2124:.
2120:.
2108:^
2098:.
2026:,
2022:-
1993:,
1955:,
389:.
334:,
330:,
286:,
2640:.
2615:.
2581:.
2550::
2527:.
2523::
2500:"
2452:.
2420:.
2414::
2398:"
2381:.
2309:.
2277::
2271:5
2250:.
2238::
2219:"
2202:.
2177:.
2165::
2102:.
1887:(
1848:†
1837:†
1804:†
1773:(
1756:†
1734:†
1718:(
1703:(
1677:(
1640:(
1581:(
1555:(
1529:(
1514:(
1489:(
1391:(
1356:(
1165:(
1130:(
1115:(
1089:(
1074:(
1037:(
1000:(
985:(
926:(
834:(
797:(
782:(
665:(
642:(
627:(
551:(
465:(
70:)
34:.
20:)
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