57:
33:
226:
have round inguinal glands restricted to the posterior half of the dorsal surface. It is unusual to find more than one species of "clickers" or "squelchers" in the same location, although finding one of each is quite frequent in northern
Australia. The species in this genus show great similarities in
211:. They are native to northern and eastern Australia and southern lowlands of New Guinea. These are small squat frogs, more commonly known as "toadlets". They have glandular skin, often with a pair of raised glands behind each eye, or on the flanks.
214:
They have bumpy, rough skin giving them the appearance of a small toad, hence the name "toadlet", although they are often called "gungans" in
Queensland. There are two distinct types of calls—
227:
body shape and colouration making them difficult to tell apart. Call analysis is often required to confirm identification. This genus is the largest of any in the family
Myobatrachidae.
936:
Catullo, R.A.; Keogh, J.S. (2014). "Aridification drove repeated episodes of diversification between
Australian biomes: Evidence from a multi-locus phylogeny of Australian toadlets (
1101:
1127:
1075:
1206:
1114:
1041:
886:
1036:
279:
1119:
784:
1023:
56:
1201:
1132:
860:
825:
toadlets (Anura: Myobatrachidae) from the western arid zone of
Australia, with a description of a new species"
1196:
477:
462:
236:
U. glandulosa, U. aspera, U. minima, U. trachyderma, U. lithomoda, U. littlejohni, U. altissima, U. mimula,
976:
372:
168:
417:
1158:
750:), a species that the Amphibian Species of the World, following Catullo and Keogh (2014), treats as a
1062:
740:
644:
598:
523:
492:
432:
1191:
751:
678:
613:
156:
51:
723:
40:
1140:
1106:
1049:
957:
708:
402:
1145:
949:
839:
755:
628:
342:
135:
1168:
659:
507:
296:
1014:
222:
have long thin inguinal glands that run along the dorsal surface, while the "squelching"
910:
820:
553:
447:
312:
208:
118:
1185:
583:
568:
387:
327:
693:
538:
1054:
1028:
1088:
1008:
953:
357:
999:
32:
844:
98:
68:
961:
993:
88:
218:
species make either a "click" or a "squelch". Generally, the "clicking"
1080:
1093:
78:
970:
1153:
865:
Amphibian
Species of the World: an Online Reference. Version 6.0
793:
Amphibian
Species of the World: an Online Reference. Version 6.0
204:
108:
974:
819:
Catullo, R.A.; Doughty, P.; Roberts, J.D.; Keogh, J.S. (2011).
1067:
262:
are also squelchers, but are distantly related. The species
248:
U. russelli, U. saxatilis, U. talpa, U. borealis, U. crassa
887:"Uperoleia Gray, 1841 | Amphibian Species of the World"
814:
812:
810:
306:
Davies, Watson, McDonald, Trenerry & Werren, 1993
983:
821:"Multi-locus phylogeny and taxonomic revision of
8:
971:
917:. University of California, Berkeley. 2017
31:
20:
843:
739:Note that the AmphibiaWeb recognizes the
501:Clulow, Anstis, Keogh, and Catullo, 2016
778:
776:
774:
770:
270:are distinct from these other groups.
942:Molecular Phylogenetics and Evolution
748:(Davies, McDonald & Corben, 1986)
264:U. mjobergi, U. micromeles, U. micra,
7:
1159:D15DE063-FFF7-B218-FF47-FBA0FD7F57C4
867:. American Museum of Natural History
795:. American Museum of Natural History
577:Davies, McDonald & Corben, 1986
486:Davies, McDonald & Corben, 1986
396:Davies, McDonald & Corben, 1986
411:Davies, Mahony & Roberts, 1985
256:U. laevigata, U. fusca, U. tyleri,
14:
687:Catullo, Doughty, and Keogh, 2014
717:Tyler, Davies & Martin, 1981
702:Tyler, Davies & Martin, 1981
592:Tyler, Davies & Martin, 1981
562:Tyler, Davies & Martin, 1981
471:Tyler, Davies & Martin, 1981
441:Tyler, Davies & Martin, 1981
366:Tyler, Davies & Martin, 1981
351:Tyler, Davies & Martin, 1981
336:Tyler, Davies & Martin, 1981
321:Tyler, Davies & Martin, 1981
55:
861:"Myobatrachidae Schlegel, 1850"
1207:Taxa named by John Edward Gray
280:Amphibian Species of the World
1:
891:amphibiansoftheworld.amnh.org
732:Davies & Littlejohn, 1986
532:Davies & Littlejohn, 1986
381:Young, Tyler & Kent, 2005
954:10.1016/j.ympev.2014.06.012
1223:
190:Wells and Wellington, 1985
183:Wells and Wellington, 1985
859:Frost, Darrel R. (2016).
845:10.11646/zootaxa.2902.1.1
783:Frost, Darrel R. (2016).
547:Doughty and Roberts, 2008
426:Catullo & Keogh, 2021
162:
155:
148:
143:
52:Scientific classification
50:
39:
30:
23:
510:, yellow-spotted toadlet
282:, there are 29 species:
254:. The eastern species
373:Howard Springs toadlet
714:Uperoleia trachyderma
631:, red-groined toadlet
483:Uperoleia littlejohni
940:: Myobatrachidae)".
745:Uperoleia capitulata
741:small-headed toadlet
619:Uperoleia orientalis
559:Uperoleia micromeles
478:Littlejohn's toadlet
463:Stonemason's toadlet
408:Uperoleia glandulosa
665:Uperoleia saxatilis
514:Uperoleia marmorata
468:Uperoleia lithomoda
453:Uperoleia laevigata
433:Flood plain toadlet
423:Uperoleia gurrumuli
318:Uperoleia arenicola
302:Uperoleia altissima
16:Genus of amphibians
684:Uperoleia stridera
679:Ratcheting toadlet
650:Uperoleia russelli
614:Alexandria toadlet
604:Uperoleia mjobergi
438:Uperoleia inundata
418:Gurrumul's toadlet
378:Uperoleia daviesae
348:Uperoleia borealis
242:. The squelching
1179:
1178:
1141:Open Tree of Life
977:Taxon identifiers
737:
736:
709:Blacksoil toadlet
653:(Loveridge, 1933)
645:Russell's toadlet
638:(Andersson, 1916)
607:(Andersson, 1913)
599:Mjoberg's toadlet
529:Uperoleia martini
498:Uperoleia mahonyi
403:Glandular toadlet
382:
196:
195:
191:
184:
177:
139:
1214:
1202:Amphibian genera
1172:
1171:
1162:
1161:
1149:
1148:
1136:
1135:
1123:
1122:
1110:
1109:
1097:
1096:
1084:
1083:
1071:
1070:
1058:
1057:
1045:
1044:
1032:
1031:
1019:
1018:
1017:
1004:
1003:
1002:
972:
966:
965:
933:
927:
926:
924:
922:
911:"Myobatrachidae"
907:
901:
900:
898:
897:
883:
877:
876:
874:
872:
856:
850:
849:
847:
829:
816:
805:
804:
802:
800:
780:
760:Uperoleia rugosa
756:wrinkled toadlet
749:
733:
729:Uperoleia tyleri
718:
703:
688:
673:
654:
639:
635:Uperoleia rugosa
629:Wrinkled toadlet
623:
608:
593:
589:Uperoleia minima
578:
574:Uperoleia mimula
563:
548:
533:
524:Martin's toadlet
518:
502:
493:Mahony's toadlet
487:
472:
457:
456:Keferstein, 1867
442:
427:
412:
397:
380:
367:
363:Uperoleia crassa
352:
343:Northern toadlet
337:
333:Uperoleia aspera
322:
307:
285:
284:
189:
182:
175:
134:
60:
59:
45:Uperoleia tyleri
35:
21:
1222:
1221:
1217:
1216:
1215:
1213:
1212:
1211:
1182:
1181:
1180:
1175:
1167:
1165:
1157:
1152:
1144:
1139:
1131:
1126:
1118:
1113:
1105:
1100:
1092:
1087:
1079:
1074:
1066:
1061:
1053:
1048:
1040:
1035:
1027:
1022:
1013:
1012:
1007:
998:
997:
992:
979:
969:
935:
934:
930:
920:
918:
909:
908:
904:
895:
893:
885:
884:
880:
870:
868:
858:
857:
853:
827:
818:
817:
808:
798:
796:
782:
781:
772:
768:
747:
731:
724:Tyler's toadlet
716:
701:
699:Uperoleia talpa
686:
667:
660:Pilbara toadlet
652:
637:
621:
606:
591:
576:
561:
546:
544:Uperoleia micra
531:
516:
508:Marbled toadlet
500:
485:
470:
455:
440:
425:
410:
395:
393:Uperoleia fusca
365:
350:
335:
320:
305:
304:
297:Montane toadlet
276:
185:
178:
176:Loveridge, 1933
171:
133:
54:
41:Tyler's toadlet
17:
12:
11:
5:
1220:
1218:
1210:
1209:
1204:
1199:
1197:Myobatrachidae
1194:
1184:
1183:
1177:
1176:
1174:
1173:
1163:
1150:
1137:
1124:
1111:
1098:
1085:
1072:
1059:
1046:
1033:
1020:
1005:
989:
987:
981:
980:
975:
968:
967:
928:
902:
878:
851:
806:
769:
767:
764:
735:
734:
726:
720:
719:
711:
705:
704:
696:
690:
689:
681:
675:
674:
662:
656:
655:
647:
641:
640:
632:
625:
624:
622:(Parker, 1940)
616:
610:
609:
601:
595:
594:
586:
580:
579:
571:
565:
564:
556:
554:Tanami toadlet
550:
549:
541:
535:
534:
526:
520:
519:
511:
504:
503:
495:
489:
488:
480:
474:
473:
465:
459:
458:
450:
448:Smooth toadlet
444:
443:
435:
429:
428:
420:
414:
413:
405:
399:
398:
390:
384:
383:
375:
369:
368:
360:
354:
353:
345:
339:
338:
330:
324:
323:
315:
313:Jabiru toadlet
309:
308:
299:
293:
292:
291:Binomial name
289:
278:Following the
275:
272:
209:Myobatrachidae
207:in the family
203:is a genus of
194:
193:
167:— unjustified
160:
159:
153:
152:
146:
145:
141:
140:
126:
122:
121:
119:Myobatrachidae
116:
112:
111:
106:
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101:
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92:
91:
86:
82:
81:
76:
72:
71:
66:
62:
61:
48:
47:
37:
36:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
1219:
1208:
1205:
1203:
1200:
1198:
1195:
1193:
1190:
1189:
1187:
1170:
1164:
1160:
1155:
1151:
1147:
1142:
1138:
1134:
1129:
1125:
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1112:
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1090:
1086:
1082:
1077:
1073:
1069:
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1038:
1034:
1030:
1025:
1021:
1016:
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1006:
1001:
995:
991:
990:
988:
986:
982:
978:
973:
963:
959:
955:
951:
947:
943:
939:
932:
929:
916:
912:
906:
903:
892:
888:
882:
879:
866:
862:
855:
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846:
841:
837:
833:
826:
824:
815:
813:
811:
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794:
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788:
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775:
771:
765:
763:
761:
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753:
746:
742:
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721:
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627:
626:
620:
617:
615:
612:
611:
605:
602:
600:
597:
596:
590:
587:
585:
584:Small toadlet
582:
581:
575:
572:
570:
569:Mimic toadlet
567:
566:
560:
557:
555:
552:
551:
545:
542:
540:
537:
536:
530:
527:
525:
522:
521:
515:
512:
509:
506:
505:
499:
496:
494:
491:
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484:
481:
479:
476:
475:
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464:
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460:
454:
451:
449:
446:
445:
439:
436:
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431:
430:
424:
421:
419:
416:
415:
409:
406:
404:
401:
400:
394:
391:
389:
388:Dusky toadlet
386:
385:
379:
376:
374:
371:
370:
364:
361:
359:
356:
355:
349:
346:
344:
341:
340:
334:
331:
329:
328:Derby toadlet
326:
325:
319:
316:
314:
311:
310:
303:
300:
298:
295:
294:
290:
287:
286:
283:
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174:
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34:
29:
26:
22:
19:
984:
945:
941:
937:
931:
919:. Retrieved
914:
905:
894:. Retrieved
890:
881:
869:. Retrieved
864:
854:
835:
831:
822:
797:. Retrieved
792:
786:
759:
744:
738:
728:
713:
698:
694:Mole toadlet
683:
669:
664:
649:
634:
618:
603:
588:
573:
558:
543:
539:Tiny toadlet
528:
513:
497:
482:
467:
452:
437:
422:
407:
392:
377:
362:
347:
332:
317:
301:
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149:
129:
128:
44:
24:
18:
1089:iNaturalist
1009:Wikispecies
948:: 106–117.
915:AmphibiaWeb
789:Gray, 1841"
358:Fat toadlet
288:Common name
268:U. daviesae
252:U. inundata
1186:Categories
921:6 February
896:2022-04-26
871:6 February
799:6 February
766:References
517:Gray, 1841
260:U. martini
169:emendation
150:See text.
1192:Uperoleia
1042:Uperoleia
1029:Uperoleia
1015:Uperoleia
985:Uperoleia
938:Uperoleia
823:Uperoleia
787:Uperoleia
244:Uperoleia
240:U. rugosa
232:Uperoleia
230:Clicking
224:Uperoleia
220:Uperoleia
216:Uperoleia
200:Uperoleia
187:Prohartia
173:Glauertia
165:Hyperolia
130:Uperoleia
75:Kingdom:
69:Eukaryota
25:Uperoleia
1000:Q2452924
994:Wikidata
962:24971737
838:: 1–43.
668:Catullo
246:include
234:include
180:Hosmeria
157:Synonyms
144:Species
115:Family:
99:Amphibia
89:Chordata
85:Phylum:
79:Animalia
65:Domain:
1107:1022954
1081:2424625
832:Zootaxa
754:of the
752:synonym
274:Species
125:Genus:
105:Order:
95:Class:
1166:uBio:
1146:940195
1120:208114
960:
672:, 2011
670:et al.
250:, and
138:, 1841
1169:31592
1154:Plazi
1102:IRMNG
1094:25263
1068:42685
828:(PDF)
205:frogs
109:Anura
1133:8366
1128:NCBI
1115:ITIS
1076:GBIF
1055:84ZK
958:PMID
923:2017
873:2017
836:2902
801:2017
266:and
258:and
238:and
136:Gray
1063:EoL
1050:CoL
1037:ASW
1024:AFD
950:doi
840:doi
762:).
1188::
1156::
1143::
1130::
1117::
1104::
1091::
1078::
1065::
1052::
1039::
1026::
1011::
996::
956:.
946:79
944:.
913:.
889:.
863:.
834:.
830:.
809:^
791:.
773:^
43:,
964:.
952::
925:.
899:.
875:.
848:.
842::
803:.
785:"
758:(
743:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
