31:
51:
2467:
269:
lacks ocelli), are very close to each other. They show a wide separation between the hind and mid coxae on the underside. Other important family characters are the presence of a structure for stridulation, the stridulitrum on the first anal vein of the hind wing, and bristles on the claws. The female
264:
This family has about 100 species, with distribution limited mainly to eastern Asia. Urostylidids are 4 to 15 mm and somewhat longer than broad with elongated legs, three tarsal segments, and a small head. The antenna has 5 segments of which the first is longer than the head. The base of the
290:. Thus the phylogeny of the group is unclear but lies within the Pentatomoidea. In the past the family was divided into two subfamilies, but one group, the Saileriolinae, has since been suggested as a distinct family,
482:
Yao, Yunzhi; Cai, Wanzhi; Rider, David A.; Ren, Dong (2012). "Primipentatomidae fam. nov. (Hemiptera: Heteroptera: Pentatomomorpha), an extinct insect family from the
Cretaceous of north-eastern China".
2400:
2413:
270:
genital structure has nine gonocoxites that form a M or W-shaped sclerite. They suck plant sap. The male reproductive anatomy suggests close affinities to the
2508:
2374:
535:
423:"Revision of the Urostylidae of Taiwan, with descriptions of three new species and one new record (Hemiptera-Heteroptera: Urostylidae)"
312:"Removal of homonymy between Urostylidae Dallas, 1851 (Insecta, Heteroptera) and Urostylidae Buetschli, 1889 (Ciliophora, Hypotrichia)"
2501:
358:"Phylogenetic relationships of family groups in Pentatomoidea based on morphology and DNA sequences (Insecta: Heteroptera)"
2532:
2537:
357:
50:
422:
2494:
2348:
2366:
2418:
245:
or "primitive" family within the stink-bug lineage. They are found only in Asia. Older works used the spelling
137:
528:
2291:
2353:
2439:
250:
242:
2527:
311:
164:
1433:
294:, by at least two studies. The position of Saileriolidae within the Pentatomoidea is still unclear.
2154:
1895:
1655:
1480:
447:
Kumar, R. (1971). "Morphology and relationships of the
Pentatomoidea (Heteroptera) 5-Urostylidae".
2164:
1853:
1827:
265:
antenna has broad tubercles and has a ringed appearance. The simple eyes or ocelli, when present (
184:
174:
154:
2329:
1454:
1331:
1322:
521:
500:
464:
388:
45:
2444:
224:
30:
337:
204:
194:
2426:
1114:
1074:
836:
380:
254:
214:
2478:
2405:
2431:
2197:
2139:
2134:
1968:
1879:
1642:
1420:
1412:
1339:
1122:
984:
492:
456:
372:
2192:
2159:
2098:
2002:
1997:
1934:
1909:
1650:
1503:
1344:
1308:
1298:
1280:
1216:
871:
604:
2361:
2218:
1976:
1858:
1621:
1592:
1394:
1303:
1288:
1159:
1146:
1053:
1024:
904:
808:
692:
651:
271:
249:
but this clashes with the name used for a protozoan family and a spelling correction (
2521:
2474:
2266:
2254:
2226:
2085:
2025:
2007:
1947:
1832:
1817:
1715:
1464:
1449:
1441:
1175:
1098:
1090:
1000:
876:
813:
778:
739:
726:
661:
641:
376:
291:
275:
122:
504:
392:
2187:
2038:
1863:
1822:
1784:
1767:
1753:
1741:
1694:
1686:
1532:
1519:
1485:
1293:
1260:
1206:
1082:
1045:
1008:
963:
788:
713:
705:
616:
588:
287:
408:
True Bugs of the World (Hemiptera:Heteroptera): Classification and
Natural History
496:
2387:
2323:
2262:
2205:
2116:
2051:
2033:
2015:
1984:
1812:
1613:
1361:
1253:
1211:
1201:
1196:
1106:
1037:
1029:
992:
861:
856:
798:
770:
753:
279:
112:
2314:
2149:
2108:
2090:
1871:
1776:
1673:
1665:
1629:
1546:
1188:
1061:
979:
866:
851:
846:
823:
731:
718:
656:
646:
625:
2169:
2129:
2121:
2061:
2046:
2020:
1918:
1792:
1571:
1563:
1527:
1378:
1229:
1221:
1167:
1066:
953:
783:
576:
564:
545:
513:
102:
82:
62:
384:
2340:
2308:
2234:
2177:
2077:
2069:
1989:
1926:
1887:
1800:
1707:
1584:
1511:
1399:
1386:
1138:
1130:
1016:
924:
916:
831:
818:
803:
793:
582:
283:
282:
has two pump flanges in two genera which is a feature found also in the
2379:
2182:
2144:
1845:
1837:
1720:
1600:
841:
468:
2466:
2392:
687:
671:
570:
558:
92:
72:
2285:
460:
2258:
937:
2289:
1250:
901:
601:
517:
253:) has been suggested that also avoids the confusion created by
356:
Grazia, Jocelia; Schuh, Randall T.; Wheeler, Ward C. (2008).
2482:
1085:(cottony cushion scales, giant coccids, ground pearls)
2265:. Heteroptera with Coleorrhyncha were referred to as
2298:
2217:
2107:
2060:
1946:
1917:
1908:
1775:
1766:
1740:
1706:
1685:
1664:
1641:
1612:
1583:
1562:
1545:
1502:
1463:
1432:
1411:
1377:
1360:
1330:
1321:
1279:
1187:
1158:
962:
936:
915:
769:
752:
704:
670:
624:
615:
406:Schuh, Randall T.; Slater, James Alexander (1995).
261:has also been used for some members in the past.
1874:(assassin bugs, wheel bugs, thread-legged bugs)
310:Berger, H.; Heiss, E.; Kerzhner, I.M. (2001).
2502:
529:
410:. Cornell University Press. pp. 245–246.
319:Annalen des Naturhistorischen Museums in Wien
241:is a family of true bugs and is considered a
8:
339:Fauna of British India. Rhynchota. Volume 1
2509:
2495:
2286:
1914:
1772:
1737:
1559:
1499:
1374:
1327:
1276:
1272:
1247:
912:
898:
766:
621:
612:
598:
536:
522:
514:
342:. London: Taylor and Francis. p. 303.
29:
20:
421:Ren, Shu-Zhi; Lin, Cheng-Shing (2003).
302:
351:
349:
7:
2463:
2461:
2028:(jewel bugs or metallic shield bugs)
1840:(plant bugs, leaf bugs, grass bugs)
1787:(minute pirate bugs or flower bugs)
1444:(marsh treaders or water measurers)
485:Journal of Systematic Palaeontology
2481:. You can help Knowledge (XXG) by
14:
1979:(giant shield bugs and relatives)
734:(typical treehoppers, thorn bugs)
2465:
377:10.1111/j.1096-0031.2008.00224.x
49:
2172:(milkweed bugs, true seed bugs)
2080:(squash bugs, leaf-footed bugs)
1347:(unique-headed bugs, gnat bugs)
1603:(water scorpions, needle bugs)
1:
497:10.1080/14772019.2011.639814
2229:(red bugs, cotton stainers)
449:American Midland Naturalist
2554:
2460:
2250:
1736:
1498:
1275:
1271:
1246:
1056:(pupillarial palm scales)
911:
897:
611:
597:
553:
150:
145:
46:Scientific classification
44:
37:
28:
23:
1170:(woolly conifer aphids)
2093:(scentless plant bugs)
1093:(Mediterranean scales)
2440:Paleobiology Database
2237:(bordered plant bugs)
2010:(includes ebony bugs)
1676:(creeping water bugs)
430:Formosan Entomologist
39:Urochela quadrinotata
2533:Heteroptera families
2208:(ash-grey leaf bugs)
2200:(atypical seed bugs)
1866:(old world bat bugs)
1795:(bed bugs, bat bugs)
1723:(pygmy backswimmers)
1632:(velvety shore bugs)
1224:(jumping plant lice)
336:Distant, WL (1902).
2538:Pentatomoidea stubs
2072:(broad-headed bugs)
1423:(velvet water bugs)
1011:(ornate pit scales)
16:Family of true bugs
1595:(giant water bugs)
1522:(spiny shore bugs)
1455:Paraphrynoveliidae
1368:(semiaquatic bugs)
1323:Enicocephalomorpha
1117:(phenacoleachiids)
1115:Phenacoleachiidae
1077:(false pit scales)
1075:Lecanodiaspididae
257:. The family name
2490:
2489:
2455:
2454:
2427:Open Tree of Life
2292:Taxon identifiers
2283:
2282:
2279:
2278:
2275:
2274:
2270:
2246:
2245:
2238:
2230:
2209:
2201:
2198:Rhyparochromidae
2173:
2125:
2094:
2081:
2073:
2042:
2029:
2011:
1993:
1980:
1972:
1969:Acanthosomatidae
1956:
1938:
1930:
1904:
1903:
1891:
1883:
1882:(royal palm bugs)
1880:Thaumastocoridae
1875:
1867:
1849:
1841:
1808:
1796:
1788:
1762:
1761:
1732:
1731:
1724:
1698:
1677:
1633:
1625:
1604:
1596:
1575:
1555:
1553:(true water bugs)
1541:
1540:
1523:
1515:
1494:
1493:
1473:
1445:
1424:
1403:
1390:
1370:
1356:
1355:
1348:
1317:
1316:
1266:
1242:
1241:
1238:
1237:
1225:
1179:
1171:
1150:
1142:
1141:(giant mealybugs)
1134:
1126:
1123:Phoenicococcidae
1118:
1110:
1102:
1094:
1086:
1078:
1070:
1057:
1049:
1041:
1033:
1020:
1012:
1004:
996:
988:
985:Asterolecaniidae
972:
946:
928:
893:
892:
889:
888:
885:
884:
837:Hypochthonellidae
827:
762:
748:
747:
735:
722:
696:
680:
634:
236:
235:
141:
2545:
2511:
2504:
2497:
2469:
2462:
2448:
2447:
2435:
2434:
2422:
2421:
2409:
2408:
2396:
2395:
2383:
2382:
2370:
2369:
2357:
2356:
2344:
2343:
2334:
2333:
2332:
2319:
2318:
2317:
2287:
2257:are a different
2252:
2236:
2228:
2207:
2199:
2171:
2140:Cryptorhamphidae
2135:Colobathristidae
2123:
2092:
2079:
2071:
2040:
2027:
2009:
1992:(burrowing bugs)
1991:
1978:
1970:
1957:
1955:
1952:
1936:
1935:Termitaphididae
1928:
1915:
1889:
1881:
1873:
1865:
1847:
1839:
1805:Curalium cronini
1802:
1794:
1786:
1773:
1746:
1738:
1722:
1696:
1675:
1643:Aphelocheiroidea
1631:
1623:
1602:
1594:
1573:
1560:
1556:
1554:
1551:
1521:
1513:
1500:
1474:
1472:
1471:(water treaders)
1469:
1443:
1422:
1401:
1389:(water striders)
1388:
1375:
1371:
1369:
1366:
1346:
1345:Enicocephalidae
1340:Aenictopecheidae
1332:Enicocephaloidea
1328:
1277:
1273:
1265:
1259:
1256:
1248:
1223:
1177:
1169:
1148:
1140:
1132:
1124:
1116:
1108:
1100:
1092:
1084:
1076:
1068:
1055:
1047:
1040:(armored scales)
1039:
1031:
1018:
1010:
1002:
994:
986:
973:
971:
968:
947:
945:
942:
926:
913:
899:
825:
767:
763:
761:
758:
733:
720:
694:
681:
679:
676:
635:
633:
630:
622:
613:
599:
538:
531:
524:
515:
509:
508:
479:
473:
472:
444:
438:
437:
427:
418:
412:
411:
403:
397:
396:
362:
353:
344:
343:
333:
327:
326:
316:
307:
230:
220:
210:
200:
190:
180:
170:
160:
136:
54:
53:
33:
21:
2553:
2552:
2548:
2547:
2546:
2544:
2543:
2542:
2518:
2517:
2516:
2515:
2458:
2456:
2451:
2443:
2438:
2430:
2425:
2417:
2412:
2404:
2399:
2391:
2386:
2378:
2373:
2365:
2360:
2352:
2347:
2339:
2337:
2328:
2327:
2322:
2313:
2312:
2307:
2294:
2284:
2271:
2242:
2213:
2193:Pachygronthidae
2160:Heterogastridae
2103:
2099:Stenocephalidae
2056:
2003:Parastrachiidae
1998:Thaumastellidae
1977:Tessaratomidae
1953:
1951:
1950:
1942:
1910:Pentatomomorpha
1900:
1758:
1745:(Coleorrhyncha)
1744:
1743:
1742:Peloridiomorpha
1728:
1702:
1681:
1660:
1651:Aphelocheiridae
1637:
1622:Gelastocoridae
1608:
1593:Belostomatidae
1579:
1574:(water boatmen)
1552:
1550:
1549:
1537:
1504:Leptopodomorpha
1490:
1470:
1468:
1467:
1459:
1428:
1407:
1367:
1365:
1364:
1352:
1313:
1309:Stemmocryptidae
1299:Hypsipterygidae
1281:Dipsocoromorpha
1267:
1263:
1257:
1234:
1217:Phacopteronidae
1183:
1154:
1149:(stictococcids)
1147:Stictococcidae
1131:Pseudococcidae
1109:(ensign scales)
1054:Halimococcidae
970:(scale insects)
969:
967:
966:
958:
943:
941:
940:
932:
907:
881:
872:Tettigometridae
759:
757:
756:
744:
700:
695:(hairy cicadas)
693:Tettigarctidae
677:
675:
674:
666:
631:
629:
628:
607:
605:Auchenorrhyncha
593:
549:
542:
512:
481:
480:
476:
461:10.2307/2423912
446:
445:
441:
425:
420:
419:
415:
405:
404:
400:
360:
355:
354:
347:
335:
334:
330:
314:
309:
308:
304:
300:
228:
218:
208:
198:
188:
178:
168:
165:Eurhynchiocoris
158:
135:
48:
17:
12:
11:
5:
2551:
2549:
2541:
2540:
2535:
2530:
2520:
2519:
2514:
2513:
2506:
2499:
2491:
2488:
2487:
2470:
2453:
2452:
2450:
2449:
2436:
2423:
2410:
2397:
2384:
2371:
2362:Fauna Europaea
2358:
2345:
2335:
2320:
2304:
2302:
2296:
2295:
2290:
2281:
2280:
2277:
2276:
2273:
2272:
2251:
2248:
2247:
2244:
2243:
2241:
2240:
2232:
2227:Pyrrhocoridae
2223:
2221:
2219:Pyrrhocoroidea
2215:
2214:
2212:
2211:
2203:
2195:
2190:
2185:
2180:
2175:
2167:
2162:
2157:
2152:
2147:
2142:
2137:
2132:
2127:
2119:
2113:
2111:
2105:
2104:
2102:
2101:
2096:
2088:
2083:
2075:
2066:
2064:
2058:
2057:
2055:
2054:
2049:
2044:
2036:
2031:
2026:Scutelleridae
2023:
2018:
2013:
2008:Thyreocoridae
2005:
2000:
1995:
1987:
1982:
1974:
1966:
1960:
1958:
1944:
1943:
1941:
1940:
1937:(termite bugs)
1932:
1923:
1921:
1912:
1906:
1905:
1902:
1901:
1899:
1898:
1893:
1885:
1877:
1869:
1861:
1859:Plokiophilidae
1856:
1851:
1843:
1835:
1830:
1825:
1820:
1815:
1810:
1798:
1790:
1781:
1779:
1770:
1764:
1763:
1760:
1759:
1757:
1756:
1750:
1748:
1734:
1733:
1730:
1729:
1727:
1726:
1718:
1712:
1710:
1704:
1703:
1701:
1700:
1697:(backswimmers)
1691:
1689:
1683:
1682:
1680:
1679:
1670:
1668:
1662:
1661:
1659:
1658:
1653:
1647:
1645:
1639:
1638:
1636:
1635:
1627:
1618:
1616:
1610:
1609:
1607:
1606:
1598:
1589:
1587:
1581:
1580:
1578:
1577:
1568:
1566:
1557:
1543:
1542:
1539:
1538:
1536:
1535:
1530:
1525:
1517:
1508:
1506:
1496:
1495:
1492:
1491:
1489:
1488:
1483:
1477:
1475:
1461:
1460:
1458:
1457:
1452:
1447:
1442:Hydrometridae
1438:
1436:
1434:Hydrometroidea
1430:
1429:
1427:
1426:
1417:
1415:
1409:
1408:
1406:
1405:
1397:
1395:Hermatobatidae
1392:
1383:
1381:
1372:
1358:
1357:
1354:
1353:
1351:
1350:
1342:
1336:
1334:
1325:
1319:
1318:
1315:
1314:
1312:
1311:
1306:
1304:Schizopteridae
1301:
1296:
1291:
1289:Ceratocombidae
1285:
1283:
1269:
1268:
1251:
1244:
1243:
1240:
1239:
1236:
1235:
1233:
1232:
1227:
1219:
1214:
1209:
1204:
1199:
1193:
1191:
1185:
1184:
1182:
1181:
1178:(phylloxerans)
1176:Phylloxeridae
1173:
1164:
1162:
1160:Phylloxeroidea
1156:
1155:
1153:
1152:
1144:
1136:
1128:
1120:
1112:
1104:
1101:(giant scales)
1099:Monophlebidae
1096:
1091:Micrococcidae
1088:
1080:
1072:
1064:
1059:
1051:
1043:
1035:
1030:Dactylopiidae
1027:
1025:Conchaspididae
1022:
1014:
1006:
1003:(carayonemids)
1001:Carayonemidae
998:
990:
982:
976:
974:
960:
959:
957:
956:
950:
948:
934:
933:
931:
930:
921:
919:
909:
908:
905:Sternorrhyncha
902:
895:
894:
891:
890:
887:
886:
883:
882:
880:
879:
874:
869:
864:
859:
854:
849:
844:
839:
834:
829:
826:(lanternflies)
821:
816:
811:
809:Dictyopharidae
806:
801:
796:
791:
786:
781:
775:
773:
764:
760:(planthoppers)
750:
749:
746:
745:
743:
742:
737:
729:
724:
716:
710:
708:
702:
701:
699:
698:
690:
684:
682:
668:
667:
665:
664:
659:
654:
652:Clastopteridae
649:
644:
638:
636:
619:
609:
608:
602:
595:
594:
592:
591:
585:
579:
573:
567:
561:
554:
551:
550:
543:
541:
540:
533:
526:
518:
511:
510:
474:
439:
413:
398:
371:(6): 932–976.
345:
328:
301:
299:
296:
272:Tessaratomidae
234:
233:
232:
231:
221:
211:
209:Westwood, 1837
201:
199:Westwood, 1837
191:
181:
179:Kirkaldy, 1908
171:
161:
148:
147:
143:
142:
130:
126:
125:
120:
116:
115:
110:
106:
105:
100:
96:
95:
90:
86:
85:
80:
76:
75:
70:
66:
65:
60:
56:
55:
42:
41:
35:
34:
26:
25:
24:Urostylididae
15:
13:
10:
9:
6:
4:
3:
2:
2550:
2539:
2536:
2534:
2531:
2529:
2526:
2525:
2523:
2512:
2507:
2505:
2500:
2498:
2493:
2492:
2486:
2484:
2480:
2477:article is a
2476:
2475:Pentatomoidea
2471:
2468:
2464:
2459:
2446:
2441:
2437:
2433:
2428:
2424:
2420:
2415:
2411:
2407:
2402:
2398:
2394:
2389:
2385:
2381:
2376:
2372:
2368:
2363:
2359:
2355:
2350:
2346:
2342:
2336:
2331:
2330:Urostylididae
2325:
2321:
2316:
2310:
2306:
2305:
2303:
2301:
2300:Urostylididae
2297:
2293:
2288:
2268:
2267:Prosorrhyncha
2264:
2260:
2256:
2255:Coleorrhyncha
2249:
2239:
2233:
2231:
2225:
2224:
2222:
2220:
2216:
2210:
2204:
2202:
2196:
2194:
2191:
2189:
2186:
2184:
2181:
2179:
2176:
2174:
2168:
2166:
2163:
2161:
2158:
2156:
2155:Henicocoridae
2153:
2151:
2148:
2146:
2143:
2141:
2138:
2136:
2133:
2131:
2128:
2126:
2120:
2118:
2115:
2114:
2112:
2110:
2106:
2100:
2097:
2095:
2089:
2087:
2086:Hyocephalidae
2084:
2082:
2076:
2074:
2068:
2067:
2065:
2063:
2059:
2053:
2050:
2048:
2045:
2043:
2039:Pentatomidae
2037:
2035:
2032:
2030:
2024:
2022:
2019:
2017:
2014:
2012:
2006:
2004:
2001:
1999:
1996:
1994:
1988:
1986:
1983:
1981:
1975:
1973:
1971:(shield bugs)
1967:
1965:
1964:Urostylididae
1962:
1961:
1959:
1954:(shield bugs)
1949:
1948:Pentatomoidea
1945:
1939:
1933:
1931:
1925:
1924:
1922:
1920:
1916:
1913:
1911:
1907:
1897:
1896:Velocipedidae
1894:
1892:
1886:
1884:
1878:
1876:
1870:
1868:
1864:Polyctenidae
1862:
1860:
1857:
1855:
1852:
1850:
1848:(damsel bugs)
1844:
1842:
1836:
1834:
1833:Microphysidae
1831:
1829:
1826:
1824:
1821:
1819:
1818:Lasiochilidae
1816:
1814:
1811:
1809:
1806:
1799:
1797:
1791:
1789:
1785:Anthocoridae
1783:
1782:
1780:
1778:
1774:
1771:
1769:
1765:
1755:
1752:
1751:
1749:
1747:
1739:
1735:
1725:
1719:
1717:
1716:Helotrephidae
1714:
1713:
1711:
1709:
1705:
1699:
1695:Notonectidae
1693:
1692:
1690:
1688:
1687:Notonectoidea
1684:
1678:
1672:
1671:
1669:
1667:
1663:
1657:
1656:Potamocoridae
1654:
1652:
1649:
1648:
1646:
1644:
1640:
1634:
1628:
1626:
1620:
1619:
1617:
1615:
1611:
1605:
1599:
1597:
1591:
1590:
1588:
1586:
1582:
1576:
1570:
1569:
1567:
1565:
1561:
1558:
1548:
1544:
1534:
1531:
1529:
1526:
1524:
1520:Leptopodidae
1518:
1516:
1510:
1509:
1507:
1505:
1501:
1497:
1487:
1484:
1482:
1481:Madeoveliidae
1479:
1478:
1476:
1466:
1465:Mesovelioidea
1462:
1456:
1453:
1451:
1450:Macroveliidae
1448:
1446:
1440:
1439:
1437:
1435:
1431:
1425:
1419:
1418:
1416:
1414:
1410:
1404:
1402:(riffle bugs)
1398:
1396:
1393:
1391:
1385:
1384:
1382:
1380:
1376:
1373:
1363:
1359:
1349:
1343:
1341:
1338:
1337:
1335:
1333:
1329:
1326:
1324:
1320:
1310:
1307:
1305:
1302:
1300:
1297:
1295:
1292:
1290:
1287:
1286:
1284:
1282:
1278:
1274:
1270:
1262:
1261:Coleorrhyncha
1255:
1249:
1245:
1231:
1228:
1226:
1220:
1218:
1215:
1213:
1210:
1208:
1205:
1203:
1200:
1198:
1195:
1194:
1192:
1190:
1186:
1180:
1174:
1172:
1166:
1165:
1163:
1161:
1157:
1151:
1145:
1143:
1137:
1135:
1129:
1127:
1125:(palm scales)
1121:
1119:
1113:
1111:
1105:
1103:
1097:
1095:
1089:
1087:
1083:Margarodidae
1081:
1079:
1073:
1071:
1065:
1063:
1060:
1058:
1052:
1050:
1048:(felt scales)
1046:Eriococcidae
1044:
1042:
1036:
1034:
1028:
1026:
1023:
1021:
1019:(soft scales)
1015:
1013:
1009:Cerococcidae
1007:
1005:
999:
997:
991:
989:
983:
981:
978:
977:
975:
965:
961:
955:
952:
951:
949:
939:
935:
929:
923:
922:
920:
918:
914:
910:
906:
900:
896:
878:
877:Tropiduchidae
875:
873:
870:
868:
865:
863:
860:
858:
855:
853:
850:
848:
845:
843:
840:
838:
835:
833:
830:
828:
822:
820:
817:
815:
814:Eurybrachidae
812:
810:
807:
805:
802:
800:
797:
795:
792:
790:
787:
785:
782:
780:
779:Acanaloniidae
777:
776:
774:
772:
768:
765:
755:
754:Fulgoromorpha
751:
741:
740:Myerslopiidae
738:
736:
730:
728:
727:Melizoderidae
725:
723:
721:(leafhoppers)
719:Cicadellidae
717:
715:
712:
711:
709:
707:
703:
697:
691:
689:
686:
685:
683:
673:
669:
663:
662:Machaerotidae
660:
658:
655:
653:
650:
648:
645:
643:
642:Aphrophoridae
640:
639:
637:
632:(froghoppers)
627:
623:
620:
618:
614:
610:
606:
600:
596:
590:
586:
584:
580:
578:
574:
572:
568:
566:
562:
560:
556:
555:
552:
547:
539:
534:
532:
527:
525:
520:
519:
516:
506:
502:
498:
494:
490:
486:
478:
475:
470:
466:
462:
458:
454:
450:
443:
440:
435:
431:
424:
417:
414:
409:
402:
399:
394:
390:
386:
382:
378:
374:
370:
366:
359:
352:
350:
346:
341:
340:
332:
329:
324:
320:
313:
306:
303:
297:
295:
293:
292:Saileriolidae
289:
285:
281:
277:
276:Natalicolinae
273:
268:
262:
260:
256:
252:
248:
244:
240:
239:Urostylididae
227:
226:
222:
217:
216:
212:
207:
206:
202:
197:
196:
192:
187:
186:
182:
177:
176:
172:
167:
166:
162:
157:
156:
152:
151:
149:
144:
139:
134:
133:Urostylididae
131:
128:
127:
124:
123:Pentatomoidea
121:
119:Superfamily:
118:
117:
114:
111:
108:
107:
104:
101:
98:
97:
94:
91:
88:
87:
84:
81:
78:
77:
74:
71:
68:
67:
64:
61:
58:
57:
52:
47:
43:
40:
36:
32:
27:
22:
19:
2483:expanding it
2472:
2457:
2299:
2206:Piesmatidae
2188:Oxycarenidae
2165:Idiostolidae
2124:(stilt bugs)
2041:(stink bugs)
1963:
1854:Pachynomidae
1828:Medocostidae
1823:Lyctocoridae
1804:
1768:Cimicomorpha
1754:Peloridiidae
1533:Aepophilidae
1514:(shore bugs)
1486:Mesoveliidae
1294:Dipsocoridae
1207:Carsidaridae
1107:Ortheziidae
1069:(lac scales)
1038:Diaspididae
1032:(cochineals)
995:(beesoniids)
993:Beesoniidae
987:(pit scales)
927:(whiteflies)
925:Aleyrodidae
917:Aleyrodoidea
789:Achilixiidae
732:Membracidae
714:Aetalionidae
706:Membracoidea
617:Cicadomorpha
589:Paraneoptera
587:Superorder:
581:Infraclass:
488:
484:
477:
455:(1): 63–73.
452:
448:
442:
433:
429:
416:
407:
401:
368:
364:
338:
331:
322:
318:
305:
288:Leptopodidae
266:
263:
259:Urolabididae
258:
246:
238:
237:
223:
219:Dallas, 1850
213:
203:
193:
185:Urochelellus
183:
175:Tessaromerus
173:
169:Reuter, 1881
163:
155:Cobbenicoris
153:
132:
38:
18:
2528:Shield bugs
2388:iNaturalist
2324:Wikispecies
2263:Heteroptera
2117:Artheneidae
2091:Rhopalidae
2052:Megarididae
2034:Plataspidae
2016:Lestoniidae
1985:Dinidoridae
1929:(flat bugs)
1890:(lace bugs)
1872:Reduviidae
1813:Joppeicidae
1801:Curaliidae
1674:Naucoridae
1666:Naucoroidea
1630:Ochteridae
1624:(toad bugs)
1614:Ochteroidea
1362:Gerromorpha
1254:Heteroptera
1212:Homotomidae
1202:Calophyidae
1197:Aphalaridae
1133:(mealybugs)
862:Nogodinidae
857:Meenoplidae
824:Fulgoridae
799:Delphacidae
771:Fulgoroidea
626:Cercopoidea
280:spermatheca
247:Urostylidae
113:Heteroptera
2522:Categories
2170:Lygaeidae
2150:Geocoridae
2122:Berytidae
2109:Lygaeoidea
1793:Cimicidae
1777:Cimicoidea
1572:Corixidae
1564:Corixoidea
1547:Nepomorpha
1222:Psyllidae
1189:Psylloidea
1168:Adelgidae
1067:Kerriidae
1062:Kermesidae
980:Aclerdidae
938:Aphidoidea
867:Ricaniidae
852:Lophopidae
847:Kinnaridae
672:Cicadoidea
657:Epipygidae
647:Cercopidae
575:Subclass:
565:Arthropoda
436:: 129–143.
365:Cladistics
325:: 301–302.
298:References
274:subfamily
251:emendation
225:Yangicoris
189:Yang, 1937
109:Suborder:
83:Arthropoda
2235:Largidae
2130:Blissidae
2078:Coreidae
2070:Alydidae
2062:Coreoidea
2047:Canopidae
2021:Phloeidae
1990:Cydnidae
1927:Aradidae
1919:Aradoidea
1888:Tingidae
1528:Omaniidae
1512:Saldidae
1421:Hebridae
1413:Hebroidea
1400:Veliidae
1387:Gerridae
1379:Gerroidea
1252:Suborder
1230:Triozidae
1139:Putoidae
1017:Coccidae
964:Coccoidea
954:Aphididae
903:Suborder
784:Achilidae
688:Cicadidae
678:(cicadas)
603:Suborder
577:Pterygota
557:Kingdom:
546:Hemiptera
267:Urolabida
229:Ren, 1998
205:Urostylis
195:Urolabida
159:Ren, 1998
103:Hemiptera
69:Kingdom:
63:Eukaryota
2338:BioLib:
2315:Q4006498
2309:Wikidata
2178:Malcidae
1846:Nabidae
1838:Miridae
1721:Pleidae
1708:Pleoidea
1601:Nepidae
1585:Nepoidea
944:(aphids)
832:Gengidae
819:Flatidae
804:Derbidae
794:Cixiidae
583:Neoptera
563:Phylum:
559:Animalia
548:families
505:86753502
491:: 1–20.
393:41951432
385:34892882
284:Aradidae
255:homonyms
215:Urochela
129:Family:
79:Phylum:
73:Animalia
59:Domain:
2380:5431354
2183:Ninidae
2145:Cymidae
842:Issidae
571:Insecta
569:Class:
544:Extant
469:2423912
146:Genera
99:Order:
93:Insecta
89:Class:
2445:226395
2432:603647
2406:105364
2393:246110
2354:674417
2253:Note:
503:
467:
391:
383:
278:. The
140:, 1851
138:Dallas
2473:This
2401:IRMNG
2367:12857
2341:17126
2261:from
2259:clade
1258:(with
501:S2CID
465:JSTOR
426:(PDF)
389:S2CID
361:(PDF)
315:(PDF)
243:basal
2479:stub
2419:7527
2414:NCBI
2375:GBIF
2349:BOLD
381:PMID
323:103B
286:and
493:doi
457:doi
373:doi
2524::
2442::
2429::
2416::
2403::
2390::
2377::
2364::
2351::
2326::
2311::
499:.
489:11
487:.
463:.
453:85
451:.
434:23
432:.
428:.
387:.
379:.
369:24
367:.
363:.
348:^
321:.
317:.
2510:e
2503:t
2496:v
2485:.
2269:.
1807:)
1803:(
1264:)
537:e
530:t
523:v
507:.
495::
471:.
459::
395:.
375::
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