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Vessel element

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199:: more or less circular regions in contact with neighbouring cells. Tracheids also have pits, but only vessel elements have openings at both ends that connect individual vessel elements to form a continuous tubular vessel. These end openings are called perforations or perforation plates. They have a variety of shapes: the most common are the simple perforation (a simple opening) and the scalariform perforation (several elongated openings in a ladder-like design). Other types include the foraminate perforation plate (several round openings) and the reticulate perforation plate (a net-like pattern, with many openings). 36: 844: 93: 180: 175:
and vessel elements. Vessel elements are the building blocks of vessels, the conducting pathways that constitute the major part of the water transporting system in flowering plants. Vessels form an efficient system for transporting water (including necessary minerals) from the root to the leaves and
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The presence of vessels in xylem has been considered to be one of the key innovations that led to the success of the flowering plants. It was once thought that vessel elements were an evolutionary innovation of flowering plants, but their absence from some basal
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suggest that this hypothesis must be re-examined; vessel elements in Gnetales may not be homologous with those of angiosperms, or vessel elements that originated in a precursor to the angiosperms may have been subsequently lost in some basal lineages (e.g.,
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At maturity, the protoplast – the living material of the cell – dies and disappears, but the lignified cell walls persist. A vessel element is then a dead cell, but one that still has a function, and is still being protected by surrounding living cells.
191:. A long cell, oriented along the axis of the stem, called a "fusiform initial", divides along its length forming new vessel elements. The cell wall of a vessel element becomes strongly "lignified", i.e. it develops reinforcing material made of 284:. In these cases, it is generally agreed that the vessels evolved independently. It is possible that vessels may have appeared more than once among the angiosperms as well. 507: 171:
that conducts water (and substances dissolved in it) upwards from the roots to the shoots. Two kinds of cell are involved in xylem transport:
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In secondary xylem – the xylem that is produced as a stem thickens rather than when it first appears – a vessel element originates from the
476: 446: 79: 57: 500: 337: 878: 96: 493: 847: 456:. Eidgenössische Forschungsanstalt für Wald, Schnee und Landscaft, Birmensdorf (Hrsg.) Haupt, Bern und Stuttgart. 50: 44: 873: 61: 260: 596: 591: 570: 532: 472: 442: 391: 383: 281: 379: 795: 557: 524: 516: 375: 234: 226: 188: 828: 809: 766: 625: 136: 123: 409: 122:, the water conducting tissue of plants. Vessel elements are found in most angiosperms ( 637: 576: 547: 168: 883: 867: 739: 697: 691: 581: 542: 222: 790: 733: 708: 277: 272: 196: 99:
image (top) and Transmission Light Microscope image (bottom) of vessel elements in
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as "primitively vesselless". Cronquist considered the vessels of
718: 489: 100: 29: 463:. Finnish Museum of Natural History, University of Helsinki. 144:. Vessel elements are the main feature distinguishing the " 454:
Anatomie europäischer Hölzer - Anatomy of European woods
441:. Chicago and London: The University of Chicago Press. 246:
Vessel-like cells have also been found in the xylem of
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Stained Xylem Vessel in Red/Orange (x400 magnification)
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Turner, Simon; Gallois, Patrick; Brown, David (2007).
338:"Vessels vs Tracheids, Similarities and Differences" 556: 523: 469:The Anatomy of Wood: Its Diversity and Variability 461:Introduction to Microscopic Wood Identification 501: 8: 243:to be convergent with those of angiosperms. 508: 494: 486: 216:and their presence in some members of the 195:. The side walls of a vessel element have 80:Learn how and when to remove this message 380:10.1146/annurev.arplant.57.032905.105236 43:This article includes a list of general 305: 7: 467:Wilson, K.; White, D. J. B. (1986). 118:) is one of the cell types found in 364:"Tracheary element differentiation" 439:The Evolutionary Biology of Plants 280:), and the enigmatic fossil group 49:it lacks sufficient corresponding 25: 471:. London: Stobart & Son Ltd. 843: 842: 34: 368:Annual Review of Plant Biology 1: 452:Schweingruber, F. H. (1990) 148:" of angiosperms from the " 900: 176:other parts of the plant. 838: 207:Evolutionary significance 314:"Vessel | plant anatomy" 459:Timonen, Tuuli (2002). 437:Niklas, Karl J. (1997) 318:Encyclopedia Britannica 64:more precise citations. 184: 103: 879:Plant stem morphology 182: 95: 755:Meristematic tissue 261:Pteridium aquilinum 517:Biological tissues 432:Further references 342:Easy Biology Class 185: 104: 27:Component of Xylem 861: 860: 648:Phloem parenchyma 282:Gigantopteridales 167:is the tissue in 90: 89: 82: 16:(Redirected from 891: 846: 845: 796:Vascular cambium 682:Xylem parenchyma 510: 503: 496: 487: 482: 425: 424: 422: 421: 406: 400: 399: 359: 353: 352: 350: 349: 334: 328: 327: 325: 324: 310: 264:(bracken fern), 258:(spike-mosses), 235:Arthur Cronquist 233:), described by 227:Trochodendraceae 189:vascular cambium 140:, but absent in 124:flowering plants 85: 78: 74: 71: 65: 60:this article by 51:inline citations 38: 37: 30: 21: 899: 898: 894: 893: 892: 890: 889: 888: 864: 863: 862: 857: 834: 767:Ground meristem 626:Vascular tissue 597:Subsidiary cell 552: 519: 514: 479: 466: 434: 429: 428: 419: 417: 408: 407: 403: 361: 360: 356: 347: 345: 336: 335: 331: 322: 320: 312: 311: 307: 302: 290: 209: 169:vascular plants 162: 114:(also called a 86: 75: 69: 66: 56:Please help to 55: 39: 35: 28: 23: 22: 18:Vessel elements 15: 12: 11: 5: 897: 895: 887: 886: 881: 876: 866: 865: 859: 858: 856: 855: 850: 839: 836: 835: 833: 832: 825: 824: 823: 818: 801: 800: 799: 798: 793: 781: 780: 779: 774: 769: 750: 749: 748: 747: 742: 730: 723: 722: 721: 716: 711: 706: 687: 686: 685: 684: 679: 674: 672:Vessel element 669: 657: 656: 655: 650: 645: 640: 638:Companion cell 621: 620: 619: 618: 613: 601: 600: 599: 594: 589: 584: 579: 577:Bulliform cell 562: 560: 554: 553: 551: 550: 545: 540: 535: 529: 527: 521: 520: 515: 513: 512: 505: 498: 490: 484: 483: 477: 464: 457: 450: 433: 430: 427: 426: 414:Biology Online 410:"Xylem vessel" 401: 354: 329: 304: 303: 301: 298: 297: 296: 289: 286: 252:(horsetails), 208: 205: 161: 158: 126:) and in some 108:vessel element 88: 87: 42: 40: 33: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 896: 885: 882: 880: 877: 875: 874:Plant anatomy 872: 871: 869: 854: 851: 849: 841: 840: 837: 831: 830: 826: 822: 819: 817: 814: 813: 812: 811: 806: 803: 802: 797: 794: 792: 789: 788: 787: 786: 782: 778: 775: 773: 770: 768: 765: 764: 763: 762: 757: 756: 752: 751: 746: 743: 741: 738: 737: 736: 735: 731: 729: 728: 724: 720: 717: 715: 712: 710: 707: 705: 702: 701: 700: 699: 694: 693: 692:Ground tissue 689: 688: 683: 680: 678: 675: 673: 670: 668: 665: 664: 663: 662: 658: 654: 651: 649: 646: 644: 641: 639: 636: 635: 634: 633: 628: 627: 623: 622: 617: 614: 612: 609: 608: 607: 606: 602: 598: 595: 593: 592:Pavement cell 590: 588: 585: 583: 580: 578: 575: 574: 573: 572: 567: 566:Dermal tissue 564: 563: 561: 559: 555: 549: 546: 544: 541: 539: 536: 534: 531: 530: 528: 526: 522: 518: 511: 506: 504: 499: 497: 492: 491: 488: 480: 478:0-85442-033-9 474: 470: 465: 462: 458: 455: 451: 448: 447:0-226-58082-2 444: 440: 436: 435: 431: 415: 411: 405: 402: 397: 393: 389: 385: 381: 377: 373: 369: 365: 358: 355: 343: 339: 333: 330: 319: 315: 309: 306: 299: 295: 292: 291: 287: 285: 283: 279: 278:aquatic ferns 275: 274: 269: 268: 263: 262: 257: 256: 251: 250: 244: 242: 241: 236: 232: 228: 224: 223:Amborellaceae 219: 215: 206: 204: 200: 198: 194: 190: 181: 177: 174: 170: 166: 159: 157: 155: 151: 147: 143: 139: 138: 133: 129: 125: 121: 117: 113: 112:vessel member 109: 102: 98: 94: 84: 81: 73: 70:November 2014 63: 59: 53: 52: 46: 41: 32: 31: 19: 827: 808: 804: 791:Cork cambium 783: 759: 753: 734:Sclerenchyma 732: 725: 709:Chlorenchyma 696: 690: 671: 659: 643:Phloem fiber 630: 624: 603: 569: 565: 468: 460: 453: 438: 418:. Retrieved 416:. 2019-10-07 413: 404: 371: 367: 357: 346:. Retrieved 344:. 2016-12-12 341: 332: 321:. Retrieved 317: 308: 273:Regnellidium 271: 265: 259: 253: 247: 245: 238: 210: 201: 186: 163: 135: 116:xylem vessel 115: 111: 107: 105: 76: 67: 48: 727:Collenchyma 677:Xylem fiber 374:: 407–433. 255:Selaginella 231:Winteraceae 214:angiosperms 128:gymnosperms 62:introducing 868:Categories 816:Endodermis 772:Procambium 704:Aerenchyma 698:Parenchyma 653:Sieve tube 616:Phelloderm 587:Guard cell 538:Epithelial 533:Connective 420:2021-12-31 348:2021-12-31 323:2021-03-30 300:References 45:references 853:Histology 821:Exodermis 785:Secondary 777:Protoderm 714:Mesophyll 571:Epidermis 388:1543-5008 249:Equisetum 173:tracheids 848:Category 745:Sclereid 667:Tracheid 605:Periderm 543:Muscular 396:17472568 294:Tracheid 288:See also 267:Marsilea 218:Gnetales 154:conifers 150:softwood 146:hardwood 142:conifers 130:such as 761:Primary 611:Phellem 582:Cuticle 548:Nervous 525:Animals 160:Anatomy 137:Ephedra 58:improve 810:Cortex 632:Phloem 558:Plants 475:  445:  394:  386:  240:Gnetum 229:, and 193:lignin 132:cycads 47:, but 829:Stele 805:Mixed 740:Fiber 661:Xylem 165:Xylem 152:" of 120:xylem 884:Wood 719:Pith 473:ISBN 443:ISBN 392:PMID 384:ISSN 270:and 197:pits 134:and 376:doi 110:or 101:Oak 97:SEM 870:: 807:: 758:: 695:: 629:: 568:: 412:. 390:. 382:. 372:58 370:. 366:. 340:. 316:. 225:, 156:. 106:A 509:e 502:t 495:v 481:. 449:. 423:. 398:. 378:: 351:. 326:. 276:( 83:) 77:( 72:) 68:( 54:. 20:)

Index

Vessel elements
references
inline citations
improve
introducing
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SEM
Oak
xylem
flowering plants
gymnosperms
cycads
Ephedra
conifers
hardwood
softwood
conifers
Xylem
vascular plants
tracheids

vascular cambium
lignin
pits
angiosperms
Gnetales
Amborellaceae
Trochodendraceae
Winteraceae

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