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Raphael Weldon

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In 1894, Weldon rolled a set of 12 dice 26,306 times. He collected the data in part, 'to judge whether the differences between a series of group frequencies and a theoretical law, taken as a whole, were or were not more than might be attributed to the chance fluctuations of random sampling.' Weldon's
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in 1899. In the years of their collaboration Pearson laid the foundations of modern statistics. Magnello emphasises this side of Weldon's career. In 1900 he took the DSc degree and as Linacre Professor he also held a Fellowship at
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gives a detailed account of the controversy. The debate lost much of its intensity with the death of Weldon in 1906, though the general debate between the biometricians and the Mendelians continued until the creation of the
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Pearson, Karl (1900). On the criterion that a given system of derivations from the probable in the case of a correlated system of variables is such that it can be reasonably supposed to have arisen from random sampling.
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had developed for he had come to the view that "the problem of animal evolution is essentially a statistical problem." Weldon began working with his University College colleague, the mathematician
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who influenced him greatly; Weldon gave up his plans for a career in medicine. In 1881 he gained a first-class honours degree in the Natural Science Tripos; in the autumn he left for the
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on the other. Bateson, who had been taught by Weldon, took a very strong line against the biometricians. This bitter dispute ranged across substantive issues of the nature of
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His interests were changing from morphology to problems in variation and organic correlation. He began using the statistical techniques that
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On his religious views, he considered himself an agnostic. He died in 1906 of acute pneumonia, and is buried at Holywell Church, Oxford.
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Weldon's Search for a Direct Proof of Natural Selection and the Tortuous Path to the Neo-Darwinian Synthesis
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Proceedings of the Royal Society, 54, (1893), 318–329. An example of Weldon's use of statistical methods
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He was through the many years the present writer knew him, like his hero Huxley, a confirmed Agnostic.
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in 1890. Royal Society records show his election supporters included the great zoologists of the day:
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W.B. Provine (1971). The origins of theoretical population genetics. University of Chicago Press.
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was rediscovered and this precipitated a conflict between Weldon and Pearson on the one side and
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dice data were used by Karl Pearson in his pioneering paper on the chi-squared statistic.
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Weldon was one of the first scientists to provide evidence of stabilizing and directional
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was established by the University of Oxford in his honour; it is awarded annually.
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Upon returning to Cambridge in 1882, he was appointed university lecturer in
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Medicine was his intended career and he spent the academic year 1876-1877 at
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Walter Frank Raphael Weldon 1860–1906: A Memoir Reprinted from Biometrika
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Walter Frank Raphael Weldon 1860–1906: A Memoir Reprinted from Biometrika
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and methodological issues such as the value of the statistical method.
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In 1889 Weldon succeeded Lankester in the Jodrell Chair of Zoology at
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Weldon was the second child of the journalist and industrial chemist,
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arising from the work of the Royal Society Committee, Weldon wrote:
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to begin the first of his studies on marine biological organisms.
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Kemp, A.W., and C.D. Kemp. (1991). Weldon's dice data revisited,
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There Weldon studied with the developmental morphologist
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Jodrell Professors of Zoology and Comparative Anatomy
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Oxford: Basil Blackwell. p. 5. 488:Grier, David Alan (1 November 2013). 7: 809:"Weldon, Walter Frank Raphael"  594:"Weldon, Walter Frank Raphael"  293:, and as curator of what is now the 921:Alumni of University College London 25: 916:Fellows of Merton College, Oxford 642:Merton College Register 1900–1964 429:Davis, Lea K. (27 October 2022). 941:Deaths from pneumonia in England 819:Dictionary of National Biography 772:Works by or about Raphael Weldon 604:. Vol. 3. pp. 629–631. 600:Dictionary of National Biography 518:Mathematics in Victorian Britain 214:. Having studied mathematics at 961:19th-century British zoologists 926:Alumni of King's College London 871:British evolutionary biologists 748:Statisticians of the Centuries 726:. Cambridge University Press. 494:. Princeton University Press. 1: 946:Linacre Professors of Zoology 394:modern evolutionary synthesis 896:Fellows of the Royal Society 640:Levens, R.G.C., ed. (1964). 448:10.1371/journal.pgen.1010379 244:St John's College, Cambridge 137:St John's College, Cambridge 96:St John's College, Cambridge 547:A Cambridge Alumni Database 167:Walter Frank Raphael Weldon 106:Fellow of the Royal Society 36:Walter Frank Raphael Weldon 977: 838:Portraits of Statisticians 549:. University of Cambridge. 684:The American Statistician 491:When Computers Were Human 297:, and was elected to the 291:University College London 255:Naples Zoological Station 228:University College London 216:Girton College, Cambridge 160: 141:University College London 111: 41: 797:University of St Andrews 654:Amitabh, Joshi. (2017). 352:in natural populations. 334:Linacre Chair of Zoology 274:Oxford University Museum 155:Francis Maitland Balfour 804:Bourne, Gilbert Charles 589:Bourne, Gilbert Charles 295:Grant Museum of Zoology 701:Philosophical Magazine 343:Merton College, Oxford 276: 234:and the mathematician 174:evolutionary biologist 881:English statisticians 755:Proc Roy Soc Series B 560:Karl Pearson (2011). 401:Weldon Memorial Prize 399:After his death, the 271: 240:King's College London 951:People from Highgate 849:UK National Archives 834:Photograph of Weldon 783:Robertson, Edmund F. 406: 375:In 1900 the work of 781:O'Connor, John J.; 688:doi:10.2307/2684294 886:English zoologists 277: 222:Life and education 876:English agnostics 733:978-1-107-60122-2 703:, 5(50), 157–175. 686:, 45(3):216–222. 528:978-0-19-162794-1 501:978-1-4008-4936-9 338:Oxford University 176:and a founder of 164: 163: 151:Academic advisors 145:Oxford University 113:Scientific career 16:(Redirected from 968: 891:Biostatisticians 857: 852: 824: 821:(2nd supplement) 811: 799: 787:"Raphael Weldon" 776:Internet Archive 737: 704: 696: 690: 680: 674: 671: 665: 664:22 (6): 525-548. 652: 646: 645: 637: 631: 630: 628: 626: 612: 606: 605: 602:(2nd supplement) 596: 585: 579: 578: 557: 551: 550: 539: 533: 532: 512: 506: 505: 485: 479: 478: 468: 450: 441:(10): e1010379. 426: 232:E. Ray Lankester 79: 63: 61: 46: 32: 21: 976: 975: 971: 970: 969: 967: 966: 965: 861: 860: 843: 802: 780: 768: 734: 716: 713: 708: 707: 697: 693: 681: 677: 672: 668: 653: 649: 639: 638: 634: 624: 622: 620:blogs.ucl.ac.uk 614: 613: 609: 587: 586: 582: 572: 559: 558: 554: 541: 540: 536: 529: 514: 513: 509: 502: 487: 486: 482: 428: 427: 423: 418: 409: 381:William Bateson 266: 251:Francis Balfour 224: 200: 143: 139: 92:Alma mater 87: 81: 77: 68: 67:London, England 65: 59: 57: 49: 37: 28: 23: 22: 18:W. F. R. Weldon 15: 12: 11: 5: 974: 972: 964: 963: 958: 953: 948: 943: 938: 933: 928: 923: 918: 913: 908: 903: 898: 893: 888: 883: 878: 873: 863: 862: 859: 858: 841: 831: 825: 800: 778: 767: 766:External links 764: 763: 762: 751: 744: 738: 732: 712: 709: 706: 705: 691: 675: 666: 647: 632: 607: 580: 570: 552: 534: 527: 521:. OUP Oxford. 507: 500: 480: 420: 419: 417: 414: 408: 405: 396:in the 1930s. 373: 372: 326:Francis Galton 265: 262: 223: 220: 199: 196: 188:Francis Galton 162: 161: 158: 157: 152: 148: 147: 134: 130: 129: 120: 116: 115: 109: 108: 103: 99: 98: 93: 89: 88: 82: 80:(aged 46) 74: 70: 69: 66: 55: 51: 50: 48:Raphael Weldon 47: 39: 38: 35: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 973: 962: 959: 957: 954: 952: 949: 947: 944: 942: 939: 937: 934: 932: 929: 927: 924: 922: 919: 917: 914: 912: 909: 907: 904: 902: 899: 897: 894: 892: 889: 887: 884: 882: 879: 877: 874: 872: 869: 868: 866: 856: 850: 846: 842: 839: 835: 832: 829: 826: 822: 820: 815: 810: 805: 801: 798: 794: 793: 788: 784: 779: 777: 773: 770: 769: 765: 760: 756: 752: 749: 745: 742: 739: 735: 729: 725: 724: 719: 718:Pearson, Karl 715: 714: 710: 702: 695: 692: 689: 685: 679: 676: 670: 667: 663: 662: 657: 651: 648: 643: 636: 633: 621: 617: 611: 608: 603: 601: 595: 590: 584: 581: 577: 573: 571:9781107601222 567: 563: 556: 553: 548: 544: 538: 535: 530: 524: 520: 519: 511: 508: 503: 497: 493: 492: 484: 481: 476: 472: 467: 462: 458: 454: 449: 444: 440: 436: 435:PLOS Genetics 432: 425: 422: 415: 413: 407:Weldon's dice 404: 402: 397: 395: 390: 386: 382: 378: 377:Gregor Mendel 370: 369: 368: 366: 362: 358: 353: 351: 346: 344: 339: 335: 331: 327: 322: 320: 316: 312: 308: 305:, Lankester, 304: 300: 299:Royal Society 296: 292: 287: 285: 282: 275: 270: 263: 261: 258: 256: 252: 247: 245: 241: 237: 236:Olaus Henrici 233: 229: 221: 219: 217: 213: 209: 208:Florence Tebb 205: 204:Walter Weldon 197: 195: 193: 189: 185: 184: 179: 175: 171: 168: 159: 156: 153: 149: 146: 142: 138: 135: 131: 128: 124: 121: 117: 114: 110: 107: 104: 100: 97: 94: 90: 85: 76:13 April 1906 75: 71: 64:15 March 1860 56: 52: 45: 40: 33: 30: 19: 817: 790: 758: 754: 747: 741:W.B. 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Index

W. F. R. Weldon

Oxford
St John's College, Cambridge
Fellow of the Royal Society
Zoology
biometry
St John's College, Cambridge
University College London
Oxford University
Francis Maitland Balfour
FRS
evolutionary biologist
biometry
Biometrika
Francis Galton
Karl Pearson
Walter Weldon
Florence Tebb
William Tebb
Girton College, Cambridge
University College London
E. Ray Lankester
Olaus Henrici
King's College London
St John's College, Cambridge
Francis Balfour
Naples Zoological Station

Oxford University Museum

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