75:
50:
31:
378:(SVL) of 105–130 mm (4.1–5.1 in). The back is dark brown with small yellow spots forming bands. The belly is light brown, and the throat and lips are mottled. Males have 10-12 femoral pores. The tail is spiny and slightly longer than the SVL.
1669:
1735:
1625:
1705:
1573:
1599:
637:; Mouton, P. Le Fras N. (2011). "Between a rock and a hard polytomy: Rapid radiation in the rupicolous girdled lizards (Squamata: Cordylidae)".
534:
676:
1730:
615:
219:
579:(1908). "On a collection of fresh-water fishes, batrachians, and reptiles from Natal and Zululand, with description of new species".
1725:
605:
959:
1604:
1715:
869:
1076:
842:
1534:
1419:
1265:
1459:
1720:
1630:
555:
1435:
1256:
1184:
1175:
1127:
1118:
421:
1411:
1403:
1301:
1151:
1142:
977:
941:
755:
669:
74:
1370:
1085:
824:
576:
281:
186:
1710:
1487:
1103:
1043:
1010:
833:
797:
737:
1539:
1310:
1283:
1274:
1247:
1067:
1001:
968:
878:
860:
504:
169:
1334:
1560:
1443:
1427:
1379:
1343:
1034:
932:
905:
851:
764:
403:
1238:
1229:
1211:
896:
806:
773:
662:
39:
1525:
1292:
1220:
1094:
923:
343:
and rock outcrops on wooded mountain slopes, at altitudes of 300–800 m (980–2,620 ft).
195:
69:
1617:
630:
1651:
1547:
1361:
1202:
815:
746:
601:
530:
375:
316:
232:
634:
593:
1664:
1656:
1352:
1193:
499:
262:
1638:
914:
481:
456:
416:
269:
446:
Warren's girdled lizard was formerly available in the pet trade, possibly exported from
1451:
1394:
1325:
950:
887:
146:
1699:
1612:
490:
409:
390:
59:
54:
30:
1552:
1025:
992:
430:
312:
1682:
288:, is in honour of British zoologist Ernest Warren (1871–1945), who collected the
1586:
1519:
1166:
1058:
728:
1510:
686:
447:
435:
360:
308:
265:
136:
654:
356:
320:
86:
1643:
1504:
788:
717:
711:
705:
304:
289:
126:
106:
1578:
340:
332:
254:
116:
1591:
699:
258:
96:
1481:
560:
1677:
600:. Sanibel Island, Florida: Ralph Curtis Books Publishing: 399 pp.
1565:
1485:
658:
475:
473:
529:. Baltimore: Johns Hopkins University Press. xiii + 296 pp.
598:
Field Guide to Snakes and other
Reptiles of Southern Africa
525:
Beolens, Bo; Watkins, Michael; Grayson, Michael (2011).
1494:
1392:
1323:
1164:
1140:
1116:
1056:
1023:
990:
786:
726:
505:
10.2305/IUCN.UK.2018-2.RLTS.T110167218A115679533.en
670:
622:. Pretoria: Transvaal Museum. xvi + 528 pp. (
8:
1482:
677:
663:
655:
48:
29:
20:
503:
319:in northeastern South Africa and eastern
469:
480:Bates, M.F.; Mouton, P.L.F.N. (2018).
1736:Taxa named by George Albert Boulenger
639:Molecular Phylogenetics and Evolution
549:
547:
7:
1706:IUCN Red List least concern species
491:IUCN Red List of Threatened Species
960:East African spiny-tailed lizard (
14:
527:The Eponym Dictionary of Reptiles
428:), and the regal girdled lizard (
73:
612:, pp. 195–196 + Plate 70).
450:. Most specimens were labeled
414:, Zoutpansberg girdled lizard (
257:of relatively large, flattened
888:Western dwarf girdled lizard (
401:The Barberton girdled lizard (
1:
1266:Orange-throated flat lizard (
1128:Blue-spotted girdled lizard (
1044:African spiny-tailed lizard (
852:Coastal spiny-tailed lizard (
407:), Waterberg girdled lizard (
1420:Zoutpansberg girdled lizard
1176:Attenborough's flat lizard (
933:N’Dolondolo girdled lizard (
870:McLachlan's girdled lizard (
438:of Warren's girdled lizard.
1077:Lawrence's girdled lizard (
1068:Campbell's girdled lizard (
924:Oelofsen's girdled lizard (
620:The Lizards of South Africa
434:) were formerly considered
268:. The species is native to
1752:
1731:Reptiles described in 1908
1436:Mozambique girdled lizard
1302:Soutpansberg flat lizard (
1152:Armadillo girdled lizard (
978:Transvaal girdled lizard (
951:Ethiopian girdled lizard (
942:Rhodesian girdled lizard (
897:Kaokoveld girdled lizard (
843:Machado's girdled lizard (
756:Large-scale grass lizard (
581:Annals of the Natal Museum
498:: e.T110167218A115679533.
452:Cordylus warreni depressus
1460:Van Dam's girdled lizard
1412:Waterberg girdled lizard
1404:Barberton girdled lizard
1371:Drakensberg crag lizard (
1035:Jordan's girdled lizard (
825:Rooiberg girdled lizard (
694:
563:www.reptile-database.org.
454:and should be considered
422:Mozambique girdled lizard
201:
194:
175:
168:
70:Scientific classification
68:
46:
37:
28:
23:
1726:Reptiles of South Africa
1468:Warren's girdled lizard
1257:Mitchell's flat lizard (
1185:Broadley's flat lizard (
1095:Peers’s girdled lizard (
1086:Namaqua girdled lizard (
861:Marungu girdled lizard (
834:Limpopo girdled lizard (
798:Angolan girdled lizard (
738:Transvaal grass lizard (
24:Warren's girdled lizard
1284:Sekukhune flat lizard (
1275:Ocellated flat lizard (
1248:Waterberg flat lizard (
1104:Herero girdled lizard (
969:Ukinga girdled lizard (
879:Mecula girdled lizard (
807:Maasai girdled lizard (
245:Warren's girdled lizard
1380:Northern crag lizard (
1344:Highveld crag lizard (
915:Nyika girdled lizard (
906:Black girdled lizard (
765:Zambian grass lizard (
331:The preferred natural
1444:Regal girdled lizard
1428:Giant girdled lizard
1311:Striped flat lizard (
1239:Spotted flat lizard (
1230:Lebombo flat lizard (
1212:Emperor flat lizard (
816:Cape girdled lizard (
633:; Jackmann, Todd R.;
1716:Reptiles of Eswatini
1452:Swazi dragon lizard
1335:Lang's crag lizard (
1293:Pungwe flat lizard (
1221:Common flat lizard (
1130:N. coeruleopunctatus
1011:Dwarf cliff lizard (
629:Stanley, Edward L.;
561:The Reptile Database
404:Smaug barbertonensis
376:snout-to-vent length
1406:(S. barbertonensis)
1362:Spiny crag lizard (
1203:Dwarf flat lizard (
1002:Cape cliff lizard (
774:Cape snake lizard (
747:Cape grass lizard (
648:, new combination).
626:, new combination).
40:Conservation status
1353:Cape crag lizard (
1194:Cape flat lizard (
635:Branch, William R.
1721:Lizards of Africa
1693:
1692:
1652:Open Tree of Life
1488:Taxon identifiers
1479:
1478:
1382:P. transvaalensis
1355:P. microlepidotus
962:C. tropidosternum
535:978-1-4214-0135-5
426:Smaug mossambicus
317:Lebombo Mountains
242:
241:
63:
16:Species of lizard
1743:
1686:
1685:
1673:
1672:
1660:
1659:
1647:
1646:
1634:
1633:
1621:
1620:
1608:
1607:
1595:
1594:
1582:
1581:
1569:
1568:
1556:
1555:
1543:
1542:
1530:
1529:
1528:
1515:
1514:
1513:
1483:
1438:(S. mossambicus)
1178:P. attenboroughi
679:
672:
665:
656:
624:Cordylus warreni
610:Cordylus warreni
564:
551:
542:
539:Cordylus warreni
523:
517:
516:
514:
512:
507:
477:
296:Geographic range
236:
223:
214:Cordylus warreni
210:
181:
78:
77:
57:
52:
51:
33:
21:
1751:
1750:
1746:
1745:
1744:
1742:
1741:
1740:
1696:
1695:
1694:
1689:
1681:
1676:
1668:
1663:
1655:
1650:
1642:
1639:Observation.org
1637:
1629:
1624:
1616:
1611:
1603:
1598:
1590:
1585:
1577:
1572:
1564:
1559:
1551:
1546:
1538:
1533:
1524:
1523:
1518:
1509:
1508:
1503:
1490:
1480:
1475:
1388:
1319:
1232:P. lebomboensis
1160:
1154:O. cataphractus
1136:
1112:
1052:
1019:
986:
863:C. marunguensis
782:
722:
690:
683:
652:
631:Bauer, Aaron M.
590:, new species).
588:Zonurus warreni
586:(3): 219–235. (
573:
571:Further reading
568:
567:
552:
545:
524:
520:
510:
508:
479:
478:
471:
466:
457:Smaug depressus
444:
417:Smaug depressus
399:
384:
369:
351:A shy species,
349:
329:
298:
278:
270:Southern Africa
230:
229:
217:
216:
209:Boulenger, 1908
208:
207:
205:Zonurus warreni
190:
183:
177:
164:
161:S. warreni
72:
64:
53:
49:
42:
17:
12:
11:
5:
1749:
1747:
1739:
1738:
1733:
1728:
1723:
1718:
1713:
1708:
1698:
1697:
1691:
1690:
1688:
1687:
1674:
1661:
1648:
1635:
1622:
1609:
1596:
1583:
1570:
1557:
1544:
1531:
1516:
1500:
1498:
1492:
1491:
1486:
1477:
1476:
1474:
1473:
1465:
1457:
1449:
1441:
1433:
1430:(S. giganteus)
1425:
1422:(S. depressus)
1417:
1409:
1400:
1398:
1390:
1389:
1387:
1386:
1377:
1368:
1359:
1350:
1341:
1331:
1329:
1326:Pseudocordylus
1321:
1320:
1318:
1317:
1308:
1299:
1295:P. pungweensis
1290:
1281:
1272:
1263:
1254:
1245:
1236:
1227:
1223:P. intermedius
1218:
1209:
1200:
1191:
1182:
1172:
1170:
1162:
1161:
1159:
1158:
1148:
1146:
1138:
1137:
1135:
1134:
1124:
1122:
1114:
1113:
1111:
1110:
1101:
1092:
1088:N. namaquensis
1083:
1074:
1064:
1062:
1054:
1053:
1051:
1050:
1041:
1031:
1029:
1021:
1020:
1018:
1017:
1008:
998:
996:
988:
987:
985:
984:
975:
966:
957:
948:
944:C. rhodesianus
939:
935:C. phonolithos
930:
921:
912:
903:
894:
885:
876:
867:
858:
854:C. macropholis
849:
840:
831:
822:
813:
804:
794:
792:
784:
783:
781:
780:
771:
762:
753:
744:
734:
732:
724:
723:
721:
720:
714:
708:
702:
695:
692:
691:
684:
682:
681:
674:
667:
659:
650:
649:
627:
613:
591:
572:
569:
566:
565:
543:
518:
468:
467:
465:
462:
443:
440:
398:
395:
383:
380:
368:
365:
348:
345:
328:
325:
303:is known from
297:
294:
277:
274:
240:
239:
238:
237:
224:
211:
199:
198:
192:
191:
184:
173:
172:
166:
165:
158:
156:
152:
151:
144:
140:
139:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
66:
65:
47:
44:
43:
38:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
1748:
1737:
1734:
1732:
1729:
1727:
1724:
1722:
1719:
1717:
1714:
1712:
1711:Smaug (genus)
1709:
1707:
1704:
1703:
1701:
1684:
1679:
1675:
1671:
1666:
1662:
1658:
1653:
1649:
1645:
1640:
1636:
1632:
1627:
1623:
1619:
1614:
1610:
1606:
1601:
1597:
1593:
1588:
1584:
1580:
1575:
1571:
1567:
1562:
1558:
1554:
1549:
1545:
1541:
1536:
1532:
1527:
1526:Smaug warreni
1521:
1517:
1512:
1506:
1502:
1501:
1499:
1497:
1496:Smaug warreni
1493:
1489:
1484:
1472:
1471:
1466:
1464:
1463:
1458:
1456:
1455:
1454:(S. swazicus)
1450:
1448:
1447:
1442:
1440:
1439:
1434:
1432:
1431:
1426:
1424:
1423:
1418:
1416:
1415:
1410:
1408:
1407:
1402:
1401:
1399:
1397:
1396:
1391:
1385:
1383:
1378:
1376:
1374:
1373:P. subviridis
1369:
1367:
1365:
1360:
1358:
1356:
1351:
1349:
1347:
1342:
1340:
1338:
1333:
1332:
1330:
1328:
1327:
1322:
1316:
1314:
1309:
1307:
1305:
1300:
1298:
1296:
1291:
1289:
1287:
1286:P. orientalis
1282:
1280:
1278:
1273:
1271:
1269:
1268:P. monotropis
1264:
1262:
1260:
1255:
1253:
1251:
1246:
1244:
1242:
1237:
1235:
1233:
1228:
1226:
1224:
1219:
1217:
1215:
1210:
1208:
1206:
1201:
1199:
1197:
1192:
1190:
1188:
1183:
1181:
1179:
1174:
1173:
1171:
1169:
1168:
1163:
1157:
1155:
1150:
1149:
1147:
1145:
1144:
1139:
1133:
1131:
1126:
1125:
1123:
1121:
1120:
1115:
1109:
1107:
1106:N. pustulatus
1102:
1100:
1098:
1093:
1091:
1089:
1084:
1082:
1080:
1075:
1073:
1071:
1066:
1065:
1063:
1061:
1060:
1055:
1049:
1047:
1042:
1040:
1038:
1033:
1032:
1030:
1028:
1027:
1022:
1016:
1014:
1009:
1007:
1005:
1000:
999:
997:
995:
994:
989:
983:
981:
976:
974:
972:
971:C. ukingensis
967:
965:
963:
958:
956:
954:
949:
947:
945:
940:
938:
936:
931:
929:
927:
922:
920:
918:
913:
911:
909:
904:
902:
900:
899:C. namakuiyus
895:
893:
891:
886:
884:
882:
877:
875:
873:
872:C. mclachlani
868:
866:
864:
859:
857:
855:
850:
848:
846:
841:
839:
837:
832:
830:
828:
823:
821:
819:
814:
812:
810:
809:C. beraduccii
805:
803:
801:
800:C. angolensis
796:
795:
793:
791:
790:
785:
779:
777:
772:
770:
768:
763:
761:
759:
758:C. macrolepis
754:
752:
750:
745:
743:
741:
736:
735:
733:
731:
730:
725:
719:
715:
713:
709:
707:
703:
701:
697:
696:
693:
688:
680:
675:
673:
668:
666:
661:
660:
657:
653:
647:
646:Smaug warreni
644:(1): 53–70. (
643:
640:
636:
632:
628:
625:
621:
617:
616:FitzSimons VF
614:
611:
607:
606:0-88359-042-5
603:
599:
595:
592:
589:
585:
582:
578:
575:
574:
570:
562:
558:
557:
556:Smaug warreni
550:
548:
544:
540:
536:
532:
528:
522:
519:
506:
501:
497:
493:
492:
487:
485:
484:Smaug warreni
476:
474:
470:
463:
461:
459:
458:
453:
449:
441:
439:
437:
433:
432:
427:
423:
419:
418:
413:
411:
410:Smaug breyeri
406:
405:
396:
394:
392:
391:ovoviviparous
388:
381:
379:
377:
373:
366:
364:
362:
358:
354:
346:
344:
342:
338:
334:
326:
324:
322:
318:
314:
310:
306:
302:
295:
293:
291:
287:
283:
282:specific name
275:
273:
271:
267:
264:
260:
256:
252:
251:
250:Smaug warreni
246:
234:
228:
227:Smaug warreni
225:
221:
220:V. FitzSimons
215:
212:
206:
203:
202:
200:
197:
193:
188:
182:
180:
179:Smaug warreni
174:
171:
170:Binomial name
167:
163:
162:
157:
154:
153:
150:
149:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
76:
71:
67:
61:
56:
55:Least Concern
45:
41:
36:
32:
27:
22:
19:
1495:
1470:(S. warreni)
1469:
1467:
1462:(S. vandami)
1461:
1453:
1445:
1437:
1429:
1421:
1414:(S. breyeri)
1413:
1405:
1393:
1381:
1372:
1363:
1354:
1346:P. melanotus
1345:
1336:
1324:
1313:P. torquatus
1312:
1303:
1294:
1285:
1277:P. ocellatus
1276:
1267:
1259:P. mitchelli
1258:
1249:
1241:P. maculatus
1240:
1231:
1222:
1214:P. imperator
1213:
1204:
1195:
1187:P. broadleyi
1186:
1177:
1165:
1153:
1141:
1129:
1117:
1105:
1096:
1087:
1078:
1070:N. campbelli
1069:
1057:
1046:K. polyzonus
1045:
1036:
1026:Karusasaurus
1024:
1013:H. nebulosus
1012:
1003:
993:Hemicordylus
991:
979:
970:
961:
952:
943:
934:
926:C. oelofseni
925:
916:
907:
898:
889:
880:
871:
862:
853:
844:
835:
826:
817:
808:
799:
787:
775:
767:C. miopropus
766:
757:
748:
739:
727:
651:
645:
641:
638:
623:
619:
609:
597:
587:
583:
580:
577:Boulenger GA
554:
538:
526:
521:
509:. Retrieved
495:
489:
483:
455:
451:
445:
431:Smaug regius
429:
425:
415:
408:
402:
400:
386:
385:
382:Reproduction
371:
370:
352:
350:
336:
330:
313:South Africa
300:
299:
285:
279:
249:
248:
244:
243:
235:et al., 2011
226:
213:
204:
178:
176:
160:
159:
147:
18:
1587:iNaturalist
1520:Wikispecies
1446:(S. regius)
1364:P. spinosus
1304:P. relictus
1205:P. guttatus
1196:P. capensis
1167:Platysaurus
1079:N. lawrenci
1059:Namazonurus
1004:H. capensis
980:C. vittifer
845:C. machadoi
818:C. cordylus
729:Chamaesaura
511:18 November
367:Description
361:vertebrates
355:eats large
1700:Categories
1037:K. jordani
881:C. meculae
836:C. jonesii
776:C. tenuior
749:C. anguina
687:Cordylidae
541:, p. 280).
464:References
448:Mozambique
436:subspecies
387:S. warreni
372:S. warreni
359:and small
357:arthropods
353:S. warreni
337:S. warreni
309:Mozambique
301:S. warreni
266:Cordylidae
137:Cordylidae
1618:110167218
1143:Ouroboros
1097:N. peersi
917:C. nyikae
827:C. imkeae
698:Kingdom:
618:(1943).
596:(2004).
442:Pet trade
276:Etymology
187:Boulenger
155:Species:
93:Kingdom:
87:Eukaryota
1678:Species+
1511:Q3007630
1505:Wikidata
1337:P. langi
1250:P. minor
953:C. rivae
908:C. niger
890:C. minor
789:Cordylus
740:C. aenea
718:Squamata
712:Reptilia
706:Chordata
704:Phylum:
700:Animalia
594:Branch B
553:Species
397:Taxonomy
333:habitats
321:Eswatini
305:Botswana
290:holotype
196:Synonyms
133:Family:
127:Squamata
117:Reptilia
107:Chordata
103:Phylum:
97:Animalia
83:Domain:
60:IUCN 3.1
1670:warreni
1605:1141848
1579:8255981
1566:1056426
1119:Ninurta
716:Order:
710:Class:
689:species
685:Extant
341:savanna
327:Habitat
286:warreni
261:in the
255:species
253:) is a
233:Stanley
189:, 1908)
143:Genus:
123:Order:
113:Class:
58: (
1657:376024
1644:939601
1631:885424
1540:751386
604:
533:
374:has a
311:, and
263:family
259:lizard
222:, 1943
1683:65690
1592:33046
1553:4XVBZ
1395:Smaug
315:(the
148:Smaug
1626:NCBI
1613:IUCN
1600:ITIS
1574:GBIF
1535:BOLD
602:ISBN
531:ISBN
513:2021
496:2018
347:Diet
339:are
280:The
1561:EoL
1548:CoL
608:. (
559:at
537:. (
500:doi
420:),
389:is
335:of
323:).
1702::
1680::
1667::
1665:RD
1654::
1641::
1628::
1615::
1602::
1589::
1576::
1563::
1550::
1537::
1522::
1507::
642:58
546:^
494:.
488:.
472:^
460:.
393:.
363:.
307:,
292:.
284:,
272:.
231:—
218:—
1384:)
1375:)
1366:)
1357:)
1348:)
1339:)
1315:)
1306:)
1297:)
1288:)
1279:)
1270:)
1261:)
1252:)
1243:)
1234:)
1225:)
1216:)
1207:)
1198:)
1189:)
1180:)
1156:)
1132:)
1108:)
1099:)
1090:)
1081:)
1072:)
1048:)
1039:)
1015:)
1006:)
982:)
973:)
964:)
955:)
946:)
937:)
928:)
919:)
910:)
901:)
892:)
883:)
874:)
865:)
856:)
847:)
838:)
829:)
820:)
811:)
802:)
778:)
769:)
760:)
751:)
742:)
678:e
671:t
664:v
584:1
515:.
502::
486:"
482:"
424:(
412:)
247:(
185:(
62:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
