559:
sticks placed in trees or shrubs. The male fetches sticks of the nest while the female places them to form the platform. The nest height varies from about 5 to 15 meters, but very low nesting (as low as 0.6 meters) has been observed in mangrove trees in salt pans. The usual clutch is three to four eggs which are light blue as in the little egret. Incubation begins when the first egg is laid resulting in chicks with size disparities. Both parents take turns to incubate and the eggs hatch after about 23 to 24 days. The chicks are white with grey dapples. Young birds sometimes die after falling off from the nest platform. Adults will shade the chicks during the hotter parts of the day. Predators of eggs include rats and young chicks may be taken by cats and crows. Adults usually guard the nest when eggs or young chicks are present. Adults feed the young by regurgitating semi-digested food at the nest. This diet includes mainly small fishes of the families
658:
444:
429:
31:
632:
583:
380:. There has been confusion related to what were thought to be dark little egrets and the possibility of hybrids. Dark morphs in little egrets are extremely rare and nearly all specimens of what were thought to be dark morphs have turned out to be western reef herons. In India some research claimed circumstantial evidence of interbreeding with little egrets, but the same researchers noted that the breeding seasons of the two species were different. Hybridization has also been claimed to occur in Morocco and Kenya (but this might refer to
601:
616:
459:
86:
471:
208:
61:
558:
rains from April to August and end of May in the former breeding colony at Chilaw in Sri Lanka. Most of the breeding colonies in
Gujarat in western India were on mangroves. They nest in colonies usually of their own species but sometimes with other egrets and smaller herons, usually on platforms of
521:
The western reef heron (nominate subspecies) occurs as a vagrant in North
America, South America and the Caribbean islands. Based on the growing number of records it is suspected that they may establish breeding colonies in Brazil. Several records around 1980–90 in Germany, Austria and France have
308:, but has a larger yellower bill, extended yellow on thicker legs, and when foraging tends to be very active, sometimes also moving its wing or using it to shade the water surface. The grey morph has a whitish throat and is unlikely to be confused with any other species within the range of this
413:
show blue and this is thought to be due to hybridization. The dark coastal form of
Madagascar, Aldabra, Comoro Islands, Seychelles and parts of East Africa (southern Kenya to Tanzania) was earlier treated as a subspecies (the mainland African form has sometimes been considered as
546:. Laboratory studies show them to be capable of making corrections for refractions but the probability of missing increases when they are forced to strike at prey at very acute angles to the water surface. Like other herons and egrets they have few vocalizations, making a low
657:
388:
genes from a sampled little egret and a western reef heron were identical and use this as evidence for demotion. The sequences and the origins of the samples are not publicly available or verifiable. In the past the Indian form which was also described as
303:
has two plumage colour forms. There is an all-white morph and a dark grey morph; morphs can also occur with intermediate shades of grey which may be related to age or particoloured in grey and white. The white morph is similar in general appearance to the
553:
The western reef heron's breeding habitat is coastal wetlands. In the Red Sea region the breeding season is from June to August. The nominate subspecies breeds in West Africa from late April to
September. In India the breeding season is during the
509:
where breeding was once recorded at Chilaw. The nominate subspecies breeds in west Africa from
Mauritania to Gabon. Birds may also be found off the mainland such as in the Canary Islands. Small numbers breed in Spain. Subspecies
312:
with beak and legs similar to that of the white morph. During the breeding season the legs and facial skin are reddish. Breeding birds have two long feathers on the sides of the nape. The nominate subspecies
631:
266:
of West Africa. There are also differences in size, structure and foraging behaviour. There have been suggestions that the species hybridizes with the Little Egret, and based on this, some authors treat
530:
These birds stalk their prey in shallow water, often running or stirring the water with their feet or flicking their wings to disturb prey; they may also stand still and wait to ambush prey. They eat
1104:"Current Status of the Breeding Population of the Western Reef Heron Egretta gularis along the Northern Coasts of the Persian Gulf and Oman Sea, and its Wintering Population in the South of Iran"
1055:"On the Relationship, Status and Range of Egretta garzetta, Demiegretta gularis, D. schistacea, D. asha, and D. dimorpha, a new subspecies and the correct type-locality of Egretta garzetta"
443:
2013:
1841:
428:
1880:
1973:
1483:"Impact of the food availability, nesting-habitat destruction and cultural variations of human settlements on the nesting distribution of a coastal bird,
1378:
Murphy, William L. (1992). "Notes on the
Occurrence of the little egret (Egretta garzetta) in the Americas, with Reference to Other Palearctic Vagrants".
336:. The dark and white morph is thought to be controlled by a single allele with the dark character being incompletely dominant over the gene for white.
1805:
600:
262:); a white form which can look very similar to the little egret although the bill tends to be paler and larger and the black form with white throat
1854:
1508:
Etezadifar, Farzaneh; Barati, Ahmad (2011). "Nestling diet of the
Western Reef Heron, Egretta gularis, in Hara Biosphere Reserve, Persian Gulf".
1103:
458:
352:
in 1792. The distinctive white throat (or gular region) in the dark morph gives it the species name. Later authors have treated it the genera
1906:
1988:
582:
952:"Cas probable d'hybridation Aigrette garzette Egretta garzetta x Aigrette des récifs Egretta gularis dans l'archipel d'Essaouira (Maroc)"
641:
321:(Hemprich & Ehrenberg, 1828) breeds from the Persian Gulf along the coast of India to the east of the India Peninsula. The bill of
30:
1753:
1993:
1359:
Cardillo, R.; Forbes-Watson, A.; Ridgely, R. (1983). "The
Western Reef–Heron (Egretta gularis) at Nantucket Island, Massachusetts".
2008:
1932:
246:
found in southern Europe, Africa and parts of Asia. It has a mainly coastal distribution and occurs in several plumage forms: a
1766:
1441:"Stationary underwater prey missed by reef herons, Egretta gularis: head position and light refraction at the moment of strike"
615:
514:
is found from the Red Sea coast east round the Indian coast. Breeding colonies are known from the east coast of India around
1859:
667:
329:
has the larger bill especially towards the base. The form on the eastern coast of South Africa is usually separated as the
349:
197:
978:
Inter-breeding between the Little Egret and the
Western Reef Heron in India, Africa and possibly Israel. Abstract No. 411
1654:
1151:
1260:
Murphy, W. L.; Nanan, W. (1987). "First confirmed record of
Western Reef-Heron (Egretta gularis) for South America".
1885:
2003:
1784:
902:
1797:
470:
85:
1983:
1413:
Fedrizzi, Carmem E.; Carlos, Caio J.; Vaske Jr., Teodoro; Bugoni, Leandro; Viana, Danielle; Véras, Dráusio P.
950:
Qninba, A.; Benhoussa, A.; Ibn Tattou, M.; El Idrissi Essougrati, A.; Haj, S. Ben; Idrissi, H. Rguibi (2011).
852:
Naik, R.M.; Parasharya, B.M. (1983). "Sequence of plumage changes and polymorphism in the Indian Reef Heron,
1998:
384:). Christidis and Boles quote a report by McCracken and Sheldon (2002) that the nucleotide sequences of the
903:"Reconsideration of a Hungarian specimen of a black-coloured egret as Western Reef Egret (Egretta gularis)"
1704:
1683:
1607:
1543:
Tere, Anika (2004). "Nesting of Western Reef-Egret Egretta gularis in the saltpans of G.H.C.L., Dholera".
951:
409:
of breeding little egrets are blue while those of the reef heron are reddish although some individuals of
251:
1315:
1231:
1440:
180:
1054:
1015:
711:
1937:
1696:
1665:
1645:
925:"The timing of breeding season and interbreeding between the colour phases in the Indian Reef Heron
1077:
394:
50:
1340:
Wilson, A. (2007). "Western Reef-Heron (Egretta gularis gularis) at Calvert Vaux Park, Brooklyn".
831:"The juvenile plumage of the Little Egret compared with that of the White-phase Indian Reef Heron"
1525:
1463:
1395:
1296:
80:
1596:
1846:
1000:
317:
has a range from West Africa to Gabon, with some birds breeding in southern Europe. Subspecies
1919:
1872:
1820:
1727:
1924:
792:
1978:
1517:
1455:
1414:
1387:
1288:
1279:
Davis Jr., William E. (1985). "Foraging Behavior of a Western Reef Heron in North America".
992:
706:
1709:
1893:
330:
220:
1670:
688:
1792:
1779:
1587:
406:
137:
1967:
1867:
993:
818:. Washington DC and Barcelona: Smithsonian Institution and Lynx Edicions. p. 55.
697:
279:. Works that consider the Western Reef Heron as a valid species include the nominate
70:
65:
1563:
1529:
1482:
1204:
1177:
924:
876:
830:
760:
734:
1732:
1521:
1467:
515:
498:
477:
385:
377:
305:
255:
207:
1950:
1771:
1719:
1828:
1639:
568:
1945:
543:
535:
522:
been attributed to birds that escaped from an animal dealer in Mittelfranken.
288:
1758:
1630:
250:-grey plumage in which it can only be confused with the rather uncommon dark
560:
506:
97:
1078:"Probable hybrids Little Egret x Indian Reef Heron in India and Sri Lanka"
1898:
1624:
564:
117:
1415:"Western Reef-Heron Egretta gularis in Brazil (Ciconiiformes: Ardeidae)"
1810:
1691:
1459:
1399:
1300:
645:
555:
539:
494:
450:
435:
157:
1678:
1316:"Trans-Atlantic vagrancy of Palearctic birds in Trindidad and Tobago"
490:
107:
1833:
1601:
1391:
1292:
1745:
1139:(2nd ed.). New Delhi: Oxford University Press. pp. 75–76.
1740:
649:
622:
502:
309:
247:
243:
147:
1016:"A further note on the Relationship, Status, and Distribution of
923:
Naik, R.M.; Parasharya, B.M.; Patel, B.H.; Mansuri, A.P. (1981).
1911:
592:
531:
300:
127:
1605:
761:"Changes in the soft part coloration of the Indian Reef Heron,
793:"Identification of western reef egrets and dark Little Egrets"
348:
from a specimen obtained in Senegal by the French naturalist
1152:"Occasional breeding by Western Reef Egret in eastern Spain"
418:) but is now raised to full species as the dimorphic egret (
372:
has however been highly debated with some authors treating
397:
has been treated as a subspecies of the little egret as
571:. The young birds leave the nest after about a month.
464:
A light grey form that has been considered as a hybrid
1439:
Katzir, Gadi; Lotem, Arnon; Intrator, Nathan (1989).
1137:
Handbook of the birds of India and Pakistan. Volume 2
1614:
550:or grating sounds when disturbed or near the nest.
712:10.2305/IUCN.UK.2016-3.RLTS.T22729692A95020328.en
666:dark morph in flight, showing white carpal area,
505:. It also occurs in the Lakshadweep Islands and
489:It occurs mainly on the coasts in tropical west
356:(in which dimorphic egrets were once included),
1053:Grant, CHB & C. W. Mackworth-Praed (1933).
816:Birds of South Asia. The Ripley Guide. Volume 2
364:until its current stable position in the genus
215:Range with subspecies and related species. For
1014:Grant, CHG & Mackworth-Praed, CW (1938).
980:. 19th International Ornithological Congress.
8:
1059:Bulletin of the British Ornithologists' Club
1036:Bulletin of the British Ornithologists' Club
995:Systematics and taxonomy of Australian birds
401:on the basis of presumed hybridization with
2014:Taxa named by Louis Augustin Guillaume Bosc
1588:IUCN Heron specialist group (treated under
1564:"Breeding biology of the Indian Reef heron"
1102:Etezadifar, Farzaneh; Amini, Hamid (2010).
791:Dubois, Philippe J.; Yésou, Pierre (1995).
786:
784:
782:
754:
752:
542:. In coastal areas, they regularly feed on
1602:
1314:Kenefick, Martyn; Hayes, Floyd E. (2006).
1150:Dies, J.I.; Prosper, J.; Dies, B. (2001).
991:Christidis, Les; Boles, Walter E. (2008).
765:(Bosc) related to age and breeding status"
518:. They occasionally occur further inland.
206:
59:
29:
20:
1232:"First record of the Western Reef-heron (
710:
896:
894:
814:Rasmussen, P.C.; Anderton, J.C. (2005).
368:. The distinctness of this species from
1076:Koparde, Pankaj; Yesou, Pierre (2017).
679:
578:
424:
1135:Ali, Salim; Ripley, S. Dillon (1978).
1130:
1128:
1126:
1124:
1562:Parasharya, B.M.; Naik, R.M. (1988).
1481:Naik, R.M.; Parasharya, B.M. (1987).
1236:) for St. Vincent and the Grenadines"
829:Parasharya, B.M.; Naik, R.M. (1984).
759:Parasharya, B.M.; Naik, R.M. (1987).
735:"Some notes on the Indian Reef Heron"
43:Dohat Arad Lagoon, Muharraq, Bahrain
7:
1798:9be1679a-2763-4e03-99eb-2b0dee58ec1e
1974:IUCN Red List least concern species
1448:Journal of Comparative Physiology A
1209:(Bosc.)] in Hyderabad District"
1205:"Occurrence of the Reef Heron [
698:IUCN Red List of Threatened Species
642:Marine National Park, Gulf of Kutch
344:The species was first described as
14:
1422:Revista Brasileira de Ornitologia
1243:Journal of Caribbean Ornithology
733:Dharmakumarsinhji, K.S. (1984).
656:
630:
614:
609:with a black beak (Pulicat Lake)
599:
581:
469:
457:
442:
427:
84:
687:BirdLife International (2016).
1597:Sound recordings on Xeno-Canto
1522:10.1080/09397140.2011.10648885
901:Magyar, G.; Yésou, P. (2000).
1:
1545:Newsletter for Ornithologists
976:Hancock, J; Naik, RM (1986).
350:Louis Augustin Guillaume Bosc
1989:Birds of the Gulf of Guinea
1203:Neavoll, George F. (1968).
399:Egretta garzetta schistacea
2030:
1510:Zoology in the Middle East
1230:Paice, Michael R. (2006).
1176:Kirkpatrick, K.M. (1961).
1182:(Bosc) on the East Coast"
501:(Iran) extending east to
214:
205:
186:
179:
81:Scientific classification
79:
57:
48:
37:
28:
23:
1994:Birds of the Middle East
1568:J. Bombay Nat. Hist. Soc
1491:J. Bombay Nat. Hist. Soc
1213:J. Bombay Nat. Hist. Soc
1186:J. Bombay Nat. Hist. Soc
933:J. Bombay Nat. Hist. Soc
881:J. Bombay Nat. Hist. Soc
835:J. Bombay Nat. Hist. Soc
769:J. Bombay Nat. Hist. Soc
739:J. Bombay Nat. Hist. Soc
705:: e.T22729692A95020328.
340:Taxonomy and systematics
2009:Birds described in 1792
485:Distribution and status
1178:"The Ashy Reef Heron,
877:"The dimorphic egrets"
325:is more pointed while
668:São Tomé and Príncipe
640:dark morph at Narara
526:Behaviour and ecology
1793:Fauna Europaea (new)
875:Berlioz, J. (1956).
621:particoloured form (
242:, is a medium-sized
1487:, in Western India"
1380:Colonial Waterbirds
1281:Colonial Waterbirds
395:William Henry Sykes
238:), also called the
51:Conservation status
24:Western reef heron
1821:western-reef-heron
1460:10.1007/BF00611243
1323:J. Carib. Ornithol
449:Slate grey morph,
240:western reef egret
230:western reef heron
2004:Birds of Pakistan
1961:
1960:
1920:Open Tree of Life
1608:Taxon identifiers
999:. CSIRO. p.
912:. 105–106: 35–40.
275:as subspecies of
226:
225:
74:
2021:
1954:
1953:
1941:
1940:
1928:
1927:
1915:
1914:
1902:
1901:
1889:
1888:
1876:
1875:
1863:
1862:
1850:
1849:
1837:
1836:
1824:
1823:
1814:
1813:
1801:
1800:
1788:
1787:
1775:
1774:
1762:
1761:
1749:
1748:
1736:
1735:
1723:
1722:
1713:
1712:
1700:
1699:
1697:2007348D400F8102
1687:
1686:
1674:
1673:
1661:
1660:
1650:
1649:
1648:
1635:
1634:
1633:
1603:
1590:Egretta garzetta
1576:
1575:
1559:
1553:
1552:
1540:
1534:
1533:
1505:
1499:
1498:
1478:
1472:
1471:
1445:
1436:
1430:
1429:
1419:
1410:
1404:
1403:
1375:
1369:
1368:
1356:
1350:
1349:
1337:
1331:
1330:
1320:
1311:
1305:
1304:
1276:
1270:
1269:
1257:
1251:
1250:
1240:
1227:
1221:
1220:
1200:
1194:
1193:
1173:
1167:
1166:
1156:
1147:
1141:
1140:
1132:
1119:
1118:
1108:
1099:
1093:
1092:
1082:
1073:
1067:
1066:
1050:
1044:
1043:
1018:Egretta garzetta
1011:
1005:
1004:
998:
988:
982:
981:
973:
967:
966:
956:
947:
941:
940:
920:
914:
913:
907:
898:
889:
888:
872:
866:
865:
849:
843:
842:
826:
820:
819:
811:
805:
804:
788:
777:
776:
756:
747:
746:
730:
724:
723:
721:
719:
714:
684:
660:
638:E. g. schistacea
634:
618:
607:E. g. schistacea
603:
585:
473:
461:
446:
431:
420:Egretta dimorpha
382:Egretta dimorpha
370:Egretta garzetta
334:Egretta dimorpha
277:Egretta garzetta
260:Egretta garzetta
210:
192:
89:
88:
68:
63:
62:
39:E. g. schistacea
33:
21:
2029:
2028:
2024:
2023:
2022:
2020:
2019:
2018:
1984:Birds of Africa
1964:
1963:
1962:
1957:
1951:Egretta-gularis
1949:
1944:
1936:
1931:
1923:
1918:
1910:
1905:
1897:
1894:Observation.org
1892:
1884:
1879:
1871:
1866:
1858:
1853:
1845:
1840:
1832:
1827:
1819:
1817:
1809:
1804:
1796:
1791:
1783:
1778:
1770:
1765:
1757:
1752:
1744:
1739:
1731:
1726:
1718:
1716:
1708:
1703:
1695:
1690:
1684:egretta-gularis
1682:
1677:
1671:Egretta_gularis
1669:
1664:
1658:
1653:
1646:Egretta gularis
1644:
1643:
1638:
1629:
1628:
1623:
1616:Egretta gularis
1610:
1584:
1579:
1561:
1560:
1556:
1542:
1541:
1537:
1507:
1506:
1502:
1485:Egretta gularis
1480:
1479:
1475:
1443:
1438:
1437:
1433:
1417:
1412:
1411:
1407:
1392:10.2307/1521360
1377:
1376:
1372:
1358:
1357:
1353:
1339:
1338:
1334:
1318:
1313:
1312:
1308:
1293:10.2307/1521198
1278:
1277:
1273:
1259:
1258:
1254:
1238:
1234:Egretta gularis
1229:
1228:
1224:
1207:Egretta gularis
1202:
1201:
1197:
1180:Egretta gularis
1175:
1174:
1170:
1154:
1149:
1148:
1144:
1134:
1133:
1122:
1106:
1101:
1100:
1096:
1080:
1075:
1074:
1070:
1052:
1051:
1047:
1013:
1012:
1008:
990:
989:
985:
975:
974:
970:
954:
949:
948:
944:
927:Egretta gularis
922:
921:
917:
905:
900:
899:
892:
874:
873:
869:
854:Egretta gularis
851:
850:
846:
828:
827:
823:
813:
812:
808:
790:
789:
780:
763:Egretta gularis
758:
757:
750:
732:
731:
727:
717:
715:
691:Egretta gularis
686:
685:
681:
677:
670:
661:
652:
635:
626:
619:
610:
604:
595:
589:Egretta gularis
586:
577:
528:
487:
480:
474:
465:
462:
453:
447:
438:
432:
403:Egretta gularis
376:as a polytypic
342:
331:dimorphic egret
297:
235:Egretta gularis
221:dimorphic egret
201:
194:
190:Egretta gularis
188:
175:
172:E. gularis
83:
75:
64:
60:
53:
17:
16:Species of bird
12:
11:
5:
2027:
2025:
2017:
2016:
2011:
2006:
2001:
1999:Birds of India
1996:
1991:
1986:
1981:
1976:
1966:
1965:
1959:
1958:
1956:
1955:
1942:
1929:
1916:
1903:
1890:
1877:
1864:
1851:
1838:
1825:
1815:
1802:
1789:
1780:Fauna Europaea
1776:
1763:
1750:
1737:
1724:
1714:
1701:
1688:
1675:
1662:
1651:
1636:
1620:
1618:
1612:
1611:
1606:
1600:
1599:
1594:
1583:
1582:External links
1580:
1578:
1577:
1554:
1535:
1516:(1): 131–133.
1500:
1473:
1454:(4): 573–576.
1431:
1405:
1386:(1): 113–123.
1370:
1361:American Birds
1351:
1332:
1306:
1271:
1262:American Birds
1252:
1222:
1195:
1168:
1142:
1120:
1094:
1068:
1045:
1006:
983:
968:
942:
915:
890:
867:
844:
821:
806:
778:
748:
725:
678:
676:
673:
672:
671:
662:
655:
653:
636:
629:
627:
620:
613:
611:
605:
598:
596:
587:
580:
576:
573:
527:
524:
486:
483:
482:
481:
475:
468:
466:
463:
456:
454:
448:
441:
439:
433:
426:
341:
338:
296:
293:
224:
223:
212:
211:
203:
202:
195:
184:
183:
177:
176:
169:
167:
163:
162:
155:
151:
150:
145:
141:
140:
138:Pelecaniformes
135:
131:
130:
125:
121:
120:
115:
111:
110:
105:
101:
100:
95:
91:
90:
77:
76:
58:
55:
54:
49:
46:
45:
35:
34:
26:
25:
15:
13:
10:
9:
6:
4:
3:
2:
2026:
2015:
2012:
2010:
2007:
2005:
2002:
2000:
1997:
1995:
1992:
1990:
1987:
1985:
1982:
1980:
1977:
1975:
1972:
1971:
1969:
1952:
1947:
1943:
1939:
1934:
1930:
1926:
1921:
1917:
1913:
1908:
1904:
1900:
1895:
1891:
1887:
1882:
1878:
1874:
1869:
1865:
1861:
1856:
1852:
1848:
1843:
1839:
1835:
1830:
1826:
1822:
1816:
1812:
1807:
1803:
1799:
1794:
1790:
1786:
1781:
1777:
1773:
1768:
1764:
1760:
1755:
1751:
1747:
1742:
1738:
1734:
1729:
1725:
1721:
1715:
1711:
1706:
1702:
1698:
1693:
1689:
1685:
1680:
1676:
1672:
1667:
1663:
1656:
1652:
1647:
1641:
1637:
1632:
1626:
1622:
1621:
1619:
1617:
1613:
1609:
1604:
1598:
1595:
1593:
1591:
1586:
1585:
1581:
1574:(2): 251–262.
1573:
1569:
1565:
1558:
1555:
1550:
1546:
1539:
1536:
1531:
1527:
1523:
1519:
1515:
1511:
1504:
1501:
1497:(2): 350–360.
1496:
1492:
1488:
1486:
1477:
1474:
1469:
1465:
1461:
1457:
1453:
1449:
1442:
1435:
1432:
1428:(3): 481–483.
1427:
1423:
1416:
1409:
1406:
1401:
1397:
1393:
1389:
1385:
1381:
1374:
1371:
1367:(5): 827–829.
1366:
1362:
1355:
1352:
1348:(4): 288–294.
1347:
1343:
1336:
1333:
1328:
1324:
1317:
1310:
1307:
1302:
1298:
1294:
1290:
1286:
1282:
1275:
1272:
1267:
1263:
1256:
1253:
1248:
1244:
1237:
1235:
1226:
1223:
1219:(1): 215–216.
1218:
1214:
1210:
1208:
1199:
1196:
1191:
1187:
1183:
1181:
1172:
1169:
1164:
1160:
1159:British Birds
1153:
1146:
1143:
1138:
1131:
1129:
1127:
1125:
1121:
1116:
1112:
1105:
1098:
1095:
1090:
1086:
1085:Dutch Birding
1079:
1072:
1069:
1064:
1060:
1056:
1049:
1046:
1042:(417): 24–26.
1041:
1037:
1033:
1031:
1027:
1026:E. schistacea
1023:
1019:
1010:
1007:
1002:
997:
996:
987:
984:
979:
972:
969:
964:
961:(in French).
960:
953:
946:
943:
939:(3): 494–497.
938:
934:
930:
928:
919:
916:
911:
904:
897:
895:
891:
887:(1): 188–190.
886:
882:
878:
871:
868:
863:
859:
855:
848:
845:
841:(3): 693–695.
840:
836:
832:
825:
822:
817:
810:
807:
802:
798:
797:British Birds
794:
787:
785:
783:
779:
774:
770:
766:
764:
755:
753:
749:
745:(1): 188–189.
744:
740:
736:
729:
726:
713:
708:
704:
700:
699:
694:
692:
683:
680:
674:
669:
665:
664:E. g. gularis
659:
654:
651:
647:
643:
639:
633:
628:
624:
617:
612:
608:
602:
597:
594:
590:
584:
579:
574:
572:
570:
566:
562:
557:
551:
549:
545:
541:
537:
533:
525:
523:
519:
517:
513:
508:
504:
500:
496:
492:
484:
479:
476:White morph,
472:
467:
460:
455:
452:
445:
440:
437:
430:
425:
423:
421:
417:
412:
408:
404:
400:
396:
392:
387:
383:
379:
375:
371:
367:
363:
359:
355:
351:
347:
346:Ardea gularis
339:
337:
335:
332:
328:
324:
320:
316:
311:
307:
302:
294:
292:
290:
286:
282:
278:
274:
270:
265:
264:E. g. gularis
261:
257:
253:
249:
245:
241:
237:
236:
231:
222:
218:
213:
209:
204:
199:
193:
191:
185:
182:
181:Binomial name
178:
174:
173:
168:
165:
164:
161:
160:
156:
153:
152:
149:
146:
143:
142:
139:
136:
133:
132:
129:
126:
123:
122:
119:
116:
113:
112:
109:
106:
103:
102:
99:
96:
93:
92:
87:
82:
78:
72:
67:
66:Least Concern
56:
52:
47:
44:
40:
36:
32:
27:
22:
19:
1615:
1589:
1571:
1567:
1557:
1548:
1544:
1538:
1513:
1509:
1503:
1494:
1490:
1484:
1476:
1451:
1447:
1434:
1425:
1421:
1408:
1383:
1379:
1373:
1364:
1360:
1354:
1345:
1341:
1335:
1326:
1322:
1309:
1287:(1): 70–73.
1284:
1280:
1274:
1265:
1261:
1255:
1246:
1242:
1233:
1225:
1216:
1212:
1206:
1198:
1189:
1185:
1179:
1171:
1162:
1158:
1145:
1136:
1114:
1110:
1097:
1088:
1084:
1071:
1062:
1058:
1048:
1039:
1035:
1029:
1025:
1021:
1017:
1009:
994:
986:
977:
971:
962:
958:
945:
936:
932:
926:
918:
909:
884:
880:
870:
861:
857:
853:
847:
838:
834:
824:
815:
809:
800:
796:
772:
768:
762:
742:
738:
728:
716:. Retrieved
702:
696:
690:
682:
663:
637:
606:
588:
552:
547:
529:
520:
516:Pulicat Lake
511:
499:Persian Gulf
488:
434:Dark morph,
419:
415:
410:
402:
398:
390:
386:cytochrome b
381:
378:superspecies
373:
369:
365:
361:
357:
353:
345:
343:
333:
326:
322:
318:
314:
306:little egret
298:
284:
280:
276:
272:
268:
263:
259:
256:Little egret
239:
234:
233:
229:
227:
216:
189:
187:
171:
170:
158:
42:
38:
18:
1829:iNaturalist
1640:Wikispecies
1551:(5): 73–74.
1249:(1): 46–48.
1117:(2): 71–80.
1030:E. dimorpha
718:11 November
569:Engraulidae
544:mudskippers
536:crustaceans
362:Lepterodius
354:Demiegretta
295:Description
41:dark morph
1968:Categories
1946:Xeno-canto
1165:: 382–386.
1091:: 238–246.
1065:: 189–196.
1022:E. gularis
965:: 241–242.
858:Sandgrouse
803:: 307–319.
675:References
512:schistacea
416:schistacea
411:schistacea
391:Ardea asha
327:schistacea
319:schistacea
289:subspecies
285:schistacea
269:schistacea
1192:(1): 275.
775:(1): 1–6.
561:Clupeidae
507:Sri Lanka
166:Species:
104:Kingdom:
98:Eukaryota
1873:22729692
1847:10586611
1710:22729692
1705:BirdLife
1625:Wikidata
1530:83867378
1342:Kingbird
1329:: 61–72.
1268:: 16–27.
864:: 75–81.
565:Gobiidae
540:molluscs
374:garzetta
358:Herodias
217:dimorpha
148:Ardeidae
144:Family:
118:Chordata
114:Phylum:
108:Animalia
94:Domain:
71:IUCN 3.1
1979:Egretta
1925:3596775
1886:1752861
1811:2480898
1692:Avibase
1631:Q392570
1468:6191096
1400:1521360
1301:1521198
1111:Podoces
929:(Bosc)"
646:Gujarat
575:Gallery
556:monsoon
495:Red Sea
451:Bahrain
436:Bahrain
366:Egretta
323:gularis
315:gularis
281:gularis
273:gularis
254:of the
200:, 1792)
159:Egretta
154:Genus:
134:Order:
124:Class:
69: (
1938:212665
1912:212665
1860:174822
1818:GNAB:
1767:EURING
1759:100728
1746:werher
1720:werher
1679:ARKive
1659:werehe
1528:
1466:
1398:
1299:
959:Alauda
910:Aquila
538:, and
497:, the
493:, the
491:Africa
478:Muscat
405:. The
219:, see
1933:WoRMS
1899:73453
1842:IRMNG
1785:96654
1754:EUNIS
1741:eBird
1733:6DZ3B
1717:BOW:
1526:S2CID
1464:S2CID
1444:(PDF)
1418:(PDF)
1396:JSTOR
1319:(PDF)
1297:JSTOR
1239:(PDF)
1155:(PDF)
1107:(PDF)
1081:(PDF)
955:(PDF)
906:(PDF)
650:India
623:Egypt
503:India
407:lores
310:egret
299:This
252:morph
248:slaty
244:heron
1907:OBIS
1881:NCBI
1868:IUCN
1855:ITIS
1834:4935
1806:GBIF
1772:1180
1028:and
720:2021
703:2016
593:MHNT
567:and
548:kwok
532:fish
360:and
301:bird
283:and
271:and
228:The
198:Bosc
128:Aves
1728:CoL
1666:ADW
1655:ABA
1518:doi
1456:doi
1452:165
1388:doi
1289:doi
1001:110
856:".
707:doi
422:).
393:by
287:as
1970::
1948::
1935::
1922::
1909::
1896::
1883::
1870::
1857::
1844::
1831::
1808::
1795::
1782::
1769::
1756::
1743::
1730::
1707::
1694::
1681::
1668::
1657::
1642::
1627::
1572:85
1570:.
1566:.
1547:.
1524:.
1514:54
1512:.
1495:84
1493:.
1489:.
1462:.
1450:.
1446:.
1426:15
1424:.
1420:.
1394:.
1384:15
1382:.
1365:37
1363:.
1346:57
1344:.
1327:19
1325:.
1321:.
1295:.
1283:.
1266:41
1264:.
1247:19
1245:.
1241:.
1217:65
1215:.
1211:.
1190:58
1188:.
1184:.
1163:94
1161:.
1157:.
1123:^
1113:.
1109:.
1089:39
1087:.
1083:.
1063:53
1061:.
1057:.
1040:59
1038:.
1034:.
1024:,
1020:,
963:79
957:.
937:78
935:.
931:.
908:.
893:^
885:54
883:.
879:.
860:.
839:81
837:.
833:.
801:88
799:.
795:.
781:^
773:84
771:.
767:.
751:^
743:81
741:.
737:.
701:.
695:.
648:,
644:,
591:-
563:,
534:,
291:.
1592:)
1549:1
1532:.
1520::
1470:.
1458::
1402:.
1390::
1303:.
1291::
1285:8
1115:5
1032:"
1003:.
862:5
722:.
709::
693:"
689:"
625:)
258:(
232:(
196:(
73:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
