266:, extinct armored jawless fish, to support evolution from arthropods to them and then to vertebrates. He also conducted other fieldwork. For example, in summer 1890 he was commissioned by the Botanical Division of the United States Department of Agriculture to collect specimens of grass from North Dakota for a project to determine if any were suitable for growing for grazing without irrigation. This assessment was undertaken by staff at the newly-established
282:
Labrador and the Baltic region. He became very skilled in revealing the fossilised remains from its stony matrix to see previously unknown fine structural details. His research supported his belief that ostracoderms were the ancestors of fish and linked invertebrates to vertebrates. Patten tended to work alone and rarely attended scientific conferences. He was a forceful lecturer.
31:
281:
to replace him. Patten continued to work on the phylogeny of ostracoderms, scorpions and primitive fish as well some a minor interests such as the embryology of molluscs and arthropods. He travelled worldwide during summer vacations to collect fossils, including to
Australia, Japan, Costa Rica, Cuba,
231:
with
Leuckart's group, developing a life-long interest in embryology and evolutionary relationships. His work was supported by a fellowship from Harvard for three year's research in Europe. He was awarded a doctoral degree in 1884. He then spent a year at the Zoological Station at Trieste and another
392:
Patten married
Elizabeth Merrill in 1883. She accompanied him on his fossil collecting expeditions. They had one child. Patten was noted for his singing in a local musical society while he was at University of North Dakota. He died in Hanover, New Hampshire on October 27, 1932.
769:
409:
in 1921. He was the chair and vice-president of the
Zoology section of the American Association for the Advancement of Science in 1919. He was awarded an honorary Doctor of Science degree when he retired by Dartmouth College.
320:
293:, to other disciplines. It was a mandatory course for all new students which was both innovative and quite controversial. At the time, some states in the USA did not allow evolution to be covered in
243:. While he was there, he hypothesised that vertebrates were derived from arachnid ancestors, and the exploration of this idea was important in his future research. Among others at the laboratory was
285:
Between 1920 and his retirement in 1931 he gave a lecture course on evolution to all new students at
Dartmouth College. The content was probably a development of his course in
188:
239:
Patten returned to the USA and between 1886 and 1889 acted as a research assistant at the Lake
Laboratory in Milwaukee, Wisconsin, the private research laboratory of
297:. In 1919 he addressed the annual meeting of American Association for the Advancement of Science about “The Message of the Biologist” and followed this with a book
247:
and Patten influenced him to study insect embryology and development using the cockroach as an example, which became an important part of his future career.
842:
847:
289:
that he started in 1898. This became a course where he and other members of the academic staff linked science, including evolution of humans and
405:, recognising excellence in the sciences and liberal arts from a small number of universities in the USA. Patten was elected as a Fellow of the
406:
219:. He graduated with a B.Sc. degree in 1883 and was awarded a Parker Travelling Fellowship. He then travelled to Europe and studied with
837:
832:
827:
419:
852:
212:
301:
He outlined a philosophy of science in service to society that invoked chemistry theory, mixed with evolution theory and ecology.
240:
495:
547:
233:
267:
812:
660:
316:
William Patten (1885) The
Embryology of Patella. Arbeiten aus dem Zoologischen Instituts zu Wein. Published Vienna.
251:
208:
172:
129:
153:
204:
54:
304:
However, by 1923 there was little opposition to the course and it was seen as a leading example of pedagogy.
402:
244:
72:
343:
William Patten (1912) The evolution of the vertebrates and their kin. P. Blakiston's Son & Company.
822:
817:
733:
694:
596:
228:
227:
as well as having some interest in insects, taking a morphological approach to both. Patten studied
751:
612:
463:
87:
83:
620:
365:
278:
274:
176:
133:
741:
702:
604:
528:
519:
Lauder, George V. (1981). "Edward Phelps Allis: discovery of his anatomical illustrations".
250:
In
October 1889 Patten was appointed the Professor of Biology and Curator of Museums at the
220:
358:
312:
Patten was the author of several books and many scientific publications. These included:
737:
698:
600:
270:
in Fargo. He also gave talks to local societies, including on the subject of evolution.
532:
418:
After his death in 1932, Patten's collection of ostracoderm fossils was donated to the
806:
770:"William Patten Dartmouth College, Zoologist, paleontologist, educator. elected 1921"
259:
755:
294:
224:
263:
180:
608:
706:
380:
290:
255:
30:
797:
624:
384:, the type genus of the ostracoderm family Dartmuthiidae was named by him.
184:
467:
451:
616:
584:
359:
The Grand
Strategy of Evolution: The Social Philosophy of a Biologist.
254:. His research focused on the origin of vertebrates, considering that
746:
721:
647:
The Grand
Strategy of Evolution: the Social Philosophy of a Biologist
299:
The Grand Strategy of Evolution: the Social Philosophy of a Biologist
207:
on March 15, 1861. He attended Harvard university (then known as the
193:
The Grand Strategy of Evolution: the Social Philosophy of a Biologist
140:
685:
Robertson, G. M. (1935). "The ostracoderm genus Dartmuthia Patten".
332:
William Patten (1890) On the Origin of Vertebrates from Arachnids.
277:
in New Hampshire, USA. The University of North Dakota appointed
258:
were more likely ancestors rather than the then-current idea of
223:
at Leipzig University in Germany. Leuckart specialised in
548:"Biographical Memoir of William Morton Wheeler 1865-1937"
456:
Proceedings of the American Academy of Arts and Sciences
497:
The Makers of Biology at the University of North Dakota
273:
In 1893 he was appointed as a professor at the private
171:(1861-1932) was an American biologist and zoologist at
346:William Patten (1920 The message of the biologist.
189:
American Association for the Advancement of Science
152:
139:
124:
109:
101:
93:
79:
61:
40:
21:
578:
576:
555:National Academy of Sciences Biographical Memoirs
445:
443:
441:
439:
437:
435:
378:He also described and named some fossil species.
489:
487:
485:
483:
481:
479:
477:
422:in New York. There were over 4,500 specimens.
8:
334:Quarterly Journal of Microscopical Sciences
665:Dartmouth Undergraduate Journal of Science
211:) and was awarded the Walker prize of the
183:, ideas on the origin of vertebrates from
29:
18:
745:
521:Biological Journal of the Linnean Society
215:during his first year for an essay about
431:
661:"The evolutionary debate at Dartmouth"
774:American Academy of Arts and Sciences
407:American Academy of Arts and Sciences
7:
97:ideas about evolution; ostracoderms
843:University of North Dakota faculty
533:10.1111/j.1095-8312.1981.tb01653.x
420:American Museum of Natural History
287:Comparative Anatomy of Vertebrates
14:
268:North Dakota Agricultural College
213:Boston Society of Natural History
848:American evolutionary biologists
321:Eyes of molluscs and arthropods.
217:Myology and osteology of the cat
262:. This led him to research the
585:"The message of the biologist"
546:Parker, George Howard (1938).
1:
154:Author abbrev. (zoology)
452:"William Patten (1861-1932)"
187:and his 1919 address to the
687:American Journal of Science
494:Kannowski, Paul B. (2015).
366:New Ostracoderms from Oesel
203:William Patten was born in
869:
609:10.1126/science.51.1309.93
450:Goldthwait, J. W. (1936).
252:University of North Dakota
209:Lawrence Scientific School
173:University of North Dakota
130:University of North Dakota
838:Dartmouth College faculty
833:Leipzig University alumni
828:Harvard University alumni
707:10.2475/ajs.s5-29.172.323
234:Naples Zoological Station
191:and follow-up book about
179:noted for work on fossil
162:
117:
28:
853:American paleontologists
645:Patten, William (1920).
583:Patten, William (1920).
205:Watertown, Massachusetts
199:Early life and education
55:Watertown, Massachusetts
401:He was a member of the
649:. Boston: R.G. Badger.
500:. University of Dakota
403:Phi Beta Kappa society
364:William Patten (1931)
357:William Patten (1920)
319:William Patten (1887)
245:William Morton Wheeler
73:Hanover, New Hampshire
324:Journal of Morphology
722:"Dr. William Patten"
295:children's education
229:marine invertebrates
813:American zoologists
738:1932Natur.130..873A
699:1935AmJS...29..323R
601:1920Sci....51...93P
720:W., A. S. (1932).
88:Leipzig University
84:Harvard University
16:American zoologist
397:Awards and honors
279:Melvin A. Brannon
275:Dartmouth College
177:Dartmouth College
166:
165:
134:Dartmouth College
128:Lake Laboratory;
119:Scientific career
105:Elizabeth Merrill
860:
785:
784:
782:
780:
766:
760:
759:
749:
747:10.1038/130873a0
717:
711:
710:
693:(172): 323–335.
682:
676:
675:
673:
671:
657:
651:
650:
642:
636:
635:
633:
631:
595:(1309): 93–102.
580:
571:
570:
568:
566:
552:
543:
537:
536:
516:
510:
509:
507:
505:
491:
472:
471:
447:
221:Rudolph Leuckart
148:
68:
65:October 27, 1932
50:
48:
33:
19:
868:
867:
863:
862:
861:
859:
858:
857:
803:
802:
794:
789:
788:
778:
776:
768:
767:
763:
719:
718:
714:
684:
683:
679:
669:
667:
659:
658:
654:
644:
643:
639:
629:
627:
582:
581:
574:
564:
562:
550:
545:
544:
540:
518:
517:
513:
503:
501:
493:
492:
475:
462:(10): 566–568.
449:
448:
433:
428:
416:
399:
390:
310:
291:Darwin's theory
201:
146:
80:Alma mater
75:
70:
66:
57:
52:
46:
44:
36:
24:
17:
12:
11:
5:
866:
864:
856:
855:
850:
845:
840:
835:
830:
825:
820:
815:
805:
804:
801:
800:
798:William Patten
793:
792:External links
790:
787:
786:
761:
712:
677:
652:
637:
572:
538:
527:(4): 283–291.
511:
473:
430:
429:
427:
424:
415:
412:
398:
395:
389:
386:
376:
375:
374:(1901):671-673
362:
355:
344:
341:
330:
317:
309:
306:
200:
197:
169:William Patten
164:
163:
160:
159:
156:
150:
149:
143:
137:
136:
126:
122:
121:
115:
114:
111:
107:
106:
103:
99:
98:
95:
94:Known for
91:
90:
81:
77:
76:
71:
69:(aged 71)
63:
59:
58:
53:
51:March 15, 1861
42:
38:
37:
35:William Patten
34:
26:
25:
23:William Patten
22:
15:
13:
10:
9:
6:
4:
3:
2:
865:
854:
851:
849:
846:
844:
841:
839:
836:
834:
831:
829:
826:
824:
821:
819:
816:
814:
811:
810:
808:
799:
796:
795:
791:
775:
771:
765:
762:
757:
753:
748:
743:
739:
735:
732:(3293): 873.
731:
727:
723:
716:
713:
708:
704:
700:
696:
692:
688:
681:
678:
666:
662:
656:
653:
648:
641:
638:
626:
622:
618:
614:
610:
606:
602:
598:
594:
590:
586:
579:
577:
573:
560:
556:
549:
542:
539:
534:
530:
526:
522:
515:
512:
499:
498:
490:
488:
486:
484:
482:
480:
478:
474:
469:
465:
461:
457:
453:
446:
444:
442:
440:
438:
436:
432:
425:
423:
421:
413:
411:
408:
404:
396:
394:
388:Personal life
387:
385:
383:
382:
373:
370:
367:
363:
360:
356:
354:(1309) 93-102
353:
349:
345:
342:
339:
335:
331:
328:
325:
322:
318:
315:
314:
313:
307:
305:
302:
300:
296:
292:
288:
283:
280:
276:
271:
269:
265:
261:
260:annelid worms
257:
253:
248:
246:
242:
237:
235:
230:
226:
222:
218:
214:
210:
206:
198:
196:
194:
190:
186:
182:
178:
174:
170:
161:
157:
155:
151:
144:
142:
138:
135:
131:
127:
123:
120:
116:
112:
108:
104:
100:
96:
92:
89:
85:
82:
78:
74:
64:
60:
56:
43:
39:
32:
27:
20:
777:. Retrieved
773:
764:
729:
725:
715:
690:
689:. Series 5.
686:
680:
668:. Retrieved
664:
655:
646:
640:
628:. Retrieved
592:
588:
563:. Retrieved
558:
554:
541:
524:
520:
514:
502:. Retrieved
496:
459:
455:
417:
400:
391:
379:
377:
371:
368:
351:
347:
340:(3):317-378.
337:
333:
326:
323:
311:
308:Publications
303:
298:
286:
284:
272:
264:ostracoderms
249:
241:Edward Allis
238:
225:parasitology
216:
202:
192:
181:ostracoderms
168:
167:
125:Institutions
118:
67:(1932-10-27)
823:1932 deaths
818:1861 births
779:25 February
670:25 February
630:26 February
565:25 February
504:29 December
807:Categories
426:References
381:Dartmuthia
361:RG Badger.
256:arthropods
47:1861-03-15
561:: 201–241
329:(1):67-92
185:arachnids
625:17798902
468:20023188
110:Children
756:4070074
734:Bibcode
695:Bibcode
617:1644674
597:Bibcode
589:Science
369:Science
348:Science
232:at the
754:
726:Nature
623:
615:
466:
414:Legacy
158:Patten
147:(1884)
145:
141:Thesis
102:Spouse
752:S2CID
613:JSTOR
551:(PDF)
464:JSTOR
781:2022
672:2022
632:2022
621:PMID
567:2022
506:2021
175:and
62:Died
41:Born
742:doi
730:130
703:doi
605:doi
529:doi
809::
772:.
750:.
740:.
728:.
724:.
701:.
691:29
663:.
619:.
611:.
603:.
593:51
591:.
587:.
575:^
559:19
557:.
553:.
525:16
523:.
476:^
460:70
458:.
454:.
434:^
372:73
352:51
350:.
338:31
336:.
236:.
195:.
132:;
86:;
783:.
758:.
744::
736::
709:.
705::
697::
674:.
634:.
607::
599::
569:.
535:.
531::
508:.
470:.
327:1
113:1
49:)
45:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.