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Xenophyophorea

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686: 825: 406: 63: 563:). Gametes are reportedly about 20 μm in diameter, with two flagella; after this, an amoeba-like stage seems to be present. It is also possible that the amoeboid stage represents amoeboid gametes, found in other foraminifera. These amoeboid structures are also sometimes found inside the granellare. Juveniles have occasionally been found in association with adults; in Psametta they are horseshoe-shaped and already covered in xenophyae. 3197: 31: 993: 3185: 584: 45: 576:. This growth occurred in phases lasting 2–3 days each; each phase was separated by a resting period of approximately two months. These growth phases were approximately synchronous between specimens, but it is unclear if this is biologically or developmentally controlled; some evidence suggests the synchrony may have been due to chance. 566:
The location of the initial plasma can sometimes be pointed out in adult xenophyophores. In some species this is denoted by a sharp change in the type of xenophyae; in others, the juvenile is regular and the adult is irregular; still others flip this pattern, so that the juvenile is irregular and the
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species, each 3–6 mm. The fragility of the xenophyophores suggests that the urchin either very carefully collected them, or that they settled and grew there. Among several possible explanations for the urchin's behaviour, perhaps the most likely are chemical camouflage and weighing itself down
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to the fossil. Supporting this notion is the similar abyssal habitat of living xenophyophores to the inferred habitat of fossil graphoglyptids; however, the large size (up to 0.5m) and regularity of many graphoglyptids as well as the apparent absence of xenophyae in their fossils casts doubt on the
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Each episode of growth occurred in three phases: first, the base becomes wider and flatter, causing the surface texture to become smoother; then, the original shape of the organism is regained (albeit larger); and finally, the surface texture is rebuilt. The rapid rate of growth observed suggests
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Xenophyophores are an important part of the deep sea-floor, as they have been found in all four major ocean basins. They are often found in areas of enhanced organic carbon flux, such as beneath productive surface waters, in sub-marine canyons, in settings with sloped topography (e.g. seamounts,
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and using the test to trap particles. Analysis of lipid concentrations within xenophyophores revealed especially high concentrations of bacteria in the stercomata, suggesting that xenophyophores utilise bacteria growing on their waste products in order to supplement their feeding.
425:) and other foreign mineral particles glued together with organic cements—that range from a few millimetres to 20 centimetres across. The softness and structure of tests varies from soft and lumpy shapes to fans and complex structures. 2813:
Rona, Peter A.; Seilacher, Adolf; de Vargas, Colomban; Gooday, Andrew J.; Bernhard, Joan M.; Bowser, Sam; Vetriani, Costantino; Wirsen, Carl O.; Mullineaux, Lauren; Sherrell, Robert; Frederick Grassle, J. (2009-09-01).
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Rothe, N.; Gooday, A. J.; Pearce, R. B. (2011-12-01). "Intracellular mineral grains in the xenophyophore Nazareammina tenera (Rhizaria, Foraminifera) from the Nazaré Canyon (Portuguese margin, NE Atlantic)".
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Meyer, Mike; Elliott, David; Wood, Andrew D.; Polys, Nicholas F.; Colbert, Matthew; Maisano, Jessica A.; Vickers-Rich, Patricia; Hall, Michael; Hoffman, Karl H.; Schneider, Gabi; Xiao, Shuhai (2014-08-01).
840:", providing habitat and serving as traps for organic particles, increasing diversity in the surrounding area. Research has shown that areas dominated by xenophyophores have 3–4 times the number of benthic 483:
are long (several mm in length), threadlike structures found outside of the granellare in some xenophyophores (genera traditionally grouped together as "stannomida"); they are flexible and form part of the
889:(Ophiuroidea) also appear to have a relationship with xenophyophores, as they are consistently found directly underneath or on top of the protozoans. They can also function as nurseries for fish; 836:
Local population densities may be as high as 2,000 individuals per 100 square metres (1,100 sq ft), making them dominant organisms in some areas. Xenophyophores have been found to be "
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Species of this group are morphologically variable, but the general structural pattern includes a test enclosing a branching system of organic tubules together with masses of waste material.
1493: 2347:"Rapid response of the giant protist xenophyophores (Foraminifera, Rhizaria) to organic matter supply at abyssal depths revealed by an in situ dual stable isotope labeling experiment" 642:, bacterial farming, deposit feeding, and trapping particulate matter inside the test. Studies have since confirmed active uptake of food from surrounding sediments using the 490:, for which the group is named, are the agglutinated particles from which the test is constructed. They vary by species; they can contain sediment particles, sponge spicules, 3150: 761:
However, analysis of the latter found neither barite crystals nor evidence of agglutinated foraminifera in the wall. A 2011 study that examined growth and development of
1326:"Five new species and two new genera of xenophyophores (Foraminifera: Rhizaria) from part of the abyssal equatorial Pacific licensed for polymetallic nodule exploration" 1813: 927:. As they occur in all the world's oceans and in great numbers, xenophyophores could be indispensable agents in the process of sediment deposition and in maintaining 798:
have been discovered; however, no evidence of tests, stercomares, grannelares, or xenophyophore DNA was found, and the trace may alternately represent a burrow or a
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Xenophyophores are difficult to study due to their extreme fragility. Specimens are invariably damaged during sampling, rendering them useless for captive study or
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Gooday, A. J.; Aranda da Silva, A.; Pawlowski, J. (2011-12-01). "Xenophyophores (Rhizaria, Foraminifera) from the Nazaré Canyon (Portuguese margin, NE Atlantic)".
4392: 3090:"Xenophyophores (Rhizaria, Foraminifera), including four new species and two new genera, from the western Clarion-Clipperton Zone (abyssal equatorial Pacific)" 1920:"Micro-CT 3D imaging reveals the internal structure of three abyssal xenophyophore species (Protista, Foraminifera) from the eastern equatorial Pacific Ocean" 1703:"Giant protists (xenophyophores, Foraminifera) are exceptionally diverse in parts of the abyssal eastern Pacific licensed for polymetallic nodule exploration" 1368:
Gooday, A.J; Aranda da Silva, A.; Pawlowski, J. (2011). "Xenophyophores (Rhizaria, Foraminifera) from the Nazare Canyon (Portuguese margin, NE Atlantic)".
954:, about 5 cm diameter, which was wearing a cloak consisting of over 1,000 protists and other creatures, including 245 living xenophyophores, mainly 666:. This study found no evidence to support a bacterial farming function for the test, and instead proposed that it aided to function in the collection of 3040:"Intracellular mineral grains in the xenophyophore Nazareammina tenera (Rhizaria, Foraminifera) from the Nazaré Canyon (Portuguese margin, NE Atlantic)" 2662:"Jensen, S. and Palacios, T. 2006. A peri-Gondwanan cradle for the trace fossil Paleodictyon. In: 22 Jornadas de Paleontologia, Comunicaciones, 132-134" 916:
Despite this abundance, the relatively low amount of protoplasm per unit of test means that xenophyophores often contribute little to total biomass.
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Hayward, B.W.; Le Coze, F.; Gross, O. (2019). World Foraminifera Database. Monothalamea. Accessed through: World Register of Marine Species at:
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Gooday, Andrew J; Holzmann, Maria; Caulle, Clémence; Goineau, Aurélie; Kamenskaya, Olga; Weber, Alexandra A. -T.; Pawlowski, Jan (2017-03-01).
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The diet and feeding ecology of xenophyophores was long the subject of speculation; the fragile tests and deepwater habitat of the group makes
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Gooday, Andrew J.; Holzmann, Maria; Goineau, Aurélie; Pearce, Richard B.; Voltski, Ivan; Weber, Alexandra A.-T.; Pawlowski, Jan (2018-08-03).
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Levin, Lisa A.; Gooday, Andrew J.; James, David W. (2001). "Dressing up for the deep: agglutinated protists adorn an irregular urchin".
2615:"Three-dimensional microCT analysis of the Ediacara fossil Pteridinium simplex sheds new light on its ecology and phylogenetic affinity" 513:, or concentrated in excretions. The selected minerals vary with species, but often include barite, lead and uranium. The granellare of 3140: 1541:"A new genus of xenophyophores (Foraminifera) from Japan Trench: morphological description, molecular phylogeny and elemental analysis" 341:. In the beginning of the 20th century they were considered an independent class of Rhizopoda, and later as a new eukaryotic phylum of 3229: 609:
Xenophyophores have been found between depths of 500 and 10,600 metres. Most are epifaunal (living atop the seabed), but one species (
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Growth is episodic; one observational study taking place over a period of eight months saw a three-to-tenfold growth in specimens of
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Bobrovskiy, Ilya; Hope, Janet M.; Ivantsov, Andrey; Nettersheim, Benjamin J.; Hallmann, Christian; Brocks, Jochen J. (2018-09-21).
2141:"Interactions between metazoans and large, agglutinating protozoans: implications for the community structure of deep-sea benthos" 4562: 3263: 3258: 812:
within the fossils as well as supposed morphological similarity; however, the barium content was later determined to be due to
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cast doubt on this identification, as these sterols are today associated only with animals. These researchers suggest that
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through recent times, could represent the remains of xenophyophores, and noted the similarity of the extant xenophyophore
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Brady, H.B. (1884). "Report on the Foraminifera. Report on the Scientific Results of the Voyage of H. M. S. Challenger".
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A deep sea community of organisms, including several xenophyophores; the two large individuals in the bottom middle have
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Studies have found unusually high concentrations of radioactive nuclides in xenophyophores; this was first reported in
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Haeckel, E. (1889). "Report on the scientific results of the voyage of H. M. S. Challenger during the years 1873–76".
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Xenophyophore densities are highest on soft sediments; however, they may still be found on rocky substrates including
2267:"A new infaunal xenophyophore (xenophyophorea, protozoa) with notes on its ecology and possible trace fossil analogs" 2901:"Deep-sea benthic megafaunal habitat suitability modelling: A global-scale maximum entropy model for xenophyophores" 685: 378:
of xenophyophores. This study also suggested that many individual genera are polyphyletic, with similar body shapes
4397: 2174:"Synoptic checklist and bibliography of the Xenophyophorea (Protista), with a zoogeopgraphical survey of the group" 541: 2126:
Possible roles for Xenophyophores in dee-sea carbon cycling. In: Deep-Sea Food Chains and the Global Carbon Cycle
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Different xenophyophore ecomorphs are found in different settings; reticulated or heavily folded genera such as
3141:"The offshore directory: Review of a selection of habitats, communities and species of the north-east Atlantic" 659: 345:. Phylogenetic studies suggest that xenophyophores are a specialized group of monothalamous (single-chambered) 2822:. Marine Benthic Ecology and Biodiversity: A Compilation of Recent Advances in Honor of J. Frederick Grassle. 974:
are more common in areas where the substrate is sloped or near canyon walls, while more fan-shaped forms like
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but has since been found to be true of many other xenophyophore species from different parts of the ocean.
4600: 4319: 3222: 4595: 3310: 2669: 1494:"Small subunit ribosomal DNA suggests that the xenophyophorean Syringammina corbicula isa Foraminiferan" 1412: 924: 148: 4249: 4168: 3088:
Gooday, Andrew J.; Durden, Jennifer M.; Holzmann, Maria; Pawlowski, Jan; Smith, Craig R. (2020-08-01).
615:), is known to be infaunal; it buries itself up to 6 centimetres (2.4 in) deep into the sediment. 2346: 1749:"Testing the protozoan hypothesis for Ediacaran fossils: a developmental analysis of Palaeopascichnus" 670:
by increasing surface area. These authors argued that xenophyophores fill a major role in ocean-floor
188: 4567: 4510: 4471: 4407: 4366: 4151: 3974: 3051: 3039: 2912: 2827: 2815: 2771: 2709: 2626: 2614: 2460: 2405: 2358: 2309: 2237: 2200:"Xenophyophoria (Rhizopoda, Protozoa) in bottom photographs from the bathyal and abyssal NE Atlantic" 2098: 1996: 1931: 1892: 1628: 1616: 1589: 1377: 928: 775: 671: 379: 269: 4254: 4163: 4157: 3964: 3735: 3610: 3452: 1245: 509:
They select certain minerals and elements from their environment that are included in its tests and
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Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition Auf dem Dampfer 'Validivia' 1898–1899
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alteration of the material and the morphology of the specimen instead supported an algal affinity.
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fossils have been suggested to represent the remains of xenophyophores due to the concentration of
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Gooday, Andrew J.; Sykes, Dan; Góral, Tomasz; Zubkov, Mikhail V.; Glover, Adrian G. (2018-08-14).
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sea cucumbers have all been observed feeding on xenophyophores; specifically, the monoplacophoran
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Gametes form in a specialised part of the granellare that may look like swollen side-branch (in
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that extract minerals from their surroundings and use them to form an exoskeleton known as a
4420: 4285: 4102: 3881: 3638: 3252: 3101: 3059: 3004: 2967: 2951: 2920: 2870: 2835: 2779: 2717: 2634: 2576: 2527: 2478: 2468: 2449:"Ancient steroids establish the Ediacaran fossil Dickinsonia as one of the earliest animals" 2413: 2366: 2317: 2245: 2152: 2106: 2051: 2004: 1955: 1939: 1900: 1849: 1760: 1714: 1636: 1597: 1552: 1505: 1385: 1337: 733: 623: 208: 4541: 4384: 4371: 4289: 3826: 3163: 2697: 2682: 1425: 885:—some of which may take up semi-permanent residence within a xenophyophore's test. 3055: 2916: 2831: 2775: 2713: 2630: 2464: 2409: 2362: 2313: 2297: 2241: 2102: 2000: 1935: 1896: 1632: 1593: 1381: 852:
than equivalent areas that lack xenophyophores. The xenophyophores themselves also play
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Lecroq, Béatrice; Gooday, Andrew John; Tsuchiya, Masashi; Pawlowski, Jan (2009-07-01).
1509: 943: 900: 418: 326: 322: 292: 281: 159: 100: 87: 393:. However, cladistic analyses based on molecular data have suggested a high amount of 4584: 4476: 4145: 4117: 3933: 3730: 3630: 3444: 3429: 2598: 2564: 2433: 2110: 2071: 2039: 1765: 1748: 1557: 1540: 879: 805: 799: 667: 592:
sp. from the Porcupine Abyssal Plain in the NE Atlantic, from a depth of about 4800m.
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Schulze, F. E. (1907). "Die Xenophyophoren, eine besondere Gruppe der Rhizopoden".
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Torres, Andrew M. (1996). "Fossil algae were very different from xenophyophores".
276:, at depths of 500 to 10,600 metres (1,600 to 34,800 ft). They are a kind of 4447: 1719: 1702: 397:, and that the division between psamminids and stannomids is not well supported. 4523: 4456: 4351: 4140: 4135: 4002: 3997: 3984: 3876: 3836: 3434: 2721: 1853: 1584:. The Geology, Geochemistry, and Biology of Submarine Canyons West of Portugal. 1342: 1306:
A Monograph of the Xenophyophoria (Rhizopodea, Protozoa) (Doctoral dissertation)
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As of 2017, no positively-identified xenophyophore fossils had been discovered.
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to examine the question of xenophyophore feeding confirmed rapid uptake of both
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are more common in areas with quieter water and/or lower primary productivity.
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A number of unique terms are used to refer to anatomical aspects of the group:
4277: 4203: 4087: 3992: 3951: 3941: 3923: 3871: 3846: 3841: 3673: 3643: 3597: 3579: 3569: 3063: 3008: 2924: 2581: 2370: 2087:"Direct observation of episodic growth in an abyssal xenophyophore (Protista)" 2055: 2040:"Natural 226Ra and 232Th radionuclides in xenophyophores of the Pacific Ocean" 1904: 1841: 1217: 1180: 947: 871: 845: 813: 724: 583: 506:
of xenophyophores contributes less than 1% of the total mass of the organism.
503: 491: 450: 338: 4342: 3115: 3071: 3016: 2963: 2882: 2847: 2791: 2729: 2646: 2590: 2541: 2492: 2425: 2378: 2331: 2282: 2226:"Effect of giant protozoans (class: Xenophyophorea) on deep-seamount benthos" 2063: 2016: 1951: 1840:
Levin, L. A.; Gooday, A. J. (1992), Rowe, Gilbert T.; Pariente, Vita (eds.),
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has been suggested as a fossil xenophyophore, but this remains controversial.
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Very little is known about xenophyophore reproduction. It is assumed that an
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London, Postgraduate Unit of Micropalaeontology, University College (2002).
1617:"New supraordinal classification of Foraminifera: Molecules meet morphology" 1235:
Tendal, O.S. (1972) A MONOGRAPH OF THE XENOPHYOPHORIA (Rhizopodea, Protozoa)
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Ashford, Oliver S.; Davies, Andrew J.; Jones, Daniel O.B. (December 2014).
2737: 2500: 2157: 2140: 1985:"High levels of natural radionuclides in a deep-sea infaunal xenophyophore" 1969: 1517: 754:
have been posited as fossil xenophyophores and linked to the Eocene fossil
44: 3207: 2516:"Ediacaran biota: The dawn of animal life in the shadow of giant protists" 1984: 1265: 4441: 4336: 4297: 4293: 4191: 3803: 3778: 3514: 3369: 3322: 3238: 1269: 1046: 896: 875: 784: 699: 602:
abyssal hills) and on continental slopes. They are not found in areas of
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that xenophyophores may not be as long-lived as previously hypothesised.
458:. Stercomares also include small, yellow-red spherical bodies known as 4281: 3907: 2799: 2532: 2515: 2483: 2322: 2298:"Use of lipids to study the trophic ecology of deep-sea xenophyophores" 2250: 2225: 882: 857: 849: 663: 454:; pellets that are bundled together in long strings are referred to as 342: 300: 110: 4528: 2124:
Levin, L. A.; Gooday, A. J. (1992). Rowe, G. T.; Pariente, V. (eds.).
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Antcliffe, Jonathan B.; Gooday, Andrew J.; Brasier, Martin D. (2011).
1481:(2nd ed.). Society of protozoologists. Lawrence, KS: Allen Press. 893:
have been found to lay eggs in the shelter of the xenophyophore test.
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Levin, Lisa A. (1994). "Paleoecology and Ecology of Xenophyophores".
2417: 2008: 1848:, NATO ASI Series, Dordrecht: Springer Netherlands, pp. 93–104, 809: 710: 655: 619: 466: 4313: 2783: 2514:
Seilacher, Adolf; Grazhdankin, Dmitri; Legouta, Anton (2003-03-31).
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Levin, L. A.; DeMaster, D. J.; McCann, L. D.; Thomas, C. L. (1986).
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Historically xenophyophores have been divided into the agglutinated
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period represent fossil xenophyophores. However, the discovery of C
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Pawlowski, J.; Holzmann, M.; Fahrni, J.; Richardson, S.L. (2003).
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The name Xenophyophora means "bearer of foreign bodies", from the
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http://www.marinespecies.org/aphia.php?p=taxdetails&id=744106
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Pawlowski, Jan; Holzmann, Maria; Tyszka, Jarosław (2013-04-01).
4317: 3211: 434:—have compartmentalized tests consisting of multiple chambers. 1842:"Possible Roles for Xenophyophores in Deep-Sea Carbon Cycling" 987: 250: 2816:"Paleodictyon nodosum: A living fossil on the deep-sea floor" 765:
concluded it was likely not a xenophyophore. A 2014 study of
2940:"Giant protists (xenophyophores) function as fish nurseries" 473:
within their cytoplasm. This is not to be confused with the
2565:"Benkovac Stone (Eocene, Croatia): a deep-sea Plattenkalk?" 253: 244: 235: 232: 223: 217: 2820:
Deep Sea Research Part II: Topical Studies in Oceanography
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Deep-Sea Research Part II: Topical Studies in Oceanography
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has been proposed as a specialised predator of the group.
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have been described, varying widely in size. The largest,
3151:"Windows to the deep exploration: Giants of the protozoa" 313:, reaching up to 20 centimetres (8 in) in diameter. 3139:
Gubbay, S., Baker, M., Bettn, B., Konnecker, G. (2002).
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Swinbanks, David D.; Shirayama, Yoshihisa (March 1986).
295:, and in some regions are the dominant species. Fifteen 3044:
Deep Sea Research Part I: Oceanographic Research Papers
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Deep Sea Research Part I: Oceanographic Research Papers
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Deep Sea Research Part I: Oceanographic Research Papers
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Tendal, Os; Swinbanks, Dd; Shirayama, Y. (1982-01-01).
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Deep Sea Research Part I: Oceanographic Research Papers
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Gooday, A.J.; Bett, B.J.; Pratt, D.N. (November 1993).
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Deep-Sea Research Part I: Oceanographic Research Papers
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Xenophyophores also commonly have abundant crystals of
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More xenophyophore photos, with a map of their habitat
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Rothe, N.; Gooday, A. J.; Pearce, R. B. (2011-12-01).
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Journal of the Marine Biological Association of the UK
1299: 1297: 1295: 1293: 1291: 1289: 1287: 1285: 3173: 2698:"Piaeodicton: The Traces of Infaunal Xenophyophores?" 241: 229: 226: 220: 214: 211: 2128:. The Netherlands: Kluwer Academic. pp. 93–104. 247: 4431: 4326: 4228: 4177: 4116: 4077: 4039: 4028: 3983: 3950: 3932: 3921: 3904: 3859: 3817: 3794: 3787: 3776: 3664: 3629: 3596: 3550: 3539: 3513: 3492: 3485: 3474: 3443: 3410: 3389: 3379: 3366: 1363: 1361: 774:Some researchers have suggested that the enigmatic 638:observation difficult. Early propositions included 238: 2563:Seilacher, Adolf; Mrinjek, Erwin (December 2011). 2345:Tsuchiya, Masashi; Nomaki, Hidetaka (2021-10-01). 1266:"Giant amoebas discovered in deepest ocean trench" 519:have been found to contain high concentrations of 1477:Lee, J. J.; Leedale, G. F.; Bradbury, P. (2000). 923:. For this reason, very little is known of their 417:. Many form delicate and elaborate agglutinated 2398:Geological Society, London, Special Publications 2296:Laureillard, J; Méjanelle, L; Sibuet, M (2004). 1846:Deep-Sea Food Chains and the Global Carbon Cycle 368:to form a monophyletic clade closely related to 272:found on the ocean floor throughout the world's 477:, which refers to the plasma body and its tube. 374:. Further molecular evidence has confirmed the 1256: 1254: 413:Xenophyophores are unicellular, but have many 3223: 421:—shells often made of calcium carbonate (CaCO 8: 1812:: CS1 maint: multiple names: authors list ( 856:host to a number of organisms—such as 444:Individual waste pellets are referred to as 4314: 4036: 4025: 3929: 3918: 3914: 3901: 3791: 3784: 3773: 3661: 3626: 3593: 3557: 3547: 3536: 3489: 3482: 3471: 3386: 3376: 3363: 3230: 3216: 3208: 938:at a depth of 3,088 metres at the Alaskan 698:It has been suggested that the mysterious 43: 29: 20: 3105: 2971: 2580: 2531: 2482: 2472: 2321: 2249: 2156: 1959: 1764: 1718: 1556: 1545:Zoological Journal of the Linnean Society 1341: 1330:Zoological Journal of the Linnean Society 823: 548:; however, this has not been confirmed. 3180: 2938:Levin, Lisa A.; Rouse, Greg W. (2020). 1228: 2678: 2667: 1805: 1696: 1694: 1692: 1690: 1688: 1686: 1684: 1682: 1680: 1678: 1421: 1410: 3164:Microscopy-UK: Note on Xenophyophores 3083: 3081: 2894: 2892: 2757: 2755: 2198:Tendal, O. S.; Gooday, A. J. (1981). 1835: 1833: 1831: 1829: 1827: 1825: 1823: 1742: 1740: 1738: 1676: 1674: 1672: 1670: 1668: 1666: 1664: 1662: 1660: 1658: 1479:The illustrated guide to the protozoa 494:tests, and even the tests of smaller 7: 4408:bcb29775-1da9-46a3-a223-75a28b94e988 1319: 1317: 1315: 337:. Later they were placed within the 52:Xenophyophore at the Galapagos Rift 731:. Other ediacaran fossils, such as 2875:10.1111/j.1502-3931.1996.tb01661.x 1510:10.1111/j.1550-7408.2003.tb00275.x 1498:Journal of Eukaryotic Microbiology 959:to avoid being moved in currents. 38:Image of a deep sea xenophyophore 14: 3195: 3183: 3094:European Journal of Protistology 1766:10.1111/j.1475-4983.2011.01058.x 1558:10.1111/j.1096-3642.2008.00493.x 991: 794:possibility. Modern examples of 409:A large 20-cm wide xenophyophore 389:and the flexible, proteinaceous 207: 61: 16:Clade of single-celled organisms 2696:Swinbanks, D. D. (1982-10-01). 2639:10.1016/j.precamres.2014.04.013 713:associated with the fossils of 2302:Marine Ecology Progress Series 2230:Marine Ecology Progress Series 1641:10.1016/j.marmicro.2013.04.002 934:Scientists in the submersible 291:in 1883. They are abundant on 1: 771:reached similar conclusions. 352:A 2013 molecular study using 333:classified them as primitive 309:, is among the largest known 287:They were first described by 2569:Swiss Journal of Geosciences 2392:Seilacher, A. (2007-01-01). 2111:10.1016/0967-0637(93)90094-J 2038:Domanov, M. M. (July 2015). 1720:10.1016/j.biocon.2017.01.006 428:Some xenophyophores—notably 2722:10.1126/science.218.4567.47 2394:"The nature of vendobionts" 1854:10.1007/978-94-011-2452-2_6 650:A 2021 study that utilised 4617: 3107:10.1016/j.ejop.2020.125715 2840:10.1016/j.dsr2.2009.05.015 2044:Geochemistry International 1944:10.1038/s41598-018-30186-2 1602:10.1016/j.dsr2.2011.04.005 1390:10.1016/j.dsr2.2011.04.005 723:and relatives are instead 573:Reticulammina labyrinthica 542:alternation of generations 4263: 4024: 3917: 3900: 3772: 3660: 3592: 3560: 3535: 3470: 3362: 3246: 3064:10.1016/j.dsr.2011.09.003 3009:10.1017/S0025315401004738 2925:10.1016/j.dsr.2014.07.012 2582:10.1007/s00015-011-0051-7 2371:10.1016/j.dsr.2021.103608 2056:10.1134/S0016702915070034 1905:10.1016/j.dsr.2011.09.003 1370:Deep-Sea Research Part II 1343:10.1093/zoolinnean/zlx093 544:takes place, as in other 317:Naming and classification 306:Syringammina fragilissima 166: 156: 58:Scientific classification 56: 51: 42: 37: 28: 23: 2520:Paleontological Research 1621:Marine Micropaleontology 660:dissolved organic matter 557:) or a stalked bulb (in 2474:10.1126/science.aat7228 1707:Biological Conservation 1308:. Danish Science Press. 931:in benthic ecosystems. 783:, known from the early 536:Growth and reproduction 2677:Cite journal requires 2172:Tendal, O. S. (1996). 1420:Cite journal requires 1304:Tendal, O. S. (1972). 833: 695: 672:biogeochemical cycling 593: 410: 371:Rhizammina algaeformis 827: 688: 612:Occultammina profunda 586: 408: 380:convergently evolving 270:unicellular organisms 265:. Xenophyophores are 185:Clade incertae sedis 4591:Foraminifera classes 2619:Precambrian Research 2158:10.1093/icb/31.6.886 2139:Levin, L. A (1991). 2097:(11–12): 2131–2143. 1588:(23–24): 2401–2419. 1376:(23–24): 2401–2419. 1005:adding missing items 929:biological diversity 622:, canyon walls, and 4276:organization type: 3056:2011DSRI...58.1189R 2917:2014DSRI...94...31A 2832:2009DSRII..56.1700R 2776:1994Palai...9...32L 2714:1982Sci...218...47S 2631:2014PreR..249...79M 2465:2018Sci...361.1246B 2459:(6408): 1246–1249. 2410:2007GSLSP.286..387S 2363:2021DSRI..17603608T 2314:2004MEPS..270..129L 2242:1986MEPS...29...99L 2103:1993DSRI...40.2131G 2001:1986Natur.320..354S 1936:2018NatSR...812103G 1897:2011DSRI...58.1189R 1633:2013MarMP.100....1P 1594:2011DSRII..58.2401G 1382:2011DSRII..58.2401G 1264:(22 October 2011). 910:Neopilina galatheae 838:ecosystem engineers 157:Orders and subtaxa 3910:    3906:    2533:10.2517/prpsj.7.43 2323:10.3354/meps270129 2251:10.3354/meps029099 2145:American Zoologist 1924:Scientific Reports 1003:; you can help by 952:Cystochinus loveni 940:continental margin 834: 696: 640:suspension feeding 594: 567:adult is regular. 411: 354:small subunit rDNA 331:Henry Bowman Brady 289:Henry Bowman Brady 4578: 4577: 4550:Open Tree of Life 4416:Open Tree of Life 4320:Taxon identifiers 4311: 4310: 4294:heliozoan amoebae 4286:amoeboflagellates 4250:Labyrinthomyxidae 4224: 4223: 4220: 4219: 4216: 4215: 4212: 4211: 4169:Globigerinitoidea 4020: 4019: 4016: 4015: 3896: 3895: 3892: 3891: 3855: 3854: 3768: 3767: 3764: 3763: 3760: 3759: 3756: 3755: 3752: 3751: 3748: 3747: 3744: 3743: 3679:Phaeogymnocellida 3656: 3655: 3652: 3651: 3588: 3587: 3575:Paracercomonadida 3531: 3530: 3466: 3465: 3143:, pp. 74–77. 3050:(12): 1189–1195. 2826:(19): 1700–1712. 2271:Oceanologica Acta 2207:Oceanologica Acta 1995:(6060): 354–358. 1891:(12): 1189–1195. 1863:978-94-011-2452-2 1021: 1020: 948:spatangoid urchin 652:isotopic labeling 597:Habitat and range 201: 200: 193: 182: 174: 152: 4608: 4571: 4570: 4558: 4557: 4545: 4544: 4532: 4531: 4519: 4518: 4506: 4505: 4493: 4492: 4480: 4479: 4467: 4466: 4465: 4452: 4451: 4450: 4424: 4423: 4411: 4410: 4401: 4400: 4388: 4387: 4375: 4374: 4362: 4361: 4360: 4347: 4346: 4345: 4315: 4103:Silicoloculinida 4037: 4033: 4026: 3930: 3926: 3919: 3915: 3911: 3902: 3882:Plasmodiophorida 3792: 3785: 3781: 3774: 3662: 3639:Thaumatomonadida 3627: 3594: 3558: 3548: 3544: 3537: 3490: 3483: 3479: 3472: 3387: 3377: 3373: 3364: 3275: 3255: 3232: 3225: 3218: 3209: 3200: 3199: 3188: 3187: 3186: 3179: 3149:Ocean Explorer. 3128: 3127: 3109: 3085: 3076: 3075: 3035: 3029: 3028: 2992: 2986: 2985: 2975: 2956:10.1002/ecy.2933 2935: 2929: 2928: 2896: 2887: 2886: 2858: 2852: 2851: 2810: 2804: 2803: 2759: 2750: 2749: 2693: 2687: 2686: 2680: 2675: 2673: 2665: 2657: 2651: 2650: 2609: 2603: 2602: 2584: 2560: 2554: 2553: 2535: 2511: 2505: 2504: 2486: 2476: 2444: 2438: 2437: 2418:10.1144/SP286.28 2389: 2383: 2382: 2342: 2336: 2335: 2325: 2293: 2287: 2286: 2262: 2256: 2255: 2253: 2221: 2215: 2214: 2204: 2195: 2189: 2188: 2178: 2169: 2163: 2162: 2160: 2136: 2130: 2129: 2121: 2115: 2114: 2082: 2076: 2075: 2035: 2029: 2028: 2009:10.1038/320354a0 1980: 1974: 1973: 1963: 1915: 1909: 1908: 1879: 1873: 1872: 1871: 1870: 1837: 1818: 1817: 1811: 1803: 1801: 1800: 1785: 1779: 1778: 1768: 1759:(5): 1157–1175. 1744: 1733: 1732: 1722: 1698: 1653: 1652: 1612: 1606: 1605: 1577: 1571: 1570: 1560: 1536: 1530: 1529: 1489: 1483: 1482: 1474: 1468: 1467: 1455: 1449: 1448: 1436: 1430: 1429: 1423: 1418: 1416: 1408: 1400: 1394: 1393: 1365: 1356: 1355: 1345: 1321: 1310: 1309: 1301: 1280: 1279: 1277: 1276: 1258: 1249: 1242: 1236: 1233: 1016: 1013: 995: 994: 988: 901:monoplacophorans 763:Palaeopascichnus 734:Palaeopascichnus 382:multiple times. 260: 259: 256: 255: 252: 249: 246: 243: 240: 237: 234: 231: 228: 225: 222: 219: 216: 213: 191: 189:Ammoclathrinidae 180: 172: 147: 66: 65: 47: 33: 21: 4616: 4615: 4611: 4610: 4609: 4607: 4606: 4605: 4581: 4580: 4579: 4574: 4566: 4561: 4553: 4548: 4540: 4535: 4527: 4522: 4514: 4509: 4501: 4496: 4488: 4483: 4475: 4470: 4461: 4460: 4455: 4446: 4445: 4440: 4427: 4419: 4414: 4406: 4404: 4396: 4391: 4383: 4378: 4370: 4365: 4356: 4355: 4350: 4341: 4340: 4335: 4322: 4312: 4307: 4290:testate amoebae 4259: 4208: 4173: 4152:Globotruncanida 4112: 4073: 4029: 4012: 3979: 3975:Symphyacanthida 3946: 3922: 3905: 3888: 3851: 3827:Claustrosporida 3813: 3777: 3740: 3648: 3625: 3584: 3540: 3527: 3509: 3475: 3462: 3439: 3406: 3367: 3358: 3357: 3273: 3251: 3242: 3236: 3206: 3194: 3184: 3182: 3174: 3160: 3136: 3134:Further reading 3131: 3087: 3086: 3079: 3037: 3036: 3032: 2994: 2993: 2989: 2937: 2936: 2932: 2898: 2897: 2890: 2860: 2859: 2855: 2812: 2811: 2807: 2784:10.2307/3515076 2761: 2760: 2753: 2708:(4567): 47–49. 2695: 2694: 2690: 2676: 2666: 2660:Jensen, Soren. 2659: 2658: 2654: 2611: 2610: 2606: 2575:(S1): 159–166. 2562: 2561: 2557: 2513: 2512: 2508: 2446: 2445: 2441: 2391: 2390: 2386: 2344: 2343: 2339: 2295: 2294: 2290: 2264: 2263: 2259: 2223: 2222: 2218: 2202: 2197: 2196: 2192: 2181:Galathea Report 2176: 2171: 2170: 2166: 2138: 2137: 2133: 2123: 2122: 2118: 2084: 2083: 2079: 2037: 2036: 2032: 1982: 1981: 1977: 1917: 1916: 1912: 1881: 1880: 1876: 1868: 1866: 1864: 1839: 1838: 1821: 1804: 1798: 1796: 1787: 1786: 1782: 1746: 1745: 1736: 1700: 1699: 1656: 1614: 1613: 1609: 1579: 1578: 1574: 1538: 1537: 1533: 1491: 1490: 1486: 1476: 1475: 1471: 1457: 1456: 1452: 1438: 1437: 1433: 1419: 1409: 1402: 1401: 1397: 1367: 1366: 1359: 1323: 1322: 1313: 1303: 1302: 1283: 1274: 1272: 1260: 1259: 1252: 1243: 1239: 1234: 1230: 1226: 1204: 1017: 1011: 1008: 992: 986: 822: 709: 680: 662:in the form of 632: 599: 538: 424: 403: 319: 210: 206: 146: 60: 24:Xenophyophorea 17: 12: 11: 5: 4614: 4612: 4604: 4603: 4598: 4593: 4583: 4582: 4576: 4575: 4573: 4572: 4559: 4546: 4533: 4520: 4507: 4494: 4481: 4468: 4453: 4437: 4435: 4429: 4428: 4426: 4425: 4412: 4402: 4389: 4376: 4363: 4358:Xenophyophorea 4348: 4332: 4330: 4328:Xenophyophorea 4324: 4323: 4318: 4309: 4308: 4306: 4305: 4298:sporozoan-like 4274: 4271: 4264: 4261: 4260: 4258: 4257: 4255:Rhizoplasmidae 4252: 4247: 4245:Gymnosphaerida 4242: 4234: 4232: 4230:Incertae sedis 4226: 4225: 4222: 4221: 4218: 4217: 4214: 4213: 4210: 4209: 4207: 4206: 4201: 4194: 4189: 4183: 4181: 4179:Incertae sedis 4175: 4174: 4172: 4171: 4166: 4164:Guembelitriida 4160: 4158:Heterohelicida 4154: 4148: 4143: 4138: 4133: 4128: 4122: 4120: 4114: 4113: 4111: 4110: 4105: 4100: 4095: 4090: 4083: 4081: 4075: 4074: 4072: 4071: 4064: 4058: 4056:Xenophyophorea 4053: 4046: 4044: 4034: 4022: 4021: 4018: 4017: 4014: 4013: 4011: 4010: 4005: 4000: 3995: 3989: 3987: 3981: 3980: 3978: 3977: 3972: 3967: 3965:Chaunocanthida 3962: 3960:Arthracanthida 3956: 3954: 3948: 3947: 3945: 3944: 3938: 3936: 3927: 3912: 3898: 3897: 3894: 3893: 3890: 3889: 3887: 3886: 3885: 3884: 3879: 3869: 3863: 3861: 3857: 3856: 3853: 3852: 3850: 3849: 3844: 3839: 3834: 3829: 3823: 3821: 3815: 3814: 3812: 3811: 3806: 3800: 3798: 3789: 3782: 3770: 3769: 3766: 3765: 3762: 3761: 3758: 3757: 3754: 3753: 3750: 3749: 3746: 3745: 3742: 3741: 3739: 3738: 3736:Ventricleftida 3733: 3728: 3727: 3726: 3721: 3716: 3708: 3707: 3706: 3701: 3696: 3691: 3689:Phaeosphaerida 3686: 3681: 3670: 3668: 3658: 3657: 3654: 3653: 3650: 3649: 3647: 3646: 3641: 3635: 3633: 3624: 3623: 3618: 3613: 3611:Spongomonadida 3608: 3602: 3600: 3590: 3589: 3586: 3585: 3583: 3582: 3577: 3572: 3567: 3565:Glissomonadida 3561: 3555: 3545: 3533: 3532: 3529: 3528: 3526: 3525: 3519: 3517: 3511: 3510: 3508: 3507: 3502: 3496: 3494: 3487: 3480: 3468: 3467: 3464: 3463: 3461: 3460: 3455: 3453:Chlorarachnida 3449: 3447: 3441: 3440: 3438: 3437: 3432: 3427: 3425:Desmothoracida 3422: 3416: 3414: 3408: 3407: 3405: 3404: 3399: 3393: 3391: 3384: 3381:Reticulofilosa 3374: 3360: 3359: 3356: 3355: 3354: 3353: 3352: 3351: 3348:Mesomycetozoea 3345: 3340: 3330: 3320: 3319: 3318: 3313: 3308: 3303: 3298: 3297: 3296: 3284:Diaphoretickes 3281: 3276: 3271: 3266: 3261: 3256: 3248: 3247: 3244: 3243: 3241:classification 3237: 3235: 3234: 3227: 3220: 3212: 3205: 3204: 3192: 3172: 3171: 3166: 3159: 3158:External links 3156: 3155: 3154: 3144: 3135: 3132: 3130: 3129: 3077: 3030: 2987: 2930: 2888: 2869:(3): 287–288. 2853: 2805: 2751: 2688: 2679:|journal= 2652: 2604: 2555: 2506: 2439: 2404:(1): 387–397. 2384: 2337: 2288: 2277:(3): 325–329. 2257: 2216: 2190: 2164: 2151:(6): 886–900. 2131: 2116: 2077: 2050:(7): 664–669. 2030: 1975: 1910: 1874: 1862: 1819: 1790:"Foraminifera" 1780: 1734: 1654: 1607: 1572: 1551:(3): 455–464. 1531: 1504:(6): 483–487. 1484: 1469: 1450: 1431: 1422:|journal= 1395: 1357: 1336:(4): 723–748. 1311: 1281: 1250: 1237: 1227: 1225: 1222: 1221: 1220: 1215: 1210: 1203: 1200: 1199: 1198: 1191: 1184: 1177: 1169: 1162: 1155: 1148: 1141: 1134: 1127: 1120: 1113: 1106: 1099: 1092: 1085: 1078: 1071: 1064: 1057: 1050: 1043: 1036: 1029: 1019: 1018: 1012:September 2020 998: 996: 985: 984:List of genera 982: 944:Gulf of Alaska 821: 818: 776:graphoglyptids 707: 679: 676: 631: 628: 598: 595: 537: 534: 500: 499: 485: 478: 463: 422: 402: 399: 318: 315: 293:abyssal plains 263:foraminiferans 261:is a clade of 204:Xenophyophorea 199: 198: 197: 196: 195: 194: 183: 175: 164: 163: 160:incertae sedis 154: 153: 144:Xenophyophorea 141: 134: 133: 128: 124: 123: 118: 114: 113: 108: 104: 103: 98: 91: 90: 88:Diaphoretickes 85: 78: 77: 72: 68: 67: 54: 53: 49: 48: 40: 39: 35: 34: 26: 25: 15: 13: 10: 9: 6: 4: 3: 2: 4613: 4602: 4601:Extremophiles 4599: 4597: 4594: 4592: 4589: 4588: 4586: 4569: 4564: 4560: 4556: 4551: 4547: 4543: 4538: 4534: 4530: 4525: 4521: 4517: 4512: 4508: 4504: 4499: 4495: 4491: 4486: 4482: 4478: 4473: 4469: 4464: 4458: 4454: 4449: 4443: 4439: 4438: 4436: 4434: 4430: 4422: 4417: 4413: 4409: 4403: 4399: 4394: 4390: 4386: 4381: 4377: 4373: 4372:Xénophyophore 4368: 4364: 4359: 4353: 4349: 4344: 4338: 4334: 4333: 4331: 4329: 4325: 4321: 4316: 4303: 4299: 4295: 4291: 4287: 4283: 4279: 4275: 4272: 4270: 4266: 4265: 4262: 4256: 4253: 4251: 4248: 4246: 4243: 4241: 4240: 4236: 4235: 4233: 4231: 4227: 4205: 4202: 4200: 4199: 4195: 4193: 4190: 4188: 4185: 4184: 4182: 4180: 4176: 4170: 4167: 4165: 4161: 4159: 4155: 4153: 4149: 4147: 4146:Globigerinida 4144: 4142: 4139: 4137: 4134: 4132: 4129: 4127: 4124: 4123: 4121: 4119: 4118:Globothalamea 4115: 4109: 4106: 4104: 4101: 4099: 4096: 4094: 4091: 4089: 4085: 4084: 4082: 4080: 4076: 4070: 4069: 4065: 4062: 4059: 4057: 4054: 4051: 4048: 4047: 4045: 4042: 4038: 4035: 4032: 4027: 4023: 4009: 4006: 4004: 4001: 3999: 3996: 3994: 3991: 3990: 3988: 3986: 3982: 3976: 3973: 3971: 3968: 3966: 3963: 3961: 3958: 3957: 3955: 3953: 3949: 3943: 3940: 3939: 3937: 3935: 3934:Sticholonchea 3931: 3928: 3925: 3920: 3916: 3913: 3909: 3903: 3899: 3883: 3880: 3878: 3875: 3874: 3873: 3870: 3868: 3865: 3864: 3862: 3858: 3848: 3845: 3843: 3840: 3838: 3835: 3833: 3830: 3828: 3825: 3824: 3822: 3820: 3816: 3810: 3807: 3805: 3802: 3801: 3799: 3797: 3793: 3790: 3786: 3783: 3780: 3775: 3771: 3737: 3734: 3732: 3731:Tectofilosida 3729: 3725: 3722: 3720: 3717: 3715: 3712: 3711: 3709: 3705: 3704:Phaeoconchida 3702: 3700: 3697: 3695: 3692: 3690: 3687: 3685: 3682: 3680: 3677: 3676: 3675: 3672: 3671: 3669: 3667: 3663: 3659: 3645: 3642: 3640: 3637: 3636: 3634: 3632: 3631:Silicofilosea 3628: 3622: 3619: 3617: 3614: 3612: 3609: 3607: 3604: 3603: 3601: 3599: 3595: 3591: 3581: 3580:Cercomonadida 3578: 3576: 3573: 3571: 3568: 3566: 3563: 3562: 3559: 3556: 3553: 3549: 3546: 3543: 3538: 3534: 3524: 3521: 3520: 3518: 3516: 3512: 3506: 3505:Metromonadida 3503: 3501: 3498: 3497: 3495: 3491: 3488: 3484: 3481: 3478: 3473: 3469: 3459: 3456: 3454: 3451: 3450: 3448: 3446: 3445:Chlorarachnea 3442: 3436: 3433: 3431: 3430:Leucodictyida 3428: 3426: 3423: 3421: 3418: 3417: 3415: 3413: 3409: 3403: 3400: 3398: 3395: 3394: 3392: 3388: 3385: 3382: 3378: 3375: 3371: 3365: 3361: 3349: 3346: 3344: 3341: 3339: 3336: 3335: 3334: 3331: 3329: 3326: 3325: 3324: 3321: 3317: 3314: 3312: 3311:Stramenopiles 3309: 3307: 3304: 3302: 3299: 3295: 3292: 3291: 3290: 3287: 3286: 3285: 3282: 3280: 3277: 3274:(major groups 3272: 3270: 3267: 3265: 3262: 3260: 3257: 3254: 3250: 3249: 3245: 3240: 3233: 3228: 3226: 3221: 3219: 3214: 3213: 3210: 3203: 3198: 3193: 3191: 3181: 3177: 3170: 3167: 3165: 3162: 3161: 3157: 3152: 3148: 3145: 3142: 3138: 3137: 3133: 3125: 3121: 3117: 3113: 3108: 3103: 3099: 3095: 3091: 3084: 3082: 3078: 3073: 3069: 3065: 3061: 3057: 3053: 3049: 3045: 3041: 3034: 3031: 3026: 3022: 3018: 3014: 3010: 3006: 3002: 2998: 2991: 2988: 2983: 2979: 2974: 2969: 2965: 2961: 2957: 2953: 2950:(4): e02933. 2949: 2945: 2941: 2934: 2931: 2926: 2922: 2918: 2914: 2910: 2906: 2902: 2895: 2893: 2889: 2884: 2880: 2876: 2872: 2868: 2864: 2857: 2854: 2849: 2845: 2841: 2837: 2833: 2829: 2825: 2821: 2817: 2809: 2806: 2801: 2797: 2793: 2789: 2785: 2781: 2777: 2773: 2769: 2765: 2758: 2756: 2752: 2747: 2743: 2739: 2735: 2731: 2727: 2723: 2719: 2715: 2711: 2707: 2703: 2699: 2692: 2689: 2684: 2671: 2663: 2656: 2653: 2648: 2644: 2640: 2636: 2632: 2628: 2624: 2620: 2616: 2608: 2605: 2600: 2596: 2592: 2588: 2583: 2578: 2574: 2570: 2566: 2559: 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1824: 1820: 1815: 1809: 1795: 1794:www.ucl.ac.uk 1791: 1784: 1781: 1776: 1772: 1767: 1762: 1758: 1754: 1753:Palaeontology 1750: 1743: 1741: 1739: 1735: 1730: 1726: 1721: 1716: 1712: 1708: 1704: 1697: 1695: 1693: 1691: 1689: 1687: 1685: 1683: 1681: 1679: 1677: 1675: 1673: 1671: 1669: 1667: 1665: 1663: 1661: 1659: 1655: 1650: 1646: 1642: 1638: 1634: 1630: 1626: 1622: 1618: 1611: 1608: 1603: 1599: 1595: 1591: 1587: 1583: 1576: 1573: 1568: 1564: 1559: 1554: 1550: 1546: 1542: 1535: 1532: 1527: 1523: 1519: 1515: 1511: 1507: 1503: 1499: 1495: 1488: 1485: 1480: 1473: 1470: 1465: 1461: 1454: 1451: 1446: 1442: 1435: 1432: 1427: 1414: 1406: 1399: 1396: 1391: 1387: 1383: 1379: 1375: 1371: 1364: 1362: 1358: 1353: 1349: 1344: 1339: 1335: 1331: 1327: 1320: 1318: 1316: 1312: 1307: 1300: 1298: 1296: 1294: 1292: 1290: 1288: 1286: 1282: 1271: 1267: 1263: 1257: 1255: 1251: 1248:on 2019-01-07 1247: 1241: 1238: 1232: 1229: 1223: 1219: 1216: 1214: 1211: 1209: 1206: 1205: 1201: 1197: 1196: 1192: 1190: 1189: 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4490:Monothalamea 4477:Monothalamea 4463:Monothalamea 4433:Monothalamea 4432: 4327: 4269:paraphyletic 4239:Lapot gusevi 4237: 4229: 4198:Heterogromia 4196: 4187:Involutinida 4178: 4126:Textulariida 4108:Ammodiscidae 4098:Spirillinida 4079:Tubothalamea 4068:Reticulomyxa 4066: 4061:Astrorhizida 4055: 4050:Allogromiida 4041:Monothalamea 4031:Foraminifera 3985:Polycystinea 3970:Holocanthida 3867:Vampyrellida 3832:Haplosporida 3819:Ascetosporea 3809:Reticulosida 3714:Cryomonadida 3699:Phaeogromida 3694:Phaeocalpida 3684:Phaeocystida 3666:Thecofilosea 3621:Variglissida 3616:Marimonadida 3552:Sarcomonadea 3542:Ventrifilosa 3493:Metromonadea 3477:Monadofilosa 3420:Cryptofilida 3412:Granofilosea 3397:Aquavolonida 3333:Opisthokonta 3153:, p. 2. 3097: 3093: 3047: 3043: 3033: 3000: 2996: 2990: 2947: 2943: 2933: 2908: 2904: 2866: 2862: 2856: 2823: 2819: 2808: 2770:(1): 32–41. 2767: 2763: 2705: 2701: 2691: 2670:cite journal 2655: 2622: 2618: 2607: 2572: 2568: 2558: 2526:(1): 43–54. 2523: 2519: 2509: 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Retrieved 1240: 1231: 1213:Extremophile 1193: 1186: 1179: 1171: 1164: 1157: 1150: 1143: 1136: 1131:Nazareammina 1129: 1122: 1115: 1110:Spiculammina 1108: 1101: 1094: 1087: 1080: 1075:Semipsammina 1073: 1066: 1061:Semipsammina 1059: 1052: 1045: 1040:Aschemonella 1038: 1033:Occultammina 1031: 1026:Syringammina 1024: 1009: 975: 971:Syringammina 969: 963: 961: 955: 951: 946:collected a 933: 925:life history 921:cell culture 918: 915: 908: 895: 880:harpacticoid 861: 835: 803: 796:Paleodictyon 795: 790:Occultammina 788: 780:Paleodictyon 778: 773: 766: 762: 755: 749: 743: 737: 732: 720: 714: 697: 691:Paleodictyon 689: 681: 649: 635: 633: 617: 610: 608: 600: 589:Occultammina 587: 578: 572: 569: 565: 558: 552: 550: 546:foraminifera 539: 529:Occultammina 527: 525: 514: 508: 501: 496:foraminifera 487: 480: 474: 470: 459: 455: 449: 445: 439: 436: 429: 427: 412: 384: 369: 363: 359:Syringammina 357: 351: 347:Foraminifera 335:Foraminifera 320: 304: 286: 203: 202: 192:Tendal, 1972 181:Tendal, 1972 158: 143: 137: 131:Monothalamea 121:Foraminifera 94: 81: 18: 4524:iNaturalist 4457:Wikispecies 4352:Wikispecies 4302:fungus-like 4278:flagellates 4141:Carterinida 4136:Robertinida 4003:Spumellaria 3998:Nassellaria 3877:Phagomyxida 3860:Proteomyxia 3837:Mikrocytida 3435:Limnofilida 3390:Skiomonadea 3190:Marine life 2484:1885/230014 2308:: 129–140. 1713:: 106–116. 1262:MSNBC Staff 1082:Homogammina 846:echinoderms 842:crustaceans 757:Benkovacina 751:Neonereites 745:Yelovichnus 729:bilaterians 721:Dickinsonia 716:Dickinsonia 644:pseudopodia 492:radiolarian 460:xanthosomes 456:stercomares 329:. In 1883, 173:Poche, 1913 117:Subphylum: 4585:Categories 4204:Komokiacea 4088:Fusulinida 3993:Collodaria 3952:Acantharea 3942:Taxopodida 3924:Radiolaria 3872:Phytomyxea 3847:Paramyxida 3842:Paradinida 3674:Phaeodarea 3644:Euglyphida 3598:Imbricatea 3570:Katabiidae 3458:Minorisida 3100:: 125715. 3003:(5): 881. 2357:: 103608. 2236:: 99–104. 2213:: 415–422. 1869:2020-09-30 1799:2018-06-17 1275:2011-10-24 1224:References 1218:Piezophile 1195:Moanammina 1181:Holopsamma 1173:Cerelpemma 1166:Maudammina 1145:Maudammina 1138:Cerelpemma 1089:Rhizammina 1001:incomplete 872:polychaete 868:sipunculan 863:Hebefustis 814:diagenetic 768:Pteridinum 700:vendozoans 504:protoplasm 475:granellare 451:stercomata 391:Stannomida 387:Psamminida 311:coenocytes 178:Stannomida 170:Psamminida 4448:Q16608712 4131:Rotaliida 4093:Miliolida 3796:Gromiidea 3788:Marimyxia 3710:Eothecia 3606:Krakenida 3523:Helkesida 3500:Metopiida 3402:Tremulida 3328:Amoebozoa 3306:Alveolata 3294:Cryptista 3269:Eukaryota 3116:0932-4739 3072:0967-0637 3017:0025-3154 2964:1939-9170 2911:: 31–44. 2883:1502-3931 2848:0967-0645 2792:0883-1351 2730:0036-8075 2647:0301-9268 2625:: 79–87. 2599:128952011 2591:1661-8726 2542:1342-8144 2493:0036-8075 2434:128619251 2426:0305-8719 2379:0967-0637 2332:0171-8630 2283:0399-1784 2187:: 79–101. 2072:127121951 2064:0016-7029 2017:0028-0836 1952:2045-2322 1775:1475-4983 1729:0006-3207 1649:0377-8398 1567:1096-3642 1352:0024-4082 1208:Coenocyte 1188:Abyssalia 1103:Cerelasma 1096:Shinkaiya 936:DSV Alvin 905:molpadiid 891:snailfish 876:nematodes 854:commensal 704:Ediacaran 624:manganese 560:Cerelasma 554:Psammetta 516:Shinkaiya 511:cytoplasm 488:Xenophyae 471:granellae 446:stercomes 395:homoplasy 376:monophyly 365:Shinkaiya 75:Eukaryota 4442:Wikidata 4337:Wikidata 4273:†extinct 4192:Lagenida 4008:Orodaria 3804:Gromiida 3779:Endomyxa 3724:Matazida 3719:Ebriacea 3515:Helkesea 3486:Eoglissa 3370:Cercozoa 3323:Amorphea 3301:Rhizaria 3289:Hacrobia 3279:Excavata 3264:Bacteria 3239:Rhizaria 3124:32585572 3025:86707195 2982:31742677 2746:28690086 2738:17776707 2550:85146476 2501:30237355 1970:30108286 1808:cite web 1627:: 1–10. 1526:39906626 1518:14733441 1407:: 1–814. 1270:NBC News 1202:See also 1159:Stannoma 1152:Psametta 1124:Tendalia 1117:Bizarria 1047:Psammina 956:Psammina 897:Starfish 883:copepods 850:molluscs 804:Certain 785:Cambrian 739:Intrites 626:crusts. 606:waters. 481:Linellae 431:Psammina 343:Protista 107:Phylum: 71:Domain: 4555:5362903 4516:2912356 4343:Q632686 4282:amoebae 3908:Retaria 3338:Animals 3259:Archaea 3202:Biology 3176:Portals 3052:Bibcode 2973:7341444 2944:Ecology 2913:Bibcode 2863:Lethaia 2828:Bibcode 2800:3515076 2772:Bibcode 2764:PALAIOS 2710:Bibcode 2702:Science 2627:Bibcode 2461:Bibcode 2453:Science 2406:Bibcode 2359:Bibcode 2310:Bibcode 2238:Bibcode 2099:Bibcode 2025:4254138 1997:Bibcode 1961:6092355 1932:Bibcode 1893:Bibcode 1629:Bibcode 1590:Bibcode 1466:: 1–55. 1447:: 1–92. 1441:Zoology 1378:Bibcode 942:in the 874:worms, 858:isopods 832:on top. 820:Ecology 711:sterols 702:of the 664:glucose 656:diatoms 636:in vivo 630:Feeding 620:basalts 604:hypoxic 521:mercury 469:called 401:Anatomy 339:sponges 301:species 299:and 75 149:Schulze 127:Class: 111:Retaria 4568:744106 4529:497927 4421:137898 4405:NZOR: 4398:212500 3372:  3368:  3316:Plants 3253:Domain 3122:  3114:  3070:  3023:  3015:  2980:  2970:  2962:  2881:  2846:  2798:  2790:  2744:  2736:  2728:  2645:  2597:  2589:  2548:  2540:  2499:  2491:  2432:  2424:  2377:  2330:  2281:  2070:  2062:  2023:  2015:  1989:Nature 1968:  1958:  1950:  1860:  1773:  1727:  1647:  1565:  1524:  1516:  1350:  903:, and 878:, and 848:, and 810:barium 748:, and 467:barite 415:nuclei 356:found 297:genera 274:oceans 151:, 1904 4563:WoRMS 4537:IRMNG 4503:8ZXG5 4380:IRMNG 3343:Fungi 3021:S2CID 2796:JSTOR 2742:S2CID 2595:S2CID 2546:S2CID 2430:S2CID 2203:(PDF) 2177:(PDF) 2068:S2CID 2021:S2CID 1522:S2CID 484:test. 419:tests 327:tests 323:Greek 138:Clade 95:Clade 82:Clade 4542:1605 4393:NCBI 4385:1033 3147:NOAA 3120:PMID 3112:ISSN 3068:ISSN 3013:ISSN 2978:PMID 2960:ISSN 2879:ISSN 2844:ISSN 2788:ISSN 2734:PMID 2726:ISSN 2683:help 2643:ISSN 2587:ISSN 2538:ISSN 2497:PMID 2489:ISSN 2422:ISSN 2375:ISSN 2328:ISSN 2279:ISSN 2060:ISSN 2013:ISSN 1966:PMID 1948:ISSN 1858:ISBN 1814:link 1771:ISSN 1725:ISSN 1645:ISSN 1563:ISSN 1514:PMID 1426:help 1348:ISSN 968:and 870:and 725:stem 658:and 502:The 362:and 282:test 4511:EoL 4498:CoL 4485:AFD 4472:ADW 4367:ADW 3102:doi 3060:doi 3005:doi 2968:PMC 2952:doi 2948:101 2921:doi 2871:doi 2836:doi 2780:doi 2718:doi 2706:218 2635:doi 2623:249 2577:doi 2573:104 2528:doi 2479:hdl 2469:doi 2457:361 2414:doi 2402:286 2367:doi 2355:176 2318:doi 2306:270 2246:doi 2153:doi 2107:doi 2052:doi 2005:doi 1993:320 1956:PMC 1940:doi 1901:doi 1850:doi 1761:doi 1715:doi 1711:207 1637:doi 1625:100 1598:doi 1553:doi 1549:156 1506:doi 1386:doi 1338:doi 1334:183 1007:. 866:), 448:or 101:SAR 4587:: 4565:: 4552:: 4539:: 4526:: 4513:: 4500:: 4487:: 4474:: 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Index



Scientific classification
Edit this classification
Eukaryota
Diaphoretickes
SAR
Retaria
Foraminifera
Monothalamea
Xenophyophorea
Schulze
incertae sedis
Psamminida
Stannomida
Ammoclathrinidae
/ˌzɛnəˌfəˈfrə/
foraminiferans
multinucleate
unicellular organisms
oceans
foraminiferan
test
Henry Bowman Brady
abyssal plains
genera
species
Syringammina fragilissima
coenocytes
Greek

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