1815:
44:
1805:
1800:
1810:
352:, and its teeth, if it had any, were likely vestigial and served no purpose. Since the whale had several double-headed vertebral joints, and modern baleen whales have a reduced number of these to increase the flexibility of the ribcage to aid in deep-sea diving,
229:
1894:
278:
in the skull used as passage for blood vesselsthough not as many as modern baleen whales–which indicate it had baleen in its mouth. The skull contains several
488:
1881:
1814:
1958:
1968:
1963:
274:, the skull is wide and flat. The skull, along the sides, has several narrow, straight grooves–eight in total–and there are several
481:
1953:
314:, and the complete series was preserved. The neural spine projecting up from the vertebra increases along the series.
43:
446:"A new mysticete from the upper Oligocene Ashiya Group, Kyushu, Japan and its significance to mysticete evolution"
474:
1804:
1948:
1834:
1799:
221:
1920:
162:
1599:
138:
445:
318:
one through seven and nine were preserved. The ribs had a double-headed joint with the vertebrae. The
1627:
947:
871:
824:
217:
1466:
1281:
1216:
1070:
957:
783:
1762:
1691:
1397:
1361:
922:
466:
361:
1193:
1170:
1051:
631:
1809:
1742:
1712:
1664:
1616:
1589:
1341:
1146:
529:
287:
38:
1925:
1722:
1223:
1202:
1156:
1096:
621:
600:
580:
1253:
1086:
1021:
642:
247:
translates to "with groove" in reference to the grooves in its mouth which indicate functional
1907:
1899:
315:
213:
1912:
1872:
610:
286:
and held no teeth, or the teeth naturally fell out over the course of the animal's life. The
1732:
1550:
1420:
385:
252:
653:
1702:
1647:
1577:
1441:
892:
756:
290:
was located midway along the snout, near the position of eyes. Unlike the closely related
1772:
1566:
1260:
851:
837:
334:
319:
297:
255:
201:
194:
118:
30:
325:
The left forearm and fragments of the right were discovered. As in modern whales, the
1942:
1427:
1331:
1315:
1041:
712:
705:
413:
405:
349:
311:
1654:
1606:
1535:
1495:
1434:
1406:
1351:
1304:
864:
726:
668:
563:
550:
498:
376:
372:
292:
279:
271:
259:
244:
197:
1857:
1866:
1752:
1505:
1413:
1372:
1237:
1031:
684:
390:
333:
is proportionally longer than that of modern baleen whales, a characteristic of
326:
322:
was concave on the side facing the ribs, a primitive feature for baleen whales.
301:
95:
1674:
1525:
1515:
1485:
1455:
1246:
1209:
1179:
1136:
994:
974:
931:
772:
765:
742:
735:
691:
677:
409:
400:
395:
367:
359:
Several marine vertebrates were found in the Ashiya Group: the baleen whales "
233:
1681:
1637:
1297:
1290:
1267:
1126:
1116:
1011:
1001:
984:
885:
878:
803:
590:
283:
55:
1851:
1448:
1383:
1106:
810:
796:
517:
75:
1886:
1475:
964:
535:
381:
330:
275:
105:
453:
Bulletin of the
Kitakyushu Museum of Natural History and Human History
698:
523:
511:
248:
225:
85:
65:
1828:
282:, but no teeth were discovered; it is possible these sockets were
237:
337:
whales, suggesting eomysticetids had less swimming capabilities.
1832:
548:
470:
296:, the head is relatively flat, and the snout is blunt. The
300:
in the ear is more similar to primitive baleen whales and
230:
Kitakyushu Museum of
Natural History and Human History
1768:
1758:
1748:
1738:
1728:
1718:
1708:
1698:
1687:
1670:
1660:
1643:
1633:
1623:
1612:
1595:
1585:
1562:
1531:
1521:
1511:
1501:
1491:
1481:
1394:
1379:
1368:
1357:
1347:
1337:
1327:
1278:
1234:
1190:
1167:
1152:
1142:
1132:
1122:
1112:
1102:
1092:
1082:
1047:
1037:
1027:
1017:
1007:
990:
980:
970:
953:
943:
848:
821:
753:
723:
665:
649:
638:
627:
617:
606:
596:
586:
576:
1841:
1576:
1549:
1465:
1393:
1314:
1277:
1233:
1189:
1166:
1069:
930:
921:
847:
820:
782:
752:
722:
664:
561:
416:, a terrestrial hippo-like animal, was also found.
216:, a partial skeleton, was discovered by curator
482:
439:
437:
435:
433:
431:
429:
8:
220:in October 1981 in the Ashiya Group of the
1829:
1555:
1320:
1075:
936:
927:
856:
829:
788:
569:
558:
545:
489:
475:
467:
398:; an unidentified crocodile; the seabirds
236:honors the old name of Japan, Yamato, and
20:
425:
7:
14:
1813:
1808:
1803:
1798:
356:likely stayed near the surface.
42:
329:probably could not rotate. The
1:
1959:Prehistoric cetacean genera
1985:
1969:Oligocene mammals of Asia
1964:Mammals described in 2012
1796:
1558:
1323:
1078:
939:
859:
832:
791:
572:
557:
544:
506:
251:. It was placed into the
193:is an extinct species of
190:Yamatocetus canaliculatus
175:Yamatocetus canaliculatus
168:
161:
39:Scientific classification
37:
23:
501:and their extinct allies
16:Extinct genus of mammals
371:, and an unidentified
258:, a primitive group of
375:; and an unidentified
1921:Paleobiology Database
154:Y. canaliculatus
444:Okazaki, Y. (2012).
391:Indotestudo takasago
228:, and stored in the
1954:Oligocene cetaceans
222:Jinnobaru Formation
1743:Plesiobalaenoptera
1590:Archaebalaenoptera
394:; an unidentified
316:Thoracic vertebrae
1936:
1935:
1908:Open Tree of Life
1835:Taxon identifiers
1826:
1825:
1822:
1821:
1794:
1793:
1790:
1789:
1786:
1785:
1782:
1781:
1600:Archaeschrichtius
1545:
1544:
1065:
1064:
1061:
1060:
917:
916:
913:
912:
909:
908:
224:on the island of
214:holotype specimen
186:
185:
142:
1976:
1929:
1928:
1916:
1915:
1903:
1902:
1890:
1889:
1877:
1876:
1875:
1862:
1861:
1860:
1830:
1817:
1812:
1807:
1802:
1770:
1760:
1750:
1740:
1733:Parabalaenoptera
1730:
1720:
1710:
1700:
1689:
1672:
1662:
1645:
1635:
1625:
1614:
1597:
1587:
1564:
1556:
1551:Balaenopteroidea
1533:
1523:
1513:
1503:
1493:
1483:
1396:
1381:
1370:
1359:
1349:
1339:
1329:
1321:
1280:
1236:
1192:
1169:
1154:
1144:
1134:
1124:
1114:
1104:
1094:
1084:
1076:
1049:
1039:
1029:
1019:
1009:
992:
982:
972:
955:
945:
937:
928:
857:
850:
830:
823:
789:
755:
725:
667:
651:
640:
629:
619:
608:
598:
588:
578:
570:
559:
546:
491:
484:
477:
468:
461:
460:
450:
441:
386:softshell turtle
177:
173:
137:
130:
117:
47:
46:
29:Temporal range:
21:
1984:
1983:
1979:
1978:
1977:
1975:
1974:
1973:
1939:
1938:
1937:
1932:
1924:
1919:
1911:
1906:
1898:
1893:
1885:
1880:
1871:
1870:
1865:
1856:
1855:
1850:
1837:
1827:
1818:
1778:
1703:Miobalaenoptera
1648:Eschrichtioides
1628:Cetoteriophanes
1578:Balaenopteridae
1572:
1541:
1461:
1442:Mithridatocetus
1389:
1310:
1273:
1229:
1185:
1162:
1057:
948:Antwerpibalaena
905:
872:Micromysticetus
843:
825:Eomysticetoidea
816:
778:
757:Mammalodontidae
748:
718:
660:
566:
553:
540:
502:
495:
465:
464:
448:
443:
442:
427:
422:
404:and an unknown
388:; the tortoise
343:
268:
243:, "whale." The
218:Toshiyuki Kamei
210:
182:
179:
171:
170:
157:
136:
128:
115:
41:
33:
27:
17:
12:
11:
5:
1982:
1980:
1972:
1971:
1966:
1961:
1956:
1951:
1941:
1940:
1934:
1933:
1931:
1930:
1917:
1904:
1891:
1878:
1863:
1847:
1845:
1839:
1838:
1833:
1824:
1823:
1820:
1819:
1797:
1795:
1792:
1791:
1788:
1787:
1784:
1783:
1780:
1779:
1777:
1776:
1773:Protororqualus
1766:
1756:
1746:
1736:
1726:
1716:
1706:
1696:
1685:
1678:
1668:
1658:
1651:
1641:
1631:
1621:
1610:
1603:
1593:
1582:
1580:
1574:
1573:
1571:
1570:
1567:Eobalaenoptera
1559:
1553:
1547:
1546:
1543:
1542:
1540:
1539:
1529:
1519:
1509:
1499:
1489:
1479:
1471:
1469:
1467:Herpetocetinae
1463:
1462:
1460:
1459:
1452:
1445:
1438:
1431:
1424:
1417:
1410:
1402:
1400:
1391:
1390:
1388:
1387:
1377:
1366:
1355:
1345:
1335:
1324:
1318:
1312:
1311:
1309:
1308:
1301:
1294:
1286:
1284:
1282:Tranatocetidae
1275:
1274:
1272:
1271:
1264:
1261:Parietobalaena
1257:
1250:
1242:
1240:
1231:
1230:
1228:
1227:
1220:
1217:Plesiocetopsis
1213:
1206:
1198:
1196:
1187:
1186:
1184:
1183:
1175:
1173:
1164:
1163:
1161:
1160:
1150:
1140:
1130:
1120:
1110:
1100:
1090:
1079:
1073:
1071:Thalassotherii
1067:
1066:
1063:
1062:
1059:
1058:
1056:
1055:
1045:
1035:
1025:
1015:
1005:
998:
988:
978:
968:
961:
958:Archaeobalaena
951:
940:
934:
925:
919:
918:
915:
914:
911:
910:
907:
906:
904:
903:
896:
889:
882:
875:
868:
860:
854:
852:Eomysticetidae
845:
844:
842:
841:
838:Cetotheriopsis
833:
827:
818:
817:
815:
814:
807:
800:
792:
786:
784:Chaeomysticeti
780:
779:
777:
776:
769:
761:
759:
750:
749:
747:
746:
739:
731:
729:
720:
719:
717:
716:
709:
702:
695:
688:
681:
673:
671:
662:
661:
659:
658:
647:
636:
625:
615:
604:
594:
584:
573:
567:
562:
555:
554:
549:
542:
541:
539:
538:
532:
526:
520:
514:
507:
504:
503:
496:
494:
493:
486:
479:
471:
463:
462:
424:
423:
421:
418:
408:; and unknown
342:
339:
320:shoulder blade
312:neck vertebrae
298:auditory bulla
267:
264:
256:Eomysticetidae
209:
206:
202:Late Oligocene
184:
183:
180:
166:
165:
159:
158:
150:
148:
144:
143:
126:
122:
121:
119:Eomysticetidae
113:
109:
108:
103:
99:
98:
93:
89:
88:
83:
79:
78:
73:
69:
68:
63:
59:
58:
53:
49:
48:
35:
34:
31:Late Oligocene
28:
15:
13:
10:
9:
6:
4:
3:
2:
1981:
1970:
1967:
1965:
1962:
1960:
1957:
1955:
1952:
1950:
1949:Baleen whales
1947:
1946:
1944:
1927:
1922:
1918:
1914:
1909:
1905:
1901:
1896:
1892:
1888:
1883:
1879:
1874:
1868:
1864:
1859:
1853:
1849:
1848:
1846:
1844:
1840:
1836:
1831:
1816:
1811:
1806:
1801:
1775:
1774:
1767:
1765:
1764:
1763:Praemegaptera
1757:
1755:
1754:
1747:
1745:
1744:
1737:
1735:
1734:
1727:
1725:
1724:
1717:
1715:
1714:
1707:
1705:
1704:
1697:
1694:
1693:
1692:Megapteropsis
1686:
1684:
1683:
1679:
1677:
1676:
1669:
1667:
1666:
1659:
1657:
1656:
1652:
1650:
1649:
1642:
1640:
1639:
1632:
1630:
1629:
1622:
1619:
1618:
1611:
1609:
1608:
1604:
1602:
1601:
1594:
1592:
1591:
1584:
1583:
1581:
1579:
1575:
1569:
1568:
1561:
1560:
1557:
1554:
1552:
1548:
1538:
1537:
1530:
1528:
1527:
1520:
1518:
1517:
1510:
1508:
1507:
1500:
1498:
1497:
1490:
1488:
1487:
1480:
1478:
1477:
1473:
1472:
1470:
1468:
1464:
1458:
1457:
1453:
1451:
1450:
1446:
1444:
1443:
1439:
1437:
1436:
1432:
1430:
1429:
1428:Eucetotherium
1425:
1423:
1422:
1418:
1416:
1415:
1411:
1409:
1408:
1404:
1403:
1401:
1399:
1398:Cetotheriinae
1392:
1386:
1385:
1378:
1375:
1374:
1367:
1364:
1363:
1362:Palaeobalaena
1356:
1354:
1353:
1346:
1344:
1343:
1336:
1334:
1333:
1332:Cephalotropis
1326:
1325:
1322:
1319:
1317:
1316:Cetotheriidae
1313:
1307:
1306:
1302:
1300:
1299:
1295:
1293:
1292:
1288:
1287:
1285:
1283:
1276:
1270:
1269:
1265:
1263:
1262:
1258:
1256:
1255:
1251:
1249:
1248:
1244:
1243:
1241:
1239:
1232:
1226:
1225:
1221:
1219:
1218:
1214:
1212:
1211:
1207:
1205:
1204:
1200:
1199:
1197:
1195:
1188:
1182:
1181:
1177:
1176:
1174:
1172:
1165:
1159:
1158:
1151:
1149:
1148:
1141:
1139:
1138:
1131:
1129:
1128:
1121:
1119:
1118:
1111:
1109:
1108:
1101:
1099:
1098:
1091:
1089:
1088:
1081:
1080:
1077:
1074:
1072:
1068:
1054:
1053:
1046:
1044:
1043:
1042:Peripolocetus
1036:
1034:
1033:
1026:
1024:
1023:
1016:
1014:
1013:
1006:
1004:
1003:
999:
997:
996:
989:
987:
986:
979:
977:
976:
969:
967:
966:
962:
960:
959:
952:
950:
949:
942:
941:
938:
935:
933:
929:
926:
924:
923:Balaenomorpha
920:
902:
901:
897:
895:
894:
890:
888:
887:
883:
881:
880:
876:
874:
873:
869:
867:
866:
862:
861:
858:
855:
853:
846:
840:
839:
835:
834:
831:
828:
826:
819:
813:
812:
808:
806:
805:
801:
799:
798:
794:
793:
790:
787:
785:
781:
775:
774:
770:
768:
767:
763:
762:
760:
758:
751:
745:
744:
740:
738:
737:
733:
732:
730:
728:
721:
715:
714:
713:Willungacetus
710:
708:
707:
706:Morawanocetus
703:
701:
700:
696:
694:
693:
689:
687:
686:
682:
680:
679:
675:
674:
672:
670:
663:
656:
655:
648:
645:
644:
637:
634:
633:
626:
624:
623:
616:
613:
612:
605:
603:
602:
595:
593:
592:
585:
583:
582:
575:
574:
571:
568:
565:
560:
556:
552:
547:
543:
537:
533:
531:
527:
525:
521:
519:
515:
513:
509:
508:
505:
500:
499:baleen whales
492:
487:
485:
480:
478:
473:
472:
469:
458:
454:
447:
440:
438:
436:
434:
432:
430:
426:
419:
417:
415:
412:. An unknown
411:
407:
406:pelagornithid
403:
402:
397:
393:
392:
387:
384:
383:
378:
374:
370:
369:
364:
363:
362:Metasqualodon
357:
355:
351:
350:filter feeder
348:was an early
347:
340:
338:
336:
332:
328:
323:
321:
317:
313:
309:
305:
303:
299:
295:
294:
289:
285:
281:
280:tooth sockets
277:
273:
272:baleen whales
265:
263:
261:
260:baleen whales
257:
254:
250:
246:
242:
239:
235:
231:
227:
223:
219:
215:
207:
205:
203:
199:
196:
192:
191:
181:Okazaki, 2012
178:
176:
167:
164:
163:Binomial name
160:
156:
155:
149:
146:
145:
140:
135:
134:
127:
124:
123:
120:
114:
111:
110:
107:
104:
101:
100:
97:
94:
91:
90:
87:
84:
81:
80:
77:
74:
71:
70:
67:
64:
61:
60:
57:
54:
51:
50:
45:
40:
36:
32:
26:
22:
19:
1842:
1771:
1761:
1751:
1741:
1731:
1721:
1711:
1701:
1690:
1680:
1673:
1663:
1655:Eschrichtius
1653:
1646:
1636:
1626:
1615:
1607:Balaenoptera
1605:
1598:
1588:
1565:
1536:Piscobalaena
1534:
1524:
1514:
1504:
1496:Herpetocetus
1494:
1484:
1474:
1454:
1447:
1440:
1435:Kurdalagonus
1433:
1426:
1419:
1412:
1407:Brandtocetus
1405:
1382:
1371:
1360:
1352:Otradnocetus
1350:
1340:
1330:
1305:Tranatocetus
1303:
1296:
1289:
1266:
1259:
1252:
1245:
1222:
1215:
1208:
1201:
1194:Diorocetidae
1178:
1171:Aglaocetidae
1155:
1145:
1135:
1125:
1115:
1105:
1095:
1085:
1052:Protobalaena
1050:
1040:
1030:
1020:
1010:
1000:
993:
983:
973:
963:
956:
946:
899:
898:
891:
884:
877:
870:
865:Eomysticetus
863:
836:
809:
802:
795:
771:
764:
741:
734:
727:Llanocetidae
711:
704:
697:
690:
683:
676:
669:Aetiocetidae
652:
641:
632:Siphonocetus
630:
620:
609:
599:
589:
579:
534:Infraorder:
530:Artiodactyla
456:
452:
399:
389:
380:
377:squalodontid
366:
360:
358:
353:
345:
344:
341:Paleoecology
335:eomysticetid
324:
307:
306:
302:archaeocetes
293:Eomysticetus
291:
270:As in other
269:
245:species name
240:
211:
198:baleen whale
195:eomysticetid
189:
188:
187:
174:
169:
153:
152:
132:
131:
102:Infraorder:
96:Artiodactyla
24:
18:
1873:Yamatocetus
1867:Wikispecies
1843:Yamatocetus
1753:Plesiocetus
1713:Nehalaennia
1665:Gricetoides
1617:Burtinopsis
1506:Metopocetus
1414:Cetotherium
1373:Titanocetus
1342:Journocetus
1238:Pelocetidae
1147:Tiphyocetus
1032:Morenocetus
900:Yamatocetus
685:Ashorocetus
414:amynodontid
379:dolphin; a
354:Yamatocetus
346:Yamatocetus
327:elbow joint
308:Yamatocetus
266:Description
133:Yamatocetus
25:Yamatocetus
1943:Categories
1723:Notiocetus
1675:Incakujira
1526:Nannocetus
1516:Miocaperea
1486:Herentalia
1456:Zygiocetus
1247:Cophocetus
1224:Thinocetus
1210:Diorocetus
1203:Amphicetus
1180:Aglaocetus
1157:Uranocetus
1137:Taikicetus
1097:Isanacetus
995:Balaenotus
975:Balaenella
932:Balaenidae
773:Mammalodon
766:Janjucetus
743:Mystacodon
736:Llanocetus
692:Chonecetus
678:Aetiocetus
622:Piscocetus
601:Imerocetus
581:Borealodon
497:Genera of
459:: 129–152.
420:References
401:Copepteryx
396:sea turtle
373:aetiocetid
368:Chonecetus
310:had seven
234:genus name
204:of Japan.
1858:Q25385774
1682:Megaptera
1638:Diunatans
1421:Ciuciulea
1298:Mixocetus
1291:Mesocetus
1268:Pelocetus
1254:Halicetus
1127:Pinocetus
1117:Mauicetus
1087:Hibacetus
1022:Mesoteras
1012:Idiocetus
1002:Eubalaena
985:Balaenula
886:Tokarahia
879:Tohoraata
804:Sitsqwayk
643:Tretulias
591:Coronodon
564:Mysticeti
551:Mysticeti
510:Kingdom:
284:vestigial
200:from the
147:Species:
62:Kingdom:
56:Eukaryota
1852:Wikidata
1449:Vampalus
1384:Tiucetus
1107:Isocetus
811:Whakakai
797:Horopeta
524:Mammalia
518:Chordata
516:Phylum:
512:Animalia
288:blowhole
276:foramina
208:Taxonomy
112:Family:
86:Mammalia
76:Chordata
72:Phylum:
66:Animalia
52:Domain:
1913:6145852
1900:1485113
1887:8550780
1476:Caperea
965:Balaena
893:Waharoa
611:Mioceta
536:Cetacea
528:Order:
522:Class:
382:Trionyx
331:humerus
172:†
151:†
139:Okazaki
125:Genus:
106:Cetacea
92:Order:
82:Class:
1926:247311
699:Fucaia
410:dugong
253:family
249:baleen
232:. The
226:Kyushu
141:, 2012
1895:IRMNG
654:Ulias
449:(PDF)
241:cetus
238:Latin
1882:GBIF
212:The
365:,"
1945::
1923::
1910::
1897::
1884::
1869::
1854::
457:10
455:.
451:.
428:^
304:.
262:.
1769:†
1759:†
1749:†
1739:†
1729:†
1719:†
1709:†
1699:†
1695:?
1688:†
1671:†
1661:†
1644:†
1634:†
1624:†
1620:?
1613:†
1596:†
1586:†
1563:†
1532:†
1522:†
1512:†
1502:†
1492:†
1482:†
1395:†
1380:†
1376:?
1369:†
1365:?
1358:†
1348:†
1338:†
1328:†
1279:†
1235:†
1191:†
1168:†
1153:†
1143:†
1133:†
1123:†
1113:†
1103:†
1093:†
1083:†
1048:†
1038:†
1028:†
1018:†
1008:†
991:†
981:†
971:†
954:†
944:†
849:†
822:†
754:†
724:†
666:†
657:?
650:†
646:?
639:†
635:?
628:†
618:†
614:?
607:†
597:†
587:†
577:†
490:e
483:t
476:v
129:†
116:†
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