37:
351:
Nine years later, Hernando and Ribera published a major redescription and organization of the genus. Previously grouped as seven species, it was reorganized into twenty species in five species groups, classified by similarities in genital shape and some external features. This 2014 revision also
730:. The head is slightly retracted into the pronotum, with long and narrow eleven-segment antennae covered in short setae. The underside of the abdomens (ventrum) has deep depressions where the long, slender legs connect with the body. Sexual dimorphism is relatively minimal; males of the
382:
subgroups. The current number of 35 species is likely to increase further, due to a relative lack of studies and material on Asian
Limnichidae. Many species are only known from a tiny number of specimens within a small area.
1343:
726:
beetles range from 2.5–5.5 mm in total length. They have ovloid bodies, ranging from brown to black in color, covered in both long and short layers of
1317:
1356:
309:. He described the species as the "largest and most remarkable of the Limnichini" and noted their close resemblance to the Central American genus
340:
in 1928; however, it is now accepted as a synonym. In 2005, Carles
Hernando and Ignacio Ribera reclassified one species described by Champion,
249:(male reproductive organ) without an articulated internal piece ("spiculum") within the median lobe; this distinguishes the genera from the
1144:
749:
They generally live in forests, inside piles of water-logged dead wood adjacent to small streams. They are most often collected using
304:
1438:
1361:
1171:
1110:
36:
612:
296:
1197:
1135:
288:
182:
125:
624:
474:
336:, in a paper describing three additional species. Pic disregarded Champion's synonymy and placed more species in
588:
484:
1433:
673:
572:
701:
667:
596:
584:
568:
449:
1400:
620:
468:
456:
357:
187:
793:
608:
604:
600:
580:
564:
560:
707:
616:
576:
556:
521:
1423:
1234:
655:
552:
478:
661:
592:
490:
329:
1387:
1428:
1304:
462:
444:
261:
738:
species have microscopic glandular pores along the central portion of their 4th sternite, while the
267:
228:
773:
241:
1272:
256:
31:
220:
251:
224:
1369:
1348:
1291:
1216:
1184:
1153:
1123:
1092:
1084:
1049:
1024:
234:
1374:
1176:
1115:
1076:
777:
1139:
765:
1417:
1296:
192:
149:
134:
1257:
1330:
1266:
321:
211:
207:
178:
174:
108:
98:
1119:
1080:
750:
245:. These genera share a number of physical features, such as a non-articulated
1220:
1188:
1157:
1127:
1088:
1053:
1028:
780:. Many species are known from a single location, although some (most notably
769:
68:
48:
1096:
1036:
Hernando, Carles; Ribera, Ignacio (2014). "Taxonomic
Revision of the Genus
1283:
1061:
Hernando, Carles; Ribera, Ignacio (2017). "Three New
Species of the Genus
1251:
1180:
734:
species have dissimilar anterior claws. Under a microscope, males of all
276:
246:
1395:
1322:
1067:
1335:
300:
88:
78:
58:
1228:
947:
945:
917:
915:
913:
911:
909:
1382:
1140:"Descriptions of Oriental Limnichini (Coleoptera, Fam. Byrrhidae)"
374:
have been identified since the 2014 redescription, all within the
170:
974:
972:
727:
1232:
896:
894:
854:
852:
1309:
932:
930:
1165:
Yoshitomi, Hiroyuki (2015). "Four new species of the genus
275:
group, but lack some distinguishing characteristics in the
317:
was the first described
Limnichinae genus native to Asia.
881:
879:
784:) have been found across a wide swath of territory. The
1104:
Matsumoto, Keita (2021). "Six new species of the genus
214:(Limnichidae). Within Limnichinae it belongs to the "
1241:
1065:Sharp, 1902 from Asia (Coleoptera: Limnichidae)".
328:, and placed in it four species of Limnichinae.
181:. The genus was first described by entomologist
1108:from the Himalayas (Coleoptera: Limnichidae)".
156:
1011:, a new genus of Limnichidae (Coleoptera) for
990:
978:
951:
921:
900:
870:
858:
831:
819:
218:group" of genera, alongside the much smaller
8:
1007:Hernando, Carles; Ribera, Ignacio (2005). "
800:group is limited to the mainland. Only the
1229:
20:
1209:Japanese Journal of Systematic Entomology
963:
936:
885:
385:
812:
352:classified one of Champion's species,
1202:(Coleoptera: Limnichidae) from China"
843:
7:
1401:CB7B4207-FD54-43A0-9617-7D15CAD62833
1388:E7618794-532C-FFC6-B2DB-2DC495DFFB48
370:As of 2021, fourteen new species of
746:groups also have these on the 5th.
332:considered this genus a synonym of
1040:Sharp (Coleoptera: Limnichidae)".
14:
792:species groups are only found in
544:denticle, or longitudinal sulcus
419:
324:described a new genus he termed
35:
1145:Entomologist's Monthly Magazine
764:species have been found across
306:Entomologist's Monthly Magazine
387:Species and species groups of
1:
804:group stretches across both.
1198:"A New Species of the Genus
1196:Yoshitomi, Hiroyuki (2018).
1172:European Journal of Taxonomy
1169:(Coleoptera, Limnichidae)".
1111:European Journal of Taxonomy
1042:Koleopterologische Rundschau
694:partial longitudinal sulcus
647:partial longitudinal sulcus
513:partial longitudinal sulcus
1013:Caccothryptus multiseriatus
271:are close relatives of the
212:minute marsh-loving beetles
1455:
991:Hernando & Ribera 2014
979:Hernando & Ribera 2014
952:Hernando & Ribera 2017
922:Hernando & Ribera 2014
901:Hernando & Ribera 2017
871:Hernando & Ribera 2014
859:Hernando & Ribera 2005
832:Hernando & Ribera 2014
820:Hernando & Ribera 2005
366:Species and species groups
344:, into a new genus dubbed
175:minute marsh-loving beetle
1120:10.5852/ejt.2021.739.1275
1081:10.11646/zootaxa.4243.2.5
691:On 4th and 5th sternites
642:On 4th and 5th sternites
437:Full longitudinal sulcus
155:
148:
140:
133:
32:Scientific classification
30:
23:
873:, pp. 281, 303–304.
410:Median lobe of aedeagus
794:Maritime Southeast Asia
645:Denticle on underside,
542:No lateral expansions,
303:) in an article in the
293:Caccothryptus compactus
1439:Taxa described in 1902
1017:Entomological Problems
287:In 1902, entomologist
330:Harry George Champion
210:, a subfamily of the
1181:10.5852/ejt.2015.147
511:Lateral expansions,
993:, pp. 303–304.
966:, pp. 138–140.
954:, pp. 366–370.
939:, pp. 168–184.
924:, pp. 281–304.
834:, pp. 284–285.
774:Indian subcontinent
391:
768:, and portions of
386:
320:In 1922 and 1923,
1411:
1410:
1370:Open Tree of Life
1235:Taxon identifiers
846:, pp. 61–63.
716:
715:
283:Taxonomic history
177:in the subfamily
162:
161:
129:
1446:
1404:
1403:
1391:
1390:
1378:
1377:
1365:
1364:
1352:
1351:
1339:
1338:
1326:
1325:
1313:
1312:
1300:
1299:
1287:
1286:
1277:
1276:
1275:
1262:
1261:
1260:
1230:
1224:
1206:
1192:
1161:
1131:
1114:(739): 168–184.
1100:
1057:
1032:
994:
988:
982:
976:
967:
961:
955:
949:
940:
934:
925:
919:
904:
898:
889:
883:
874:
868:
862:
856:
847:
841:
835:
829:
823:
817:
776:, including the
613:C. tardarsauceae
539:On 4th sternite
508:On 4th sternite
475:C. schillhammeri
434:On 4th sternite
421:
392:
356:as a synonym of
342:C. multiseriatus
124:
40:
39:
21:
16:Genus of beetles
1454:
1453:
1449:
1448:
1447:
1445:
1444:
1443:
1434:Beetles of Asia
1414:
1413:
1412:
1407:
1399:
1394:
1386:
1381:
1373:
1368:
1360:
1355:
1347:
1342:
1334:
1329:
1321:
1316:
1308:
1303:
1295:
1290:
1282:
1280:
1271:
1270:
1265:
1256:
1255:
1250:
1237:
1227:
1204:
1195:
1164:
1134:
1103:
1060:
1035:
1006:
1002:
997:
989:
985:
977:
970:
962:
958:
950:
943:
935:
928:
920:
907:
899:
892:
884:
877:
869:
865:
857:
850:
842:
838:
830:
826:
818:
814:
810:
786:jaechi, zetelli
778:Andaman Islands
759:
721:
589:C. occidentalis
485:C. sulawesianus
398:Anterior claws
368:
285:
262:Tricholimnichus
201:
144:
123:
34:
17:
12:
11:
5:
1452:
1450:
1442:
1441:
1436:
1431:
1426:
1416:
1415:
1409:
1408:
1406:
1405:
1392:
1379:
1366:
1353:
1340:
1327:
1314:
1301:
1288:
1278:
1263:
1247:
1245:
1239:
1238:
1233:
1226:
1225:
1215:(1): 138–140.
1193:
1162:
1132:
1101:
1075:(2): 366–370.
1058:
1033:
1023:(2): 131–135.
1009:Pseudothryptus
1003:
1001:
998:
996:
995:
983:
981:, p. 283.
968:
964:Yoshitomi 2018
956:
941:
937:Matsumoto 2021
926:
905:
903:, p. 370.
890:
886:Yoshitomi 2015
875:
863:
861:, p. 131.
848:
836:
824:
822:, p. 135.
811:
809:
806:
766:Southeast Asia
758:
755:
720:
717:
714:
713:
711:
698:
695:
692:
689:
686:
680:
679:
677:
674:C. wooldridgei
652:
649:
643:
640:
637:
631:
630:
628:
625:C. yunnanensis
573:C. fujianensis
549:
546:
540:
537:
534:
528:
527:
525:
518:
515:
509:
506:
503:
497:
496:
494:
441:
438:
435:
432:
429:
423:
422:
417:
414:
411:
408:
402:
396:
367:
364:
346:Pseudothryptus
338:Macrobyrrhinus
326:Macrobyrrhinus
284:
281:
268:Cyclolimnichus
229:Pseudothryptus
206:is a genus of
200:
197:
185:in 1902, with
160:
159:
153:
152:
146:
145:
138:
137:
131:
130:
116:
112:
111:
106:
102:
101:
96:
92:
91:
86:
82:
81:
76:
72:
71:
66:
62:
61:
56:
52:
51:
46:
42:
41:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
1451:
1440:
1437:
1435:
1432:
1430:
1427:
1425:
1424:Beetle genera
1422:
1421:
1419:
1402:
1397:
1393:
1389:
1384:
1380:
1376:
1371:
1367:
1363:
1358:
1354:
1350:
1345:
1341:
1337:
1332:
1328:
1324:
1319:
1315:
1311:
1306:
1302:
1298:
1293:
1289:
1285:
1279:
1274:
1273:Caccothryptus
1268:
1264:
1259:
1253:
1249:
1248:
1246:
1244:
1243:Caccothryptus
1240:
1236:
1231:
1222:
1218:
1214:
1210:
1203:
1201:
1200:Caccothryptus
1194:
1190:
1186:
1182:
1178:
1175:(147): 1–17.
1174:
1173:
1168:
1167:Caccothryptus
1163:
1159:
1155:
1151:
1147:
1146:
1141:
1137:
1133:
1129:
1125:
1121:
1117:
1113:
1112:
1107:
1106:Caccothryptus
1102:
1098:
1094:
1090:
1086:
1082:
1078:
1074:
1070:
1069:
1064:
1063:Caccothryptus
1059:
1055:
1051:
1047:
1043:
1039:
1038:Caccothryptus
1034:
1030:
1026:
1022:
1018:
1014:
1010:
1005:
1004:
999:
992:
987:
984:
980:
975:
973:
969:
965:
960:
957:
953:
948:
946:
942:
938:
933:
931:
927:
923:
918:
916:
914:
912:
910:
906:
902:
897:
895:
891:
887:
882:
880:
876:
872:
867:
864:
860:
855:
853:
849:
845:
840:
837:
833:
828:
825:
821:
816:
813:
807:
805:
803:
799:
795:
791:
787:
783:
779:
775:
771:
767:
763:
762:Caccothryptus
756:
754:
752:
747:
745:
741:
737:
736:Caccothryptus
733:
729:
725:
724:Caccothryptus
718:
712:
710:
709:
704:
703:
702:C. luzonensis
699:
696:
693:
690:
687:
685:
682:
681:
678:
676:
675:
670:
669:
668:C. ticaoensis
664:
663:
658:
657:
653:
650:
648:
644:
641:
638:
636:
633:
632:
629:
627:
626:
622:
618:
614:
610:
606:
602:
598:
594:
590:
586:
585:C. nepalensis
582:
578:
574:
570:
569:C. chayuensis
566:
562:
558:
554:
550:
547:
545:
541:
538:
535:
533:
530:
529:
526:
524:
523:
519:
516:
514:
510:
507:
504:
502:
499:
498:
495:
493:
492:
487:
486:
481:
480:
476:
471:
470:
465:
464:
459:
458:
453:
452:
451:C. chayuensis
447:
446:
442:
439:
436:
433:
430:
428:
425:
424:
418:
415:
412:
409:
407:
406:pores (male)
403:
401:
397:
394:
393:
390:
389:Caccothryptus
384:
381:
377:
373:
372:Caccothryptus
365:
363:
361:
360:
355:
354:C. laosensis,
349:
347:
343:
339:
335:
334:Caccothryptus
331:
327:
323:
318:
316:
315:Caccothryptus
312:
308:
307:
302:
298:
294:
290:
282:
280:
278:
274:
270:
269:
264:
263:
258:
254:
253:
248:
244:
243:
238:
236:
231:
230:
226:
222:
217:
213:
209:
205:
204:Caccothryptus
198:
196:
194:
190:
189:
184:
180:
176:
172:
168:
167:
166:Caccothryptus
158:
154:
151:
147:
143:
139:
136:
132:
127:
122:
121:
120:Caccothryptus
117:
114:
113:
110:
107:
104:
103:
100:
97:
94:
93:
90:
87:
84:
83:
80:
77:
74:
73:
70:
67:
64:
63:
60:
57:
54:
53:
50:
47:
44:
43:
38:
33:
29:
26:
25:Caccothryptus
22:
19:
1242:
1212:
1208:
1199:
1170:
1166:
1149:
1143:
1136:Sharp, David
1109:
1105:
1072:
1066:
1062:
1045:
1041:
1037:
1020:
1016:
1012:
1008:
1000:Bibliography
986:
959:
888:, p. 1.
866:
839:
827:
815:
801:
797:
796:, while the
789:
785:
782:C. maculosus
781:
761:
760:
748:
743:
739:
735:
731:
723:
722:
706:
700:
683:
672:
666:
660:
654:
646:
634:
621:C. tibetanus
597:C. punctatus
565:C. championi
561:C. brendelli
551:
543:
531:
520:
512:
500:
489:
483:
473:
469:C. maculosus
467:
461:
457:C. compactus
454:
450:
443:
426:
405:
399:
388:
379:
375:
371:
369:
359:C. maculosus
358:
353:
350:
345:
341:
337:
333:
325:
319:
314:
310:
305:
292:
286:
272:
266:
260:
257:Afrotropical
250:
242:Simplocarina
240:
233:
219:
215:
203:
202:
193:type species
188:C. compactus
186:
165:
164:
163:
142:C. compactus
141:
135:Type species
119:
118:
24:
18:
1429:Limnichidae
1331:iNaturalist
1267:Wikispecies
751:light traps
719:Description
609:C. taiwanus
605:C. sinensis
601:C. ripicola
581:C. malickyi
553:C. arakawae
431:Dissimilar
395:Group name
322:Maurice Pic
289:David Sharp
208:Limnichinae
183:David Sharp
179:Limnichinae
109:Limnichinae
105:Subfamily:
99:Limnichidae
1418:Categories
844:Sharp 1902
808:References
708:C. zetteli
617:C. testudo
577:C. jendeki
557:C. auratus
548:Separated
522:C. rouyeri
440:Separated
413:Parameres
311:Euthryptus
295:(found in
291:described
221:Euthryptus
157:35 species
89:Coleoptera
69:Arthropoda
1258:Q14932675
1221:1341-1160
1189:2118-9773
1158:0013-8908
1152:: 61–64.
1128:2118-9773
1089:1175-5326
1054:0075-6547
1029:1335-5899
802:compactus
770:East Asia
732:compactus
697:(varies)
656:C. jaechi
651:(varies)
479:C. schuhi
463:C. larryi
445:C. abboti
427:compactus
404:Sternite
376:compactus
297:Martapura
273:Mandersia
252:Byrrhinus
225:Mandersia
216:Mandersia
150:Diversity
55:Kingdom:
49:Eukaryota
1281:BioLib:
1252:Wikidata
1138:(1902).
1097:28610152
772:and the
688:Similar
662:C. nanus
639:Similar
593:C. orion
536:Similar
505:Similar
416:Species
277:pronotum
247:aedeagus
235:Resachus
199:Taxonomy
95:Family:
65:Phylum:
59:Animalia
45:Domain:
1396:ZooBank
1375:4643620
1349:1151142
1323:1033259
1068:Zootaxa
798:testudo
790:rouyeri
744:zetteli
684:zetteli
532:testudo
501:rouyeri
491:C. thai
400:(male)
380:testudo
259:genera
191:as the
115:Genus:
85:Order:
79:Insecta
75:Class:
1362:722575
1336:733310
1284:539614
1219:
1187:
1156:
1126:
1095:
1087:
1052:
1027:
788:, and
740:jaechi
635:jaechi
517:Fused
301:Borneo
255:. The
128:, 1902
1383:Plazi
1344:IRMNG
1310:39587
1297:62K47
1205:(PDF)
757:Range
728:setae
171:genus
169:is a
126:Sharp
1357:ITIS
1318:GBIF
1217:ISSN
1185:ISSN
1154:ISSN
1124:ISSN
1093:PMID
1085:ISSN
1073:4243
1050:ISSN
1025:ISSN
742:and
378:and
265:and
239:and
1305:EoL
1292:CoL
1177:doi
1116:doi
1077:doi
1015:".
420:Ref
173:of
1420::
1398::
1385::
1372::
1359::
1346::
1333::
1320::
1307::
1294::
1269::
1254::
1213:24
1211:.
1207:.
1183:.
1150:38
1148:.
1142:.
1122:.
1091:.
1083:.
1071:.
1048:.
1046:84
1044:.
1021:35
1019:.
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