447:
progressively taller as they approach the dorsal vertebrae. The hypapophysis of the cervicals are knob-like. The first two dorsal vertebrae are similar in shape to the cervicals with strongly compressed neural spines. The neural spine of the 3rd dorsal is blade like with a knob on the dorsal tip and the hypapophysis is prominent with a stronger keel than the other vertebrae. The 4th dorsal vertebra is the last with a distinct hypapophysis and the first with the parapophysis is entirely located on the neural spine. The posterior dorsal vertebrae are marked by massive centra and short transverse processes. The neural spines of the posterior dorsal vertebrae progressively grow shorter and anteroposteriorly elongated. The single preserved saccral is stout. The first caudal vertebra lacks a neural spine while the anterior vertebra possesses a tall spines and short transverse processes. The other caudals are from more posterior regions of the tail and notably elongated with relatively high neural spines. 28 ribs from various areas of the skeleton are known. Of the pectoral girdle and forelimbs both scapula is known, the left is complete while of the right only the distal end has been found. Neither coracoid is known. The left humerus is known as well as the shaft of the right. The shaft of the humerus is almost straight and slender, but the curvature of the proximal head and distal condylar region give the bone an overall sigmoid shape. Further material includes a complete left ulna and the distal three fourths of the right ulna, elements of both radius, complete left radiale and ulnare as well as distal phallanges and metacarpals. From the pelvic girdle the complete right ilium has been preserved as well as both ischia, however both are damaged. The articular area for the pubis is well developed. The pubis had a tear-drop shaped articular surface for the pubis and a more blade-like distal end. The femora, tibiae and fibulae resemble those of modern crocodilians. The pes are known from the right calcaneum and 2 distal metatarsals.
438:, in which the antorbital fenestra is almost entirely located within the lacrimal. Along the contact with the prefrontals the lacrimals swell to form the posterolateral portion of the prominent rostral ridge. The posteromedial portion is formed by the elongated prefrontals. The frontal is broad between the orbits and narrows significantly between the prefrontals before making contact with the nasals. It only extends slightly between the supratemporal fenestrae. The medial wall and floor of the supratemporal fenestrae is made up by the parietal. In specimen FMNH PR 2448 the lateral edges of the fenestrae are strongly everted, creating a narrow medial groove. This feature however seems to vary between specimen, as FMNH PR 2389 is flat in comparison. A thin lamina of the parietal overhangs the posterior and medial edge of the supratemporals. The dentary has 14 alveoli on each half, the biggest of which being the 4th and the 9th. The first dentary tooth is elevated above the others.
432:
evenly sized amongst themselves before the 10th and 11th (and potentially 12th) would have marked a further decrease in size. There is a notable diastema located between the 4th and 5th maxillary alveoli, most likely to make space for a large dentary tooth. Medially to the antorbital fenestra the maxilla forms a prominent ridge that runs laterally to the maxillary nasal contact. The nasals are fused, however it is uncertain if they reached up to the external nares. The nasals did not contact the lacrimal, which is anteriorly bifurcated and forms most of the posterior edge of the antorbital fenestra, a condition also seen in
1089:
155:
668:
1747:
2319:
1766:
2312:
129:
2326:
2334:
1759:
1753:
372:, consisted of a complete left and partial right mandible, vertebrae of the cervical, dorsal, saccral and caudal regions, several ribs as well as material of the pectoral, pelvic girdle and limb bones. Osteoderms and isolated teeth have also been found in association with the skeletal remains. Although this specimen was originally believed to represent
403:
was a medium to large sized crocodylomorph, characterized by a strongly arched jugal, depressions beneath the orbit, a broad platyrostral snout, a massive choanal septum, a broad and rounded anterior edge of the dentary, ziphodont teeth and short mandibular symphysis, reaching posteriorly only to the
679:
Although
Notosuchians are known for their terrestrial lifestyle, mahajangasuchids present a deviation from this ecology by having evolved multiple traits indicative of a more semi-aquatic lifestyle. The high-sided skull table with ventrally positioned jaw articulation differs from the dorsoventrally
391:
The generic name means "crocodile from
Mahajanga Basin" in reference to the area the fossils had been found in. The species epithet insignis is Latin and means "remarkable" or "extraordinary", chosen to not only reflect the preservation of the holotype specimen but also due to the strange morphology
431:
are very low in lateral view, appearing almost flat, and are heavily sculpted. They are roughly as long as they are wide with 11 to 12 alveoli on each side. The first 4 maxillary teeth were closely spaced with the 3rd being the largest. The 5th to 9th were smaller than the previous 4, but roughly
446:
Of the postcranial skeletal several elements are known. The holotype preserves 5 cervical vertebrae, thought to be the 1st to 4th and 5th postaxial cervicals which are amphicoelous to slightly amphiplatyan in shape. The intercentrum is wider than long and the neural spine of the cervical grows
454:
had deep irregular pits over their surface except for the anterior margin, where they are overlapped by the prior osteoderm. They have strong keels and are subquadrangular to ovate in shape, suggesting that the dorsal armor did not consist of more than two rows of osteoderms.
680:
compressed crania with posteriorly located jaw articulation of modern
Eusuchians, while the broad and shallow rostrum appears to have been obtained convergently to similar morphology seen in the Neosuchians. The palate of
366:. This included a variety of crocodylomorphs with the largest specimen being a well preserved disarticulated skeleton discovered in 1995 roughly 1 km south-east of the village of Berivotra. This skeleton, specimen
702:) represent a lineage of Notosuchian that independently adapted from a terrestrial to a semi-aquatic lifestyle, citing features such as the dorsally located nares and orbits as well as the platyrostrine skull shape.
426:
the skull is noticeably platyrostral, being flat and wide. The premaxilla possesses 1 to 4, potentially 5, tooth positions on each side, with the 3rd premaxillary tooth being the largest. The maxilla of
355:
With the inception of the
Mahajanga Basin Project (MBP) in 1993, led by Dr. David Krause, came a significant increase of discoveries and research into the fauna of the
2455:
692:
determine supports the idea that this morphology evolved to resist torsional forces during feeding. A semi-aquatic lifestyle was also recovered by
Wilberg
902:"A second peirosaurid crocodyliform from the Mid-Cretaceous Kem Kem Group of Morocco and the diversity of Gondwanan notosuchians outside South America"
502:
by Sereno and
Larrson (2009), who considered the family to be basal neosuchians not closely allied with peirosaurids. However, later analysis returned
2534:
2514:
2442:
388:
by
Buckley and Brochu in 1999. While the holotype specimen lacked cranial remains, further excavations did uncover skull material of the animal.
2539:
2519:
1122:
2361:
518:(2021) which recovered mahajangasuchids as a sister clade to peirosaurids, the resulting grouping in turn taking a sister position to
141:
719:
David W. Krause; Patrick M. O’Connor; Kristina Curry Rogers; Scott D. Sampson; Gregory A. Buckley; Raymond R. Rogers (2006).
154:
2529:
720:
2318:
1746:
1044:"Crocodyliform biogeography during the Cretaceous: evidence of Gondwanan vicariance from biogeographical analysis"
1115:
1765:
510:, to its sister position to peirosaurids and furthermore recovered this clade to be a branch of early diverging
2524:
2311:
838:
739:
2325:
820:(Crocodyliformes: Mesoeucrocodylia) cranial anatomy and new data on the origin of the eusuchian-style palate"
2382:
467:
has oftentimes been grouped closest to peirosaurids, such as in both
Buckley and Brochu (1999) and Turner
382:), further research concluded that this specimen instead represented a new genus of crocodylomorph, named
2333:
2509:
2481:
1207:
721:"Late Cretaceous Terrestrial Vertebrates from Madagascar: Implications for Latin American Biogeography"
2429:
2420:
1758:
1108:
973:
913:
869:
1094:
359:
340:
2061:
842:
751:
743:
149:
2486:
1752:
2468:
2460:
1621:
1351:
1073:
999:
939:
580:
493:
485:
261:
2473:
2205:
1167:
1063:
1055:
989:
981:
929:
921:
877:
834:
735:
41:
2264:
2239:
2077:
2050:
1995:
1970:
1963:
1917:
1870:
1844:
1656:
1628:
539:
336:
248:
235:
977:
917:
873:
667:
2280:
2230:
2192:
2138:
2116:
2002:
1940:
1887:
1614:
1538:
1517:
1415:
1332:
1068:
1043:
994:
961:
934:
901:
698:
519:
473:
434:
815:
2503:
2273:
2246:
2212:
2173:
2158:
2131:
2124:
2070:
2036:
2029:
2020:
1982:
1947:
1859:
1829:
1822:
1691:
1607:
1600:
1584:
1510:
1482:
783:
561:
489:
378:
356:
79:
846:
128:
2253:
2149:
2103:
1894:
1837:
1663:
1635:
1593:
1552:
1545:
1529:
1502:
1495:
1457:
1394:
1192:
755:
511:
328:
287:
222:
696:(2019), proposing that mahajangasuchids (in their phylogeny clading together with
2414:
2166:
2084:
2043:
1955:
1927:
1814:
1684:
1670:
1573:
1471:
1422:
1408:
1401:
1387:
1360:
1318:
1287:
1269:
1256:
1219:
1131:
594:
514:. The phylogeny below is a simplified cladogram based on the results of Nicholl
498:
478:
410:
324:
54:
17:
2405:
985:
2219:
2181:
1901:
1880:
1852:
1796:
1783:
1714:
1677:
1649:
1642:
1566:
1450:
1443:
1436:
1340:
1176:
1161:
1084:
962:"Evolutionary structure and timing of major habitat shifts in Crocodylomorpha"
685:
363:
347:. It was a fairly large predator, measuring up to 4 metres (13 ft) long.
344:
99:
64:
2009:
1559:
1429:
1378:
1325:
209:
166:
104:
48:
1077:
1059:
1003:
943:
882:
861:
2399:
1155:
405:
186:
94:
89:
74:
69:
59:
925:
747:
2447:
1464:
1304:
317:
196:
109:
84:
1149:
176:
2376:
900:
Nicholl, C.S.C.; Hunt, E.S.E.; Ouarhache, D.; Mannion, P.D. (2021).
784:"An enigmatic new crocodile from the Upper Cretaceous of Madagascar"
404:
posterior margin of the 2nd alveolous, superficially resembling the
666:
488:, while a later study by Turner and Calvo (2005) placed it within
320:
1048:
Proceedings of the Royal
Society of London B: Biological Sciences
2434:
2380:
2350:
1781:
1254:
1142:
1104:
1100:
1024:(Crocodyliformes) from the Upper Cretaceous of Madagascar".
839:
10.1671/0272-4634(2008)28[382:micmca]2.0.co;2
740:
10.3417/0026-6493(2006)93[178:LCTVFM]2.0.CO;2
294:
2389:
2263:
2229:
2191:
2148:
2102:
2060:
2019:
1981:
1926:
1915:
1869:
1794:
1713:
1583:
1528:
1481:
1377:
1350:
1302:
1267:
1218:
1191:
960:Wilberg, E.W.; Turner, A.H.; Brochu, C.A. (2019).
492:. It was placed in the newly constructed family
1116:
8:
484:. (2001) placed the genus within the family
862:"Cretaceous crocodyliforms from the Sahara"
2377:
2347:
2197:
2108:
1987:
1932:
1923:
1806:
1802:
1791:
1778:
1710:
1487:
1310:
1279:
1275:
1264:
1251:
1197:
1188:
1139:
1123:
1109:
1101:
127:
31:
1067:
993:
933:
881:
860:Sereno, P. C.; Larsson, H. C. E. (2009).
728:Annals of the Missouri Botanical Garden
711:
955:
953:
895:
893:
809:
807:
805:
803:
801:
777:
775:
773:
771:
769:
767:
765:
339:; its fossils have been found in the
7:
814:Turner, A.H.; Buckley, G.A. (2008).
327:which had blunt, conical teeth. The
586:
576:
554:
532:
525:
1026:Journal of Vertebrate Paleontology
1020:Buckley, G.A. (2001). "A skull of
827:Journal of Vertebrate Paleontology
782:Buckley, G.A.; Brochu, C. (1999).
25:
2535:Prehistoric pseudosuchian genera
2332:
2324:
2317:
2310:
1764:
1757:
1751:
1745:
1087:
153:
2515:Late Cretaceous crocodylomorphs
471:(2008) (under the inclusion of
142:Field Museum of Natural History
1:
2540:Fossil taxa described in 1998
2520:Crocodylomorphs of Madagascar
788:Cretaceous Fossil Vertebrates
684:is similar to that of modern
2362:Terrestrial crocodylomorphs
2556:
986:10.1038/s41598-018-36795-1
304:Buckley & Brochu, 1998
281:Buckley & Brochu, 1998
2357:
2346:
2308:
2200:
2111:
1990:
1935:
1809:
1805:
1790:
1777:
1743:
1709:
1490:
1313:
1282:
1278:
1263:
1250:
1200:
1187:
1138:
608:
591:
584:
574:
559:
552:
537:
530:
293:
286:
150:Scientific classification
148:
135:
126:
34:
27:Extinct genus of reptiles
1042:Turner, Alan H. (2004).
1022:Mahajangasuchus insignis
818:Mahajangasuchus insignis
385:Mahajangasuchus insignis
299:Mahajangasuchus insignis
138:Mahajangasuchus insignis
1060:10.1098/rspb.2004.2840
883:10.3897/zookeys.28.325
676:
474:Trematochampsa taqueti
2482:Paleobiology Database
670:
496:along with the genus
374:Trematochampsa oblita
1031:(3), supplement: A36
442:Postcranial skeleton
379:Miadanasuchus oblita
351:Discovery and naming
1095:Paleontology portal
1054:(1552): 2003–2009.
978:2019NatSR...9..514W
926:10.1098/rsos.211254
918:2021RSOS....811254N
874:2009ZooK...28....1S
360:Maevarano Formation
341:Maevarano Formation
335:, lived during the
2530:Maastrichtian life
2352:Related categories
2062:Pissarrachampsinae
966:Scientific Reports
677:
450:The osteoderms of
2497:
2496:
2469:Open Tree of Life
2383:Taxon identifiers
2374:
2373:
2370:
2369:
2342:
2341:
2306:
2305:
2302:
2301:
2298:
2297:
2294:
2293:
2290:
2289:
2098:
2097:
2094:
2093:
1911:
1910:
1773:
1772:
1741:
1740:
1737:
1736:
1733:
1732:
1705:
1704:
1701:
1700:
1622:Colhuehuapisuchus
1352:Mahajangasuchidae
1298:
1297:
1246:
1245:
1242:
1241:
1238:
1237:
906:The Royal Society
659:
658:
650:
649:
641:
640:
632:
631:
623:
622:
581:Mahajangasuchidae
494:Mahajangasuchidae
486:Trematochampsidae
392:of the mandible.
309:
308:
282:
262:Mahajangasuchidae
16:(Redirected from
2547:
2490:
2489:
2477:
2476:
2464:
2463:
2451:
2450:
2438:
2437:
2425:
2424:
2423:
2410:
2409:
2408:
2378:
2348:
2337:
2336:
2329:
2328:
2322:
2321:
2314:
2206:Adamantinasuchus
2198:
2109:
1988:
1933:
1924:
1807:
1803:
1792:
1779:
1768:
1762:
1761:
1755:
1749:
1711:
1488:
1311:
1280:
1276:
1265:
1252:
1198:
1189:
1182:
1181:
1168:Mesoeucrocodylia
1140:
1125:
1118:
1111:
1102:
1097:
1092:
1091:
1090:
1081:
1071:
1008:
1007:
997:
957:
948:
947:
937:
897:
888:
887:
885:
857:
851:
850:
824:
811:
796:
795:
779:
760:
759:
725:
716:
587:
577:
555:
533:
526:
296:
280:
273:
260:
158:
157:
131:
121:
51:
42:Upper Cretaceous
40:Temporal range:
32:
21:
18:Mahajangasuchini
2555:
2554:
2550:
2549:
2548:
2546:
2545:
2544:
2525:Maevarano fauna
2500:
2499:
2498:
2493:
2485:
2480:
2472:
2467:
2459:
2454:
2446:
2441:
2433:
2428:
2421:Mahajangasuchus
2419:
2418:
2413:
2404:
2403:
2398:
2391:Mahajangasuchus
2385:
2375:
2366:
2353:
2338:
2331:
2323:
2316:
2286:
2265:Caipirasuchinae
2259:
2240:Armadillosuchus
2225:
2187:
2144:
2090:
2078:Pissarrachampsa
2056:
2051:Stratiotosuchus
2015:
1996:Cynodontosuchus
1977:
1971:Razanandrongobe
1964:Pehuenchesuchus
1920:
1918:Xenodontosuchia
1907:
1871:Candidodontidae
1865:
1845:Razanandrongobe
1799:
1786:
1769:
1756:
1729:
1697:
1657:Montealtosuchus
1629:Gasparinisuchus
1579:
1524:
1477:
1373:
1368:Mahajangasuchus
1346:
1307:
1294:
1272:
1259:
1234:
1214:
1183:
1145:
1144:
1134:
1129:
1093:
1088:
1086:
1041:
1038:
1017:
1012:
1011:
959:
958:
951:
899:
898:
891:
859:
858:
854:
822:
813:
812:
799:
781:
780:
763:
723:
718:
717:
713:
708:
688:, which Turner
682:Mahajangasuchus
673:Mahajangasuchus
671:Restoration of
665:
660:
651:
642:
633:
624:
612:Mahajangasuchus
540:Uruguaysuchidae
504:Mahajangasuchus
465:Mahajangasuchus
461:
452:Mahajangasuchus
444:
429:Mahajangasuchus
424:Mahajangasuchus
420:
401:Mahajangasuchus
398:
353:
337:Late Cretaceous
313:Mahajangasuchus
305:
302:
279:
276:Mahajangasuchus
271:
258:
249:Crocodyliformes
236:Crocodylomorpha
152:
122:
120:
119:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
46:
45:
38:
36:Mahajangasuchus
28:
23:
22:
15:
12:
11:
5:
2553:
2551:
2543:
2542:
2537:
2532:
2527:
2522:
2517:
2512:
2502:
2501:
2495:
2494:
2492:
2491:
2478:
2465:
2452:
2439:
2426:
2411:
2395:
2393:
2387:
2386:
2381:
2372:
2371:
2368:
2367:
2365:
2364:
2358:
2355:
2354:
2351:
2344:
2343:
2340:
2339:
2309:
2307:
2304:
2303:
2300:
2299:
2296:
2295:
2292:
2291:
2288:
2287:
2285:
2284:
2281:Morrinhosuchus
2277:
2269:
2267:
2261:
2260:
2258:
2257:
2250:
2243:
2235:
2233:
2231:Sphagesaurinae
2227:
2226:
2224:
2223:
2216:
2209:
2201:
2195:
2193:Sphagesauridae
2189:
2188:
2186:
2185:
2178:
2170:
2163:
2154:
2152:
2146:
2145:
2143:
2142:
2139:Eptalofosuchus
2135:
2128:
2121:
2117:Brasileosaurus
2112:
2106:
2100:
2099:
2096:
2095:
2092:
2091:
2089:
2088:
2081:
2074:
2066:
2064:
2058:
2057:
2055:
2054:
2047:
2040:
2033:
2025:
2023:
2017:
2016:
2014:
2013:
2006:
2003:Gondwanasuchus
1999:
1991:
1985:
1979:
1978:
1976:
1975:
1967:
1960:
1952:
1944:
1941:Chimaerasuchus
1936:
1930:
1921:
1916:
1913:
1912:
1909:
1908:
1906:
1905:
1898:
1891:
1888:Lavocatchampsa
1884:
1876:
1874:
1867:
1866:
1864:
1863:
1856:
1849:
1841:
1834:
1826:
1819:
1810:
1800:
1795:
1788:
1787:
1782:
1775:
1774:
1771:
1770:
1744:
1742:
1739:
1738:
1735:
1734:
1731:
1730:
1728:
1727:
1726:
1725:
1719:
1717:
1707:
1706:
1703:
1702:
1699:
1698:
1696:
1695:
1688:
1681:
1674:
1667:
1660:
1653:
1646:
1639:
1632:
1625:
1618:
1615:Bayomesasuchus
1611:
1604:
1597:
1589:
1587:
1581:
1580:
1578:
1577:
1570:
1563:
1556:
1549:
1542:
1539:Barreirosuchus
1534:
1532:
1526:
1525:
1523:
1522:
1518:Stolokrosuchus
1514:
1507:
1499:
1491:
1485:
1479:
1478:
1476:
1475:
1468:
1461:
1454:
1447:
1440:
1433:
1426:
1419:
1416:Dentaneosuchus
1412:
1405:
1398:
1391:
1383:
1381:
1375:
1374:
1372:
1371:
1364:
1356:
1354:
1348:
1347:
1345:
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889:
852:
833:(2): 382–408.
797:
761:
734:(2): 178–208.
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699:Stolokrosuchus
675:sunning itself
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2203:
2202:
2199:
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2179:
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2171:
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2159:Coronelsuchus
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2147:
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2132:Coronelsuchus
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2125:Coringasuchus
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2110:
2107:
2105:
2101:
2087:
2086:
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2071:Campinasuchus
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2048:
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2039:
2038:
2037:Aplestosuchus
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2030:Aphaurosuchus
2027:
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2022:
2021:Baurusuchinae
2018:
2012:
2011:
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463:Historically
458:
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81:
76:
71:
66:
61:
56:
50:
43:
37:
33:
30:
19:
2510:Notosuchians
2390:
2315:
2279:
2272:
2254:Sphagesaurus
2252:
2245:
2238:
2218:
2211:
2204:
2180:
2172:
2165:
2157:
2150:Notosuchidae
2137:
2130:
2123:
2115:
2104:Sphagesauria
2083:
2076:
2069:
2049:
2042:
2035:
2028:
2008:
2001:
1994:
1969:
1962:
1954:
1946:
1939:
1900:
1895:Malawisuchus
1893:
1886:
1879:
1858:
1851:
1843:
1838:Libycosuchus
1836:
1828:
1821:
1813:
1750:
1690:
1683:
1676:
1669:
1664:Patagosuchus
1662:
1655:
1648:
1641:
1636:Hamadasuchus
1634:
1627:
1620:
1613:
1606:
1599:
1594:Antaeusuchus
1592:
1572:
1565:
1558:
1553:Epoidesuchus
1551:
1546:Caririsuchus
1544:
1537:
1530:Pepesuchinae
1516:
1509:
1503:Fortignathus
1501:
1496:Amargasuchus
1494:
1470:
1463:
1458:Sahitisuchus
1456:
1449:
1442:
1435:
1428:
1421:
1414:
1407:
1400:
1395:Barinasuchus
1393:
1386:
1367:
1366:
1359:
1339:
1331:
1324:
1317:
1286:
1208:Pseudosuchia
1193:Pseudosuchia
1172:
1051:
1047:
1028:
1025:
1021:
1015:Bibliography
969:
965:
909:
905:
865:
855:
830:
826:
817:
791:
787:
731:
727:
714:
697:
693:
689:
681:
678:
672:
663:Paleobiology
611:
610:
609:
593:
592:
560:
538:
515:
512:notosuchians
507:
506:, alongside
503:
497:
481:
472:
468:
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451:
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445:
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428:
423:
421:
409:
400:
399:
390:
384:
383:
377:
373:
368:
367:
362:in northern
354:
343:in northern
332:
329:type species
312:
311:
310:
298:
297:
288:Type species
275:
274:
242:
229:
223:Pseudosuchia
216:
203:
137:
35:
29:
2415:Wikispecies
2167:Llanosuchus
2085:Wargosuchus
2044:Baurusuchus
1956:Eremosuchus
1928:Baurusuchia
1815:Anatosuchus
1685:Rukwasuchus
1671:Peirosaurus
1574:Roxochampsa
1472:Zulmasuchus
1423:Iberosuchus
1409:Bretesuchus
1402:Bergisuchus
1388:Ayllusuchus
1361:Kaprosuchus
1319:Baharijodon
1288:Microsuchus
1270:Ziphosuchia
1257:Ziphosuchia
1220:Ziphosuchia
1132:Ziphosuchia
595:Kaprosuchus
508:Kaprosuchus
499:Kaprosuchus
411:Mourasuchus
396:Description
333:M. insignis
210:Archosauria
47:70–66
2504:Categories
2220:Yacarerani
2182:Notosuchus
1902:Pakasuchus
1881:Candidodon
1853:Simosuchus
1797:Notosuchia
1784:Notosuchia
1724:see below↓
1715:Notosuchia
1678:Pepesuchus
1650:Lomasuchus
1643:Kinesuchus
1567:Pepesuchus
1451:Ogresuchus
1444:Lorosuchus
1437:Langstonia
1341:Wanosuchus
1229:see below↓
1177:Metasuchia
1162:Sauropsida
972:(1): 514.
794:: 149–175.
706:References
686:Eusuchians
364:Madagascar
345:Madagascar
2010:Pabwehshi
1560:Itasuchus
1430:Ilchunaia
1379:Sebecidae
1326:Doratodon
1148:Kingdom:
459:Phylogeny
173:Kingdom:
167:Eukaryota
136:Skull of
2406:Q1066937
2400:Wikidata
1156:Chordata
1154:Phylum:
1150:Animalia
1078:15451689
1004:30679529
944:34659786
847:85634099
748:40035721
406:Cenozoic
255:Family:
197:Reptilia
187:Chordata
183:Phylum:
177:Animalia
163:Domain:
2474:4132248
2461:1161490
2448:4822106
2435:4473007
1465:Sebecus
1305:Sebecia
1166:Order:
1160:Class:
1069:1691824
995:6346023
974:Bibcode
935:8511751
914:Bibcode
870:Bibcode
866:ZooKeys
756:9166607
408:caiman
369:UA 8654
318:extinct
268:Genus:
193:Class:
140:in the
116:↓
2487:114396
1076:
1066:
1002:
992:
942:
932:
845:
754:
746:
694:et al.
690:et al.
516:et al.
482:et al.
479:Sereno
469:et al.
316:is an
2456:IRMNG
1173:Clade
843:S2CID
823:(PDF)
752:S2CID
744:JSTOR
724:(PDF)
418:Skull
376:(now
321:genus
243:Clade
230:Clade
217:Clade
204:Clade
2443:GBIF
1206:see
1074:PMID
1000:PMID
940:PMID
55:PreꞒ
2430:EoL
1064:PMC
1056:doi
1052:271
990:PMC
982:doi
930:PMC
922:doi
878:doi
835:doi
736:doi
477:).
422:In
323:of
2506::
2484::
2471::
2458::
2445::
2432::
2417::
2402::
1175::
1072:.
1062:.
1050:.
1046:.
1029:21
998:.
988:.
980:.
968:.
964:.
952:^
938:.
928:.
920:.
908:.
904:.
892:^
876:.
864:.
841:.
831:28
829:.
825:.
800:^
792:60
790:.
786:.
764:^
750:.
742:.
732:93
730:.
726:.
522:.
414:.
331:,
245::
232::
219::
206::
144:.
105:Pg
49:Ma
44:,
2177:?
2162:?
2120:?
1974:?
1959:?
1951:?
1873:?
1848:?
1833:?
1818:?
1521:?
1506:?
1336:?
1291:?
1124:e
1117:t
1110:v
1080:.
1058::
1006:.
984::
976::
970:9
946:.
924::
916::
910:8
886:.
880::
872::
849:.
837::
816:"
758:.
738::
295:†
272:†
259:†
110:N
100:K
95:J
90:T
85:P
80:C
75:D
70:S
65:O
60:Ꞓ
20:)
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