1185:
668:
1049:
1101:
839:
1543:
781:, though they are not known from the eastern Pacific coast of the Americas. Their eyestalks possess styles, and they are frequently confused with similar species which share their distribution range (especially juveniles). They can be distinguished by a stridulating ridge of 10 to 11 interspaced tubercles on the upper third, 8 thick striae on the central third, and 20 to 30 closely spaced tubercles on the lower third; as well as by their slender gonopod with a palp, and the pointed tip of their smaller claw.
51:
1141:
82:
1228:
786:
1659:
1353:
729:
1438:
949:
1627:
1180:. In adult males, the outer edges of the eye orbits are tooth-shaped and project forward. The palm of the claws are densely covered with tubercles on the upper surface, with distinctively serrated upper and lower margins. The stridulating ridge has 35 to 56 tubercles with striae. The tip of the smaller claw is characteristically blunt. Their eyestalks possess styles.
1001:
888:
463:
the eyestalks are tipped with a tuft of bristles (setae). The eyestalks are held vertically when the crab is active. Most species have pale-colored bodies that blend in well with the sand, though they are capable of gradually changing body coloration to match their environment and the time of day.
834:. They can be distinguished by the forward-facing triangular outer edges of their eye orbits, pointed smaller claws, a stridulating ridge of 19 to 24 tubercles, and a fringe of bristles on the upper margin of the propodi of the first and second walking legs.
1223:). Its carapace is covered densely with rough tubercles on the upper surface and wider than it is long. The outer edges of the eye orbits are broadly triangular and project forward. Their stridulating ridge has 20 to 30 tubercles.
475:(sound-producing) ridges which they use for communication. These ridges are also important morphological characters useful for identifying species. The chelipeds are shorter than the walking legs. The last pair of walking legs (
1136:. They can be distinguished by a stridulating ridge of about 10 interspaced tubercles. The propodi of their first to fourth pair of walking legs do not possess bristles on the upper and lower surfaces.
1497:
but can be distinguished by having 10 to 15 irregularly spaced elongated tubercles with striae on their stridulating ridges and a thumb-like palp on their gonopods. Their eyestalks possess styles.
1348:, but it remains unclear without further molecular genetic tests. Their stridulating ridges are composed of 30 to 42 thick striae in males and 17 to 29 in females. The smaller claw is pointed.
479:) is also usually shorter and thinner than the other pairs of walking legs. A cavity, with edges fringed by long setae are also found in between the bases of the second and third walking legs.
303:
regions throughout the world. They have a box-like body, thick and elongated eyestalks, and one claw is larger than the other in both males and females. They inhabit deep burrows in the
2549:
1586:
has also been recorded in the Red Sea, but only rarely). Also known from
Madagascar through Indian Ocean to the Western Pacific. Their eyestalks possess styles.
2588:
1774:
2155:
496:
ghost crabs construct simple to complex deep burrows in soft sandy and/or muddy substrates. They can be found in sandy beaches, rubble flats, and in
2523:
1263:
which is also found in the same regions, but can be distinguished by the presence of styles on their eyestalks. They also share the same range with
2562:
883:
in the western
Pacific. As their common name implies, they can be distinguished by the absence of stridulating ridges on the palm of their claws.
2262:
2132:
2383:
2346:
2047:
2014:
396:
by examining their gonopods. In the former, the first gonopod has a complex hoof-shaped tip, while in the latter they are simple and curved.
724:
of their first and second pair of walking legs and by the 23 to 28 tubercles on their stridulating ridge. Their eyestalks possess styles.
1898:
1184:
2660:
2064:
1825:
2072:. FAO Species Identification Sheets for Fishery Puporses. Food and Agriculture Organization of the United Nations. p. 183.
922:
799:
342:
190:
2567:
2575:
2083:
415:
occupying most of the lower portion. The eyestalks are also tipped with horn-like projections (styles) in seven species (
2209:
2113:
1048:
838:
667:
2484:
2471:
2211:
Quaternary coastal evolution adjacent to southern
Ningaloo Reef, Western Australia: Implications for land use planning
1703:
1100:
1538:. They can easily be distinguished by the presence of a characteristic red band along the inner edges of their legs.
2159:
1271:
have 35 to 71 striae on their stridulating ridges and their smaller claws have broadly rounded to flattened tips.
2593:
2182:
1362:
827:
663:
but can easily be distinguished from the latter by the absence of the tuft of bristles on the tip of their eyes.
81:
2241:
1232:
1195:
1040:
567:
2678:
471:) of both sexes are unequal in size, with one much larger than the other. The palm of the claws also possess
2706:
1542:
2424:
2409:
790:
752:
423:
350:
207:
60:
1392:
to
Madagascar. They possess 7 to 13 tubercles on their stridulating ridges. The smaller claw is pointed.
1044:
but are easily distinguishable by their bright coloration and the presence of styles on their eyestalks.
2624:
2489:
2006:
1447:
65:
2642:
2274:
944:. They are easily distinguished by the characteristic tuft of bristles on the end of their eyestalks.
2701:
2665:
2603:
2091:
1813:
1053:
1017:
429:
1296:
1072:
733:
688:
417:
50:
2319:
1942:
1664:
1337:
981:
892:
862:
795:
594:
505:
358:
312:
236:
76:
1695:
2629:
2375:
1890:
1140:
2611:
2554:
2510:
2379:
2371:
2342:
2043:
2031:
2010:
1894:
1631:
1594:
1474:
1461:
1422:
1357:
1315:
1160:
1062:
1058:
985:
933:
932:
Large-sized species found in the eastern
Atlantic, from Mauritania to Namibia; as well as the
823:
778:
539:
447:
435:
2652:
1998:
2616:
2363:
2311:
2039:
1977:
1934:
1882:
1863:
1611:
1442:
1400:
1376:
1292:
1133:
1005:
968:
880:
672:
634:
382:
1851:
1566:
1345:
1279:
1247:
1227:
897:
843:
809:
785:
713:
453:
441:
372:
304:
2401:
2229:
546:
coast of the
Americas. The rest of the species are found in the western Pacific and the
1547:
1527:
1505:
1220:
1145:
1111:
1030:
Medium to large-sized species found in the eastern
Pacific coast of the Americas, from
953:
911:
535:
459:
1259:
Small-sized species found in South China and southern Japan. It is closely related to
2695:
2462:
2364:
1915:
1883:
1855:
1418:
1414:
918:
721:
543:
1582:. Very common species in the Red Sea, and one of only two species known from there (
2515:
2296:
Thomas J. Trott (1988). "Note on the foraging activities of the painted ghost crab
1531:
1490:
1486:
1352:
1089:
1085:
728:
547:
472:
457:), though these may be shorter or even absent altogether in juvenile specimens. In
346:
339:
199:
123:
1658:
411:
are usually not clearly defined. They have thick and elongated eyestalks with the
2447:
2580:
2536:
2456:
2302:
1519:
677:
573:
531:
377:
300:
143:
2476:
1437:
1000:
948:
1654:
1535:
1328:
Small to medium-sized species. They are one of the most widespread species in
1237:
1216:
1208:
765:
Medium to large-sized species. They are one of the most widespread species in
648:
488:
385:
368:
355:
292:
173:
163:
1867:
1622:
but can easily be distinguished because they lack styles on their eyestalks.
2413:
2315:
1673:
1626:
1366:
1341:
1150:
1129:
1031:
989:
901:
852:
774:
709:
508:
501:
497:
476:
315:
308:
113:
93:
879:, being found in the western Indian Ocean (including the Red Sea) to the
2497:
2441:
1885:
Marine
Decapod Crustacea of Southern Australia: A Guide to Identification
1636:
1556:
1552:
1344:. It has been proposed that this species may actually be composed of two
1173:
1125:
848:
656:
527:
512:
468:
408:
319:
296:
133:
2502:
2323:
1694:
Sammy De Grave; N. Dean
Pentcheff; Shane T. Ahyong; et al. (2009).
647:
Medium-sized species found in the eastern
Atlantic coast of Africa from
367:
was previously the only genus classified under the ghost crab subfamily
2673:
2528:
1946:
1579:
1523:
1451:
1389:
1177:
770:
738:
652:
323:
219:
31:
17:
1982:
1961:
887:
565:
currently contains 21 valid species. The ghost crab formerly known as
2541:
1696:"A classification of living and fossil genera of decapod crustaceans"
1426:
1333:
1300:
1212:
1124:
Medium to large-sized species found in the Nicobar Islands, southern
958:
941:
875:
Medium-sized species. They are one of the most widespread species in
551:
412:
103:
69:
2418:
2066:
Field Guide to the Commercial Marine Resources of the Gulf of Guinea
1938:
371:
until 2013, when Katsushi Sakai and Michael Türkay reclassified the
2362:
G. M. Branch; M. L. Branch; C. L. Griffiths; L. E. Buckley (2007).
2300:
H. Milne Edwards & Lucas in Costa Rica (Decapoda, Brachyura)".
2637:
2133:"Creature Feature - Haunted Beaches: The fleet-footed Ghost Crabs"
2030:
George Karleskint, Richard Keith Turner & James Small (2009).
1640:
1625:
1615:
1607:
1541:
1478:
1436:
1413:
Medium to large-sized species found in the western Atlantic, from
1351:
1226:
1183:
1139:
1099:
1047:
1035:
999:
947:
937:
886:
837:
831:
784:
742:
727:
705:
666:
288:
1482:
1429:. They are easily distinguished as they are the only species of
701:
153:
2422:
769:, being found in the entirety of the Indian Ocean (except the
1518:
Medium-sized species found in the east coast of Africa, from
534:
regions throughout the world. Three species are found in the
819:
golden ghost crab, western ghost crab, or yellow ghost crab
2088:
Marine Invertebrates of the National Park of American Samoa
2003:
Ocracoke Wild: A Naturalist's Year on an Outer Banks Island
1960:
Martin Stevens; Cheo Pei Rong & Peter A. Todd (2013).
407:
ghost crabs have deep box-like bodies. The regions on the
2366:
Two Oceans: a Guide to the Marine Life of Southern Africa
2341:. Johannesburg, South Africa: Jacana Media. p. 72.
1176:, India (including the Nicobar Islands), Sri Lanka, and
1962:"Colour change and camouflage in the horned ghost crab
1916:"Morphological color change in the Hawaiian ghost crab
1473:
Large-sized species found in the southern coast of the
2189:. Department of the Environment, Australian Government
2431:
1307:but they do not share the same distribution range.
996:, but they are not found from the same localities.
716:. They can be distinguished from other species of
2158:. Government of Western Australia. Archived from
1096:, but they are not found in the same localities.
1084:Small to medium-sized species found only in the
1881:Gary C. B. Poore & Shane T. Ahyong (2004).
1860:Bulletin of the United States National Museum
27:Genus of tropical and subtropical crustaceans
8:
1773:Katsushi Sakai & Michael Türkay (2013).
1578:Medium to large-sized species found in the
1038:. They share their distribution range with
2419:
2217:(Thesis). Curtin University of Technology.
762:horned ghost crab or horn-eyed ghost crab
49:
38:
1981:
1970:Biological Journal of the Linnean Society
1846:
1844:
1842:
1790:Memoirs of the Queensland Museum – Nature
1768:
1766:
1764:
1762:
1760:
1758:
1756:
1754:
1752:
1750:
1748:
1746:
1744:
1742:
1740:
1738:
1736:
1734:
615:
322:of small animals. The genus contains 21
1807:
1805:
1803:
1732:
1730:
1728:
1726:
1724:
1722:
1720:
1718:
1716:
1714:
1686:
30:For the harpy in Greek mythology, see
2154:Naturaliste Marine Discovery Centre.
1779:with the description of a new genus,
7:
2679:C18FB670-34E5-4586-AAAD-B47D5E5C0398
2643:49810D0F-B08E-6B02-C4F7-48504C2EBA6D
2604:45bb5ba2-f23c-4ea8-b36e-a11410bcddea
338:was first established by the German
1999:"Phantoms in the Surf: Ghost Crabs"
392:can be distinguished from those in
2131:Western Australian Museum (2011).
25:
2339:The Coastal Guide of South Africa
2135:. Government of Western Australia
1783:(Crustacea: Decapoda: Brachyura)"
1291:Small-sized species found in the
1267:(which also possess styles), but
1023:H. Mile-Edwards & Lucas, 1843
720:in the Indian Ocean by the naked
571:was transferred to its own genus
2238:Great Barrier Reef Invertebrates
2181:SpeciesBank (February 7, 2007).
1826:World Register of Marine Species
1657:
80:
1856:"The Grapsoid crabs of America"
2273:. UBC - Canada. Archived from
2036:Introduction to Marine Biology
1211:and the southeastern coast of
1207:Medium-sized species found in
1172:Medium-sized species found in
980:Medium-sized species found in
800:British Indian Ocean Territory
295:found in the sandy shores of
1:
2208:Cherie Louise Leeden (2003).
1606:Small-sized species found in
1009:dorsal aspect (legs excluded)
822:Large-sized species found in
700:Large-sized species found in
1889:. CSIRO Publishing. p.
984:and Western Australia, from
228:
2120:. Wild Singapore. May 2009.
2063:Wolfgang Schneider (1990).
1704:Raffles Bulletin of Zoology
1493:. They are very similar to
1303:. They are very similar to
1092:. They are very similar to
388:. Ghost crabs of the genus
61:Ocypode ceratophthalma
2723:
1914:Jonathan P. Green (1964).
486:
29:
1433:in the western Atlantic.
1363:Point Lookout, Queensland
872:smooth-handed ghost crab
526:ghost crabs are found in
242:
235:
225:
218:
205:
198:
77:Scientific classification
75:
57:
48:
41:
2242:University of Queensland
2032:"Intertidal communities"
1868:10.5479/si.03629236.97.i
1618:. They closely resemble
1233:Ocypode madagascariensis
1196:Ocypode madagascariensis
1041:Hoplocypode occidentalis
992:. They closely resemble
957:from Dor-Habonim Beach,
815:Quoy & Gaimard, 1824
773:); and eastwards toward
354:described by the German
2337:Lynne Matthews (2007).
2316:10.1163/156854088x00546
1964:Ocypode ceratophthalmus
1927:The Biological Bulletin
1775:"Revision of the genus
550:to the tip of southern
542:and one in the eastern
467:The claw-bearing legs (
375:into a separate genus,
2408:eating a moon crab in
2406:Ocypode ceratophthalma
2228:Sharon Edgley (2011).
2116:Ocypode ceratophthalma
2114:"Horn-eyed ghost crab
1918:Ocypode ceratophthalma
1643:
1559:
1454:
1369:
1240:
1188:
1166:H. Milne-Edwards, 1852
1153:
1104:
1065:
1010:
974:H. Milne-Edwards, 1837
961:
904:
855:
802:
791:Ocypode ceratophthalma
753:Ocypode ceratophthalma
745:
694:H. Milne-Edwards, 1837
681:
593:were determined to be
351:Cancer ceratophthalmus
208:Cancer ceratophthalmus
2625:Paleobiology Database
2277:on September 24, 2015
1629:
1545:
1440:
1388:Medium-sized species
1355:
1230:
1187:
1143:
1103:
1051:
1003:
951:
890:
841:
788:
731:
670:
487:Further information:
307:. They are primarily
2298:Ocypode gaudichaudii
2187:(Family Ocypodidae)"
2162:on November 12, 2013
2092:University of Hawaii
2042:. pp. 356–411.
1677:- the semaphore crab
1410:Atlantic ghost crab
1149:from Palabuhanratu,
1054:Ocypode gaudichaudii
1018:Ocypode gaudichaudii
381:. It belongs to the
2410:Saint John's Island
2156:"Golden Ghost Crab"
2094:. November 10, 2009
1997:Pat Garber (2006).
1707:. Suppl. 21: 1–109.
1332:, being found from
1297:Malaysian Peninsula
1221:Natal, South Africa
1073:Ocypode jousseaumei
1027:painted ghost crab
734:Ocypode brevicornis
689:Ocypode brevicornis
644:African ghost crab
418:Ocypode brevicornis
345:in 1795, using the
58:Horned ghost crab (
2009:. pp. 94–98.
2007:Parkway Publishers
1665:Crustaceans portal
1644:
1560:
1455:
1370:
1340:to Madagascar and
1338:Great Barrier Reef
1325:pallid ghost crab
1265:O. ceratophthalmus
1241:
1189:
1154:
1121:Kuhl's ghost crab
1105:
1066:
1011:
962:
929:tufted ghost crab
905:
893:Ocypode cordimanus
863:Ocypode cordimanus
856:
803:
796:Chagos Archipelago
746:
682:
515:of small animals.
359:Peter Simon Pallas
2689:
2688:
2612:Open Tree of Life
2425:Taxon identifiers
2385:978-1-77007-633-4
2372:Struik Publishers
2348:978-1-77009-248-8
2232:Ocypode cordimana
2049:978-0-495-56197-2
2016:978-1-933251-31-8
1983:10.1111/bij.12039
1648:
1647:
1632:Ocypode stimpsoni
1601:
1595:Ocypode stimpsoni
1573:
1512:
1489:), including the
1475:Arabian Peninsula
1468:
1462:Ocypode rotundata
1423:Rio Grande do Sul
1407:
1406:(Fabricius, 1787)
1383:
1358:Ocypode pallidula
1322:
1316:Ocypode pallidula
1286:
1254:
1202:
1167:
1161:Ocypode macrocera
1118:
1079:
1063:Galapagos Islands
1024:
975:
934:Mediterranean Sea
926:
869:
824:Western Australia
816:
779:Clipperton Island
759:
695:
641:
599:O. ceratophthalma
540:Mediterranean Sea
424:O. ceratophthalma
280:
279:
274:
266:
258:
250:
194:
16:(Redirected from
2714:
2682:
2681:
2669:
2668:
2656:
2655:
2646:
2645:
2633:
2632:
2620:
2619:
2607:
2606:
2597:
2596:
2584:
2583:
2581:NHMSYS0020704371
2571:
2570:
2558:
2557:
2545:
2544:
2532:
2531:
2519:
2518:
2506:
2505:
2493:
2492:
2480:
2479:
2467:
2466:
2465:
2452:
2451:
2450:
2420:
2390:
2389:
2369:
2359:
2353:
2352:
2334:
2328:
2327:
2293:
2287:
2286:
2284:
2282:
2259:
2253:
2252:
2250:
2248:
2225:
2219:
2218:
2216:
2205:
2199:
2198:
2196:
2194:
2178:
2172:
2171:
2169:
2167:
2151:
2145:
2144:
2142:
2140:
2128:
2122:
2121:
2110:
2104:
2103:
2101:
2099:
2080:
2074:
2073:
2071:
2060:
2054:
2053:
2040:Cengage Learning
2038:(3rd ed.).
2027:
2021:
2020:
1994:
1988:
1987:
1985:
1957:
1951:
1950:
1924:
1911:
1905:
1904:
1888:
1878:
1872:
1871:
1848:
1837:
1836:
1834:
1832:
1812:DecaNet (2023).
1809:
1798:
1797:
1787:
1770:
1709:
1708:
1700:
1691:
1667:
1662:
1661:
1612:Korean Peninsula
1599:
1571:
1515:pink ghost crab
1510:
1466:
1443:Ocypode quadrata
1405:
1401:Ocypode quadrata
1381:
1377:Ocypode pauliani
1320:
1293:Gulf of Thailand
1284:
1252:
1200:
1165:
1134:Papua New Guinea
1116:
1077:
1022:
1006:Ocypode fabricii
973:
969:Ocypode fabricii
916:
881:French Polynesia
867:
814:
757:
693:
673:Ocypode africana
639:
635:Ocypode africana
616:
500:areas. They are
272:
264:
256:
248:
189:
85:
84:
53:
39:
21:
2722:
2721:
2717:
2716:
2715:
2713:
2712:
2711:
2692:
2691:
2690:
2685:
2677:
2672:
2664:
2659:
2651:
2649:
2641:
2636:
2628:
2623:
2615:
2610:
2602:
2600:
2592:
2587:
2579:
2574:
2566:
2561:
2553:
2548:
2540:
2535:
2527:
2522:
2514:
2509:
2501:
2496:
2488:
2483:
2475:
2470:
2461:
2460:
2455:
2446:
2445:
2440:
2427:
2398:
2393:
2386:
2361:
2360:
2356:
2349:
2336:
2335:
2331:
2295:
2294:
2290:
2280:
2278:
2261:
2260:
2256:
2246:
2244:
2227:
2226:
2222:
2214:
2207:
2206:
2202:
2192:
2190:
2185:Ocypode convexa
2180:
2179:
2175:
2165:
2163:
2153:
2152:
2148:
2138:
2136:
2130:
2129:
2125:
2112:
2111:
2107:
2097:
2095:
2082:
2081:
2077:
2069:
2062:
2061:
2057:
2050:
2029:
2028:
2024:
2017:
1996:
1995:
1991:
1959:
1958:
1954:
1939:10.2307/1539309
1922:
1913:
1912:
1908:
1901:
1880:
1879:
1875:
1852:Mary J. Rathbun
1850:
1849:
1840:
1830:
1828:
1811:
1810:
1801:
1785:
1772:
1771:
1712:
1698:
1693:
1692:
1688:
1684:
1663:
1656:
1653:
1598:
1572:(Forskål, 1775)
1570:
1567:Ocypode saratan
1534:, and also the
1509:
1465:
1404:
1380:
1346:cryptic species
1321:Jacquinot, 1846
1319:
1283:
1280:Ocypode nobilii
1251:
1248:Ocypode mortoni
1199:
1164:
1115:
1076:
1021:
972:
915:
898:New South Wales
868:Latreille, 1818
866:
844:Ocypode convexa
813:
810:Ocypode convexa
756:
714:Nicobar Islands
692:
638:
579:O. longicornuta
568:O. occidentalis
560:
521:
491:
485:
430:O. gaudichaudii
402:
373:gulf ghost crab
343:Friedrich Weber
332:
305:intertidal zone
214:
211:
188:
79:
35:
28:
23:
22:
15:
12:
11:
5:
2720:
2718:
2710:
2709:
2707:Decapod genera
2704:
2694:
2693:
2687:
2686:
2684:
2683:
2670:
2657:
2647:
2634:
2621:
2608:
2598:
2585:
2572:
2559:
2546:
2533:
2520:
2507:
2494:
2481:
2468:
2453:
2437:
2435:
2429:
2428:
2423:
2417:
2416:
2397:
2396:External links
2394:
2392:
2391:
2384:
2354:
2347:
2329:
2310:(2): 217–219.
2288:
2265:Ocypode cursor
2254:
2220:
2200:
2173:
2146:
2123:
2105:
2075:
2055:
2048:
2022:
2015:
1989:
1976:(2): 257–270.
1952:
1933:(3): 407–413.
1906:
1899:
1873:
1838:
1799:
1710:
1685:
1683:
1680:
1679:
1678:
1669:
1668:
1652:
1649:
1646:
1645:
1623:
1604:
1602:
1590:
1589:
1587:
1576:
1574:
1562:
1561:
1548:Ocypode ryderi
1539:
1528:Port Elizabeth
1516:
1513:
1511:Kingsley, 1880
1506:Ocypode ryderi
1501:
1500:
1498:
1471:
1469:
1457:
1456:
1434:
1411:
1408:
1396:
1395:
1393:
1386:
1384:
1382:Crosnier, 1965
1372:
1371:
1349:
1326:
1323:
1311:
1310:
1308:
1289:
1287:
1275:
1274:
1272:
1257:
1255:
1243:
1242:
1224:
1205:
1203:
1201:Crosnier, 1965
1191:
1190:
1181:
1170:
1168:
1156:
1155:
1146:Ocypode kuhlii
1137:
1122:
1119:
1112:Ocypode kuhlii
1107:
1106:
1097:
1082:
1080:
1078:(Nobili, 1905)
1068:
1067:
1045:
1028:
1025:
1013:
1012:
997:
994:O. jousseaumei
978:
976:
964:
963:
954:Ocypode cursor
945:
930:
927:
912:Ocypode cursor
907:
906:
884:
873:
870:
858:
857:
835:
820:
817:
805:
804:
782:
763:
760:
758:(Pallas, 1772)
748:
747:
725:
698:
696:
684:
683:
664:
645:
642:
630:
629:
626:
623:
620:
613:respectively.
603:O. brevicornis
583:O. platytarsis
559:
556:
536:Atlantic Ocean
520:
517:
484:
481:
401:
398:
331:
328:
278:
277:
276:
275:
267:
259:
251:
246:Ceratophthalma
240:
239:
233:
232:
223:
222:
216:
215:
212:
203:
202:
196:
195:
181:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
141:
137:
136:
131:
127:
126:
121:
117:
116:
111:
107:
106:
101:
97:
96:
91:
87:
86:
73:
72:
55:
54:
46:
45:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
2719:
2708:
2705:
2703:
2700:
2699:
2697:
2680:
2675:
2671:
2667:
2662:
2658:
2654:
2648:
2644:
2639:
2635:
2631:
2626:
2622:
2618:
2613:
2609:
2605:
2599:
2595:
2590:
2586:
2582:
2577:
2573:
2569:
2564:
2560:
2556:
2551:
2547:
2543:
2538:
2534:
2530:
2525:
2521:
2517:
2512:
2508:
2504:
2499:
2495:
2491:
2486:
2482:
2478:
2473:
2469:
2464:
2458:
2454:
2449:
2443:
2439:
2438:
2436:
2434:
2430:
2426:
2421:
2415:
2411:
2407:
2403:
2400:
2399:
2395:
2387:
2381:
2377:
2373:
2368:
2367:
2358:
2355:
2350:
2344:
2340:
2333:
2330:
2325:
2321:
2317:
2313:
2309:
2305:
2304:
2299:
2292:
2289:
2276:
2272:
2268:
2266:
2258:
2255:
2243:
2239:
2235:
2233:
2224:
2221:
2213:
2212:
2204:
2201:
2188:
2186:
2177:
2174:
2161:
2157:
2150:
2147:
2134:
2127:
2124:
2119:
2117:
2109:
2106:
2093:
2089:
2085:
2084:"Ghost crabs"
2079:
2076:
2068:
2067:
2059:
2056:
2051:
2045:
2041:
2037:
2033:
2026:
2023:
2018:
2012:
2008:
2004:
2000:
1993:
1990:
1984:
1979:
1975:
1971:
1967:
1965:
1956:
1953:
1948:
1944:
1940:
1936:
1932:
1928:
1921:
1919:
1910:
1907:
1902:
1900:9780643069060
1896:
1892:
1887:
1886:
1877:
1874:
1869:
1865:
1862:(97): 1–461.
1861:
1857:
1853:
1847:
1845:
1843:
1839:
1827:
1823:
1819:
1817:
1808:
1806:
1804:
1800:
1796:(2): 665–793.
1795:
1791:
1784:
1782:
1778:
1769:
1767:
1765:
1763:
1761:
1759:
1757:
1755:
1753:
1751:
1749:
1747:
1745:
1743:
1741:
1739:
1737:
1735:
1733:
1731:
1729:
1727:
1725:
1723:
1721:
1719:
1717:
1715:
1711:
1706:
1705:
1697:
1690:
1687:
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1676:
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1670:
1666:
1660:
1655:
1650:
1642:
1638:
1634:
1633:
1628:
1624:
1621:
1617:
1613:
1609:
1605:
1603:
1600:Ortmann, 1897
1597:
1596:
1592:
1591:
1588:
1585:
1584:O. cordimanus
1581:
1577:
1575:
1569:
1568:
1564:
1563:
1558:
1554:
1550:
1549:
1544:
1540:
1537:
1533:
1529:
1525:
1521:
1517:
1514:
1508:
1507:
1503:
1502:
1499:
1496:
1492:
1488:
1484:
1480:
1476:
1472:
1470:
1464:
1463:
1459:
1458:
1453:
1449:
1445:
1444:
1439:
1435:
1432:
1428:
1424:
1420:
1419:United States
1416:
1415:Massachusetts
1412:
1409:
1403:
1402:
1398:
1397:
1394:
1391:
1387:
1385:
1379:
1378:
1374:
1373:
1368:
1364:
1360:
1359:
1354:
1350:
1347:
1343:
1339:
1335:
1331:
1327:
1324:
1318:
1317:
1313:
1312:
1309:
1306:
1302:
1298:
1294:
1290:
1288:
1282:
1281:
1277:
1276:
1273:
1270:
1266:
1262:
1258:
1256:
1250:
1249:
1245:
1244:
1239:
1235:
1234:
1229:
1225:
1222:
1218:
1214:
1210:
1206:
1204:
1198:
1197:
1193:
1192:
1186:
1182:
1179:
1175:
1171:
1169:
1163:
1162:
1158:
1157:
1152:
1148:
1147:
1142:
1138:
1135:
1131:
1127:
1123:
1120:
1117:De Haan, 1835
1114:
1113:
1109:
1108:
1102:
1098:
1095:
1091:
1087:
1083:
1081:
1075:
1074:
1070:
1069:
1064:
1060:
1056:
1055:
1050:
1046:
1043:
1042:
1037:
1033:
1029:
1026:
1020:
1019:
1015:
1014:
1008:
1007:
1002:
998:
995:
991:
987:
983:
979:
977:
971:
970:
966:
965:
960:
956:
955:
950:
946:
943:
939:
935:
931:
928:
924:
920:
914:
913:
909:
908:
903:
899:
895:
894:
889:
885:
882:
878:
874:
871:
865:
864:
860:
859:
854:
850:
846:
845:
840:
836:
833:
829:
825:
821:
818:
812:
811:
807:
806:
801:
797:
793:
792:
787:
783:
780:
776:
772:
768:
764:
761:
755:
754:
750:
749:
744:
740:
736:
735:
730:
726:
723:
719:
715:
711:
707:
703:
699:
697:
691:
690:
686:
685:
679:
675:
674:
669:
665:
662:
658:
654:
650:
646:
643:
637:
636:
632:
631:
627:
624:
622:Common names
621:
618:
617:
614:
612:
611:O. cordimanus
608:
604:
600:
596:
592:
588:
584:
580:
576:
575:
570:
569:
564:
557:
555:
553:
549:
545:
541:
537:
533:
529:
525:
518:
516:
514:
510:
507:
503:
499:
495:
490:
482:
480:
478:
474:
470:
465:
462:
461:
456:
455:
450:
449:
444:
443:
438:
437:
432:
431:
426:
425:
420:
419:
414:
410:
406:
399:
397:
395:
391:
387:
384:
380:
379:
374:
370:
366:
362:
360:
357:
353:
352:
348:
344:
341:
337:
329:
327:
325:
321:
317:
314:
310:
306:
302:
298:
294:
290:
286:
285:
273:Neumann, 1878
271:
268:
265:Lamarck, 1801
263:
260:
255:
252:
249:MacLeay, 1838
247:
244:
243:
241:
238:
234:
231:
230:
224:
221:
217:
210:
209:
204:
201:
197:
192:
187:
186:
182:
179:
178:
175:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
145:
142:
139:
138:
135:
132:
129:
128:
125:
122:
119:
118:
115:
112:
109:
108:
105:
102:
99:
98:
95:
92:
89:
88:
83:
78:
74:
71:
67:
63:
62:
56:
52:
47:
44:
40:
37:
33:
19:
2432:
2405:
2365:
2357:
2338:
2332:
2307:
2301:
2297:
2291:
2279:. Retrieved
2275:the original
2270:
2264:
2257:
2245:. Retrieved
2237:
2231:
2223:
2210:
2203:
2193:November 12,
2191:. Retrieved
2184:
2176:
2166:November 12,
2164:. Retrieved
2160:the original
2149:
2139:November 12,
2137:. Retrieved
2126:
2115:
2108:
2096:. Retrieved
2087:
2078:
2065:
2058:
2035:
2025:
2002:
1992:
1973:
1969:
1963:
1955:
1930:
1926:
1917:
1909:
1884:
1876:
1859:
1829:. Retrieved
1821:
1818:Weber, 1795"
1815:
1793:
1789:
1780:
1776:
1702:
1689:
1672:
1630:
1619:
1593:
1583:
1565:
1546:
1532:South Africa
1522:of southern
1504:
1494:
1491:Persian Gulf
1477:to northern
1460:
1448:Ormond Beach
1441:
1430:
1399:
1375:
1356:
1329:
1314:
1305:O. stimpsoni
1304:
1299:to northern
1285:De Man, 1902
1278:
1268:
1264:
1261:O. stimpsoni
1260:
1253:George, 1982
1246:
1231:
1194:
1159:
1144:
1110:
1093:
1090:Gulf of Oman
1086:Gulf of Aden
1071:
1052:
1039:
1016:
1004:
993:
967:
952:
940:to southern
910:
891:
876:
861:
842:
808:
789:
766:
751:
732:
717:
687:
671:
660:
640:De Man, 1881
633:
625:Description
610:
606:
602:
598:
590:
586:
582:
578:
572:
566:
562:
561:
548:Indian Ocean
523:
522:
519:Distribution
493:
492:
473:stridulating
466:
458:
452:
448:O. rotundata
446:
440:
436:O. macrocera
434:
428:
422:
416:
404:
403:
393:
389:
376:
364:
363:
349:
347:type species
340:entomologist
335:
333:
283:
282:
281:
269:
261:
253:
245:
226:
213:Pallas, 1772
206:
200:Type species
193:, 1795
184:
183:
150:Infraorder:
124:Malacostraca
59:
42:
36:
2702:Ocypodoidea
2537:iNaturalist
2457:Wikispecies
2303:Crustaceana
2271:SeaLifeBase
1781:Hoplocypode
1520:Abd al Kuri
1467:Miers, 1882
1094:O. fabricii
678:Congo River
591:O. sinensis
587:O. pygoides
574:Hoplocypode
532:subtropical
400:Description
390:Hoplocypode
378:Hoplocypode
301:subtropical
293:ghost crabs
170:Subfamily:
144:Pleocyemata
2696:Categories
2374:. p.
2247:August 12,
2098:January 2,
1831:24 October
1682:References
1620:O. mortoni
1536:Seychelles
1495:O. saratan
1295:, and the
1269:O. mortoni
1238:Madagascar
1217:Mozambique
1209:Madagascar
1059:Santa Cruz
712:, and the
649:Mauritania
607:O. convexa
509:scavengers
506:generalist
489:Ghost crab
477:pereiopods
454:O. saratan
442:O. mortoni
386:Ocypodidae
369:Ocypodinae
356:naturalist
334:The genus
316:scavengers
313:generalist
311:, and are
270:Parocypoda
174:Ocypodinae
164:Ocypodidae
140:Suborder:
114:Arthropoda
2448:Q13908705
2414:Singapore
2281:August 4,
1920:(Pallas)"
1674:Heloecius
1367:Australia
1342:Mauritius
1151:West Java
1130:Indonesia
1032:Guatemala
990:Shark Bay
902:Australia
853:Australia
794:from the
775:Polynesia
710:Sri Lanka
676:from the
661:O. cursor
657:sympatric
577:in 2013.
513:predators
502:nocturnal
498:estuarine
469:chelipeds
460:O. cursor
361:in 1772.
320:predators
309:nocturnal
257:Say, 1817
254:Monolepis
154:Brachyura
100:Kingdom:
94:Eukaryota
2498:BugGuide
2442:Wikidata
2324:20104392
1854:(1918).
1651:See also
1637:Nagasaki
1557:Tanzania
1553:Zanzibar
1174:Pakistan
1126:Thailand
1088:and the
982:Northern
919:Linnaeus
849:Gnaraloo
655:. It is
619:Species
595:synonyms
538:and the
528:tropical
504:and are
409:carapace
330:Taxonomy
297:tropical
237:Synonyms
160:Family:
134:Decapoda
110:Phylum:
104:Animalia
90:Domain:
2674:ZooBank
2653:4263161
2555:1401318
2529:2221525
2477:Ocypode
2463:Ocypode
2433:Ocypode
1947:1539309
1816:Ocypode
1777:Ocypode
1580:Red Sea
1524:Somalia
1452:Florida
1431:Ocypode
1417:in the
1390:endemic
1336:to the
1330:Ocypode
1178:Myanmar
936:, from
877:Ocypode
826:, from
771:Red Sea
767:Ocypode
739:Chennai
722:propodi
718:Ocypode
680:estuary
653:Namibia
563:Ocypode
558:Species
544:Pacific
524:Ocypode
494:Ocypode
483:Ecology
405:Ocypode
394:Ocypode
365:Ocypode
336:Ocypode
324:species
284:Ocypode
262:Ocypoda
220:Species
185:Ocypode
180:Genus:
130:Order:
120:Class:
64:) from
43:Ocypode
32:Ocypete
18:Ocypoda
2666:106970
2650:uBio:
2617:509870
2601:NZOR:
2503:354361
2382:
2345:
2322:
2046:
2013:
1945:
1897:
1614:, and
1610:, the
1487:Bombay
1481:(from
1427:Brazil
1334:Hawaii
1301:Borneo
1215:(from
1213:Africa
1132:, and
986:Darwin
959:Israel
942:Greece
828:Broome
628:Image
552:Africa
451:, and
413:cornea
383:family
70:Hawaii
2661:WoRMS
2638:Plazi
2630:22582
2594:53309
2568:99081
2550:IRMNG
2542:53738
2516:7PMY7
2490:84193
2402:Video
2320:JSTOR
2215:(PDF)
2070:(PDF)
1943:JSTOR
1923:(PDF)
1822:WoRMS
1786:(PDF)
1699:(PDF)
1641:Japan
1635:from
1616:Japan
1608:China
1551:from
1479:India
1446:from
1361:from
1236:from
1057:from
1036:Chile
938:Egypt
896:from
847:from
832:Perth
743:India
737:from
706:India
659:with
289:genus
287:is a
191:Weber
2589:NCBI
2563:ITIS
2524:GBIF
2485:BOLD
2380:ISBN
2343:ISBN
2283:2013
2249:2012
2195:2013
2168:2013
2141:2013
2100:2010
2044:ISBN
2011:ISBN
1895:ISBN
1833:2023
1483:Oman
923:1758
777:and
702:Oman
609:and
589:and
530:and
511:and
318:and
299:and
229:text
227:See
66:Kona
2576:NBN
2511:CoL
2472:AFD
2404:of
2312:doi
1978:doi
1974:109
1935:doi
1931:126
1891:496
1864:doi
1530:of
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