731:
750:
130:
102:
1359:
773:
272:
Myxophaga have several diagnostic features: the antennae are more or less distinctly clubbed with usually fewer than nine segments, mesocoxal cavities are open laterally and bordered by a mesepimeron and metanepisternum, the hind wings are rolled apically in the resting positions. Internally, they
604:
Living members of
Lepiceridae are confined to northern South America and Central America. Members of Sphaeriusidae occur on all continents except Antarctica, while Hydroscaphidae occurs on all continents except Australia and Antarctica. Torridincolidae occurs in Africa, Asia, and South America.
730:
632:) aged sediments in Poland. It was placed in its own monotypic family Triamyxidae, and was resolved as either the most basal myxophagan or sister to Hydroscaphidae. The next oldest is
941:
Fikáček, Martin; Beutel, Rolf G.; Cai, Chenyang; Lawrence, John F.; Newton, Alfred F.; Solodovnikov, Alexey; Ślipiński, Adam; Thayer, Margaret K.; Yamamoto, Shûhei (January 2020).
1130:
Qvarnström, Martin; Fikáček, Martin; Vikberg
Wernström, Joel; Huld, Sigrid; Beutel, Rolf G.; Arriaga-Varela, Emmanuel; Ahlberg, Per E.; Niedźwiedzki, Grzegorz (June 2021).
292:
There are four extant families in the suborder
Myxophaga divided between two superfamilies, containing about 65 described species, and at least one extinct family.
3457:
3278:
799:
3405:
3431:
1221:
Cai, C.; Short, A. E. Z.; Huang, D. (2012). "The First Skiff Beetle (Coleoptera: Myxophaga: Hydroscaphidae) from Early
Cretaceous Jehol Biota".
1001:"Phylogeny and systematics of Sphaeriusidae (Coleoptera: Myxophaga): minute living fossils with underestimated past and present-day diversity"
1346:
1200:
749:
650:
but in 2019 was reinterpreted as an early diverging relative of the family
Hydroscaphidae. A fossil impression assigned to the living genus
1391:
772:
3508:
1264:
Kirejtshuk, A. G.; Poinar, G. (2006). "Haplochelidae, a new family of cretaceous beetles (Coleoptera: Myxophaga) from
Burmese amber".
943:"Reliable placement of beetle fossils via phylogenetic analyses – Triassic Leehermania as a case study (Staphylinidae or Myxophaga?)"
3513:
856:
3436:
3444:
1313:
JaĹ‚oszyĹ„ski, PaweĹ‚; Luo, Xiao-Zhu; Hammel, Jörg U.; Yamamoto, ShĂ»hei & Beutel, Rolf G. (2020). "The mid-Cretaceous â€
1080:"New fossils of Sphaeriusidae from mid-Cretaceous Burmese amber revealed by confocal microscopy (Coleoptera: Myxophaga)"
3340:
531:
1284:
839:
Beutel, R.; Leschen, R. (2005). "Morphology and
Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim)".
3384:
419:
352:
129:
3462:
3503:
3397:
3518:
1384:
274:
478:
3293:
614:
1285:"On the systematic position and taxonomic rank of the extinct myxophagan dagger Haplochelus (Coleoptera)"
3470:
365:
1317:
gen. nov. and the evolution of the relict beetle family
Lepiceridae (Insecta: Coleoptera: Myxophaga)".
999:
Fikáček, Martin; Yamamoto, Shûhei; Matsumoto, Keita; Beutel, Rolf G.; Maddison, David R. (2022-12-07).
1051:"A new genus and species of Sphaeriusidae (Coleoptera, Myxophaga) from Lower Cretaceous Burmese amber"
280:
Beetles of this suborder are adapted to feed on algae. Their mouthparts are characteristic in lacking
3366:
2530:
1230:
1143:
1091:
954:
643:
1180:
348:
2749:
1377:
712:
462:
3498:
1246:
1028:
978:
706:
688:
634:
589:
451:
335:
326:
124:
700:
440:
3475:
906:
3018:
2435:
1196:
1161:
1109:
1050:
1020:
970:
852:
848:
281:
101:
3115:
2894:
2889:
2812:
2570:
1586:
1326:
1296:
1238:
1188:
1151:
1099:
1012:
962:
914:
665:
657:
613:
The fossil record of myxophagan beetles is sparse, likely due to their small size limiting
2804:
2757:
2602:
2586:
2262:
2022:
1991:
1906:
1824:
1715:
1683:
1132:"Exceptionally preserved beetles in a Triassic coprolite of putative dinosauriform origin"
474:
223:
116:
3331:
1363:
1000:
942:
1234:
1147:
1095:
958:
3392:
3379:
3207:
3158:
3013:
2820:
2488:
2472:
2411:
2306:
2277:
2205:
2169:
2153:
2069:
2014:
1919:
1816:
1665:
1627:
1607:
1503:
1192:
763:
361:
253:
211:
3492:
3189:
2993:
2984:
2725:
2696:
2688:
2680:
2654:
2629:
2525:
2512:
2451:
2369:
2272:
2267:
2148:
2082:
2053:
2037:
2001:
1971:
1924:
1878:
1845:
1761:
1753:
1724:
1562:
1250:
1032:
982:
677:
415:
393:
59:
3166:
3090:
2836:
2828:
2783:
2767:
2717:
2638:
2594:
2546:
2464:
2377:
2246:
2233:
2213:
2174:
2074:
2061:
2032:
1966:
1954:
1850:
1807:
1790:
1782:
1732:
1570:
1540:
1471:
1460:
1444:
907:"Sphaeriusidae (Coleoptera, Myxophaga): A new beetle family to the fauna of Serbia"
549:
539:
516:
432:
313:
300:
296:
245:
1330:
3345:
3449:
3418:
3325:
3252:
3223:
3197:
3184:
3124:
3082:
3066:
3058:
3050:
3034:
3026:
2962:
2920:
2912:
2881:
2796:
2775:
2741:
2733:
2646:
2578:
2562:
2554:
2517:
2504:
2480:
2398:
2390:
2340:
2332:
2290:
2254:
2225:
2197:
2192:
2108:
2103:
2095:
2045:
1981:
1976:
1890:
1855:
1840:
1832:
1748:
1740:
1557:
757:
661:
652:
647:
562:
373:
309:
257:
228:
215:
196:
34:
1358:
3179:
3140:
3132:
3074:
2937:
2899:
2852:
2844:
2664:
2612:
2496:
2427:
2298:
2282:
2238:
2179:
2143:
2123:
2087:
1996:
1986:
1860:
1594:
1578:
1156:
1131:
1104:
1079:
673:
486:
79:
44:
3316:
1369:
1300:
1113:
1024:
974:
646:
of North
Carolina, which had previously been interpreted as the oldest known
3244:
3236:
3228:
3215:
3171:
2970:
2945:
2873:
2701:
2672:
2356:
2187:
2161:
2113:
2027:
1929:
1898:
1870:
1703:
1678:
1673:
1619:
1508:
1491:
1483:
1432:
1420:
780:
738:
694:
682:
625:
427:
383:
321:
219:
161:
141:
84:
1165:
1078:
Li, Yan-Da; Ślipiński, Adam; Huang, Di-Ying; Cai, Chen-Yang (2023-01-11).
919:
3353:
3310:
3148:
3098:
3042:
3001:
2950:
2607:
2538:
2419:
2131:
1937:
1911:
1769:
1602:
1549:
1528:
1438:
619:
573:
526:
403:
110:
74:
69:
54:
49:
39:
3410:
3358:
2865:
2857:
2709:
2443:
2364:
2348:
1016:
966:
629:
496:
89:
64:
3423:
2456:
2316:
1949:
1426:
1414:
1401:
1242:
669:
639:
249:
241:
181:
171:
151:
3287:
884:
506:
261:
3371:
841:
Band 4 Part 38: Arthropoda, Hälfte: Insecta, Coleoptera, Beetles
3291:
1700:
1651:
1525:
1457:
1373:
1772:(furniture beetles, death watch beetles, spider beetles)
624:
described in 2021 from numerous specimens of found in a
1266:
248:, consisting of roughly 65 species of small to minute
680:
from Myanmar, have been assigned to the extant genus
3300:
3206:
3157:
3123:
3114:
2992:
2983:
2936:
2911:
2795:
2766:
2663:
2637:
2628:
2410:
2389:
2315:
2122:
2013:
1869:
1815:
1806:
1781:
1723:
1714:
1539:
252:in four families. The members of this suborder are
284:and having a mobile tooth on their left mandible.
1179:Sampaio, Brunno H.L.; Short, Andrew E.Z. (2018),
2649:(jewel beetles, or metallic wood-boring beetles)
1283:Ge, S. Q.; Friedrich, F.; Beutel, R. G. (2010).
2372:(true weevils, bark beetles, ambrosia beetles)
1385:
1181:"Families Hydroscaphidae and Torridincolidae"
871:
8:
2615:(ironclad beetles, cylindrical bark beetles)
1185:Thorp and Covich's Freshwater Invertebrates
672:). Fossils from the early Late Cretaceous (
3288:
3120:
2989:
2634:
1812:
1720:
1711:
1697:
1648:
1536:
1522:
1454:
1392:
1378:
1370:
692:within Lepiceridae, as well as the extant
100:
20:
3279:List of subgroups of the order Coleoptera
2359:(straight snout weevils, New York weevil)
1155:
1103:
918:
845:Handbuch der Zoologie/Handbook of Zoology
800:List of subgroups of the order Coleoptera
511:Py-Daniel, da Fonseca & Barbosa, 1993
277:and the testes are tube-like and coiled.
273:are characterised by the presence of six
2786:(cicada beetle, cicada parasite beetles)
408:Short, Joly, GarcĂa & Maddison, 2015
831:
811:
726:
240:is the second-smallest suborder of the
2876:(false fireflies, long-lipped beetles)
911:Bulletin of the Natural History Museum
1125:
1123:
617:. The currently oldest myxophagan is
7:
3398:42f89ae7-71c5-45ac-8492-825030358e4e
2902:(false metallic wood-boring beetles)
1073:
1071:
1044:
1042:
994:
992:
936:
934:
932:
930:
656:in Hydroscaphidae is known from the
1319:Journal of Systematic Palaeontology
818:†Denotes that the group is extinct
1193:10.1016/b978-0-12-804223-6.00022-6
14:
2513:Perimylopidae, or Promecheilidae
2098:(minute brown scavenger beetles)
1357:
1289:Insect Systematics and Evolution
887:. Tree of Life Web Project. 2007
771:
748:
743:(Sphaeriusidae) in various views
729:
128:
2728:(variegated mud-loving beetles)
1187:, Elsevier, pp. 519–525,
467:Li & Cai in Li et al. 2023
1:
2557:(narrow-waisted bark beetles)
2249:(parasitic flat bark beetles)
2200:(short-winged flower beetles)
2056:(lady beetles, or God's cows)
1793:(tooth-necked fungus beetles)
1331:10.1080/14772019.2020.1747561
716:, belonging to Sphaerusidae.
3069:(sand-loving scarab beetles)
2459:(minute tree-fungus beetles)
2285:(silvanid flat bark beetles)
398:Reichardt & Hinton, 1976
3218:(primitive carrion beetles)
3174:(minute mud-loving beetles)
3037:(bumble bee scarab beetles)
2807:(soft-bodied plant beetles)
2778:(soft bodied plant beetles)
1581:(predaceous diving beetles)
491:Reichardt & Vanin, 1976
3535:
3053:(scavenger scarab beetles)
3045:(enigmatic scarab beetles)
2218:Lamingtonium binnaberrense
1932:(soft-wing flower beetles)
1084:Frontiers in Earth Science
1049:Kirejtshuk, A. G. (2009).
3509:Beetles described in 1955
3276:
3192:(water scavenger beetles)
2925:Rhinorhipus tamborinensis
2704:(long-toed water beetles)
2597:(polypore fungus beetles)
2499:(palm and flower beetles)
2483:(tumbling flower beetles)
2430:(ant-like flower beetles)
2208:(lined flat bark beetles)
2182:(pleasing fungus beetles)
2077:(handsome fungus beetles)
1710:
1696:
1661:
1647:
1622:(burrowing water beetles)
1535:
1521:
1467:
1453:
1409:
1351:Atlas of Living Australia
1157:10.1016/j.cub.2021.05.015
1105:10.3389/feart.2022.901573
872:Beutel & Leschen 2005
594:Chatzimanolis et al. 2012
209:
204:
125:Scientific classification
123:
108:
99:
23:
3514:Beetles of North America
3239:(feather-winged beetles)
2573:(false longhorn beetles)
2520:(jugular-horned beetles)
2467:(false darkling beetles)
2438:(cryptic fungus beetles)
2257:(shining flower beetles)
1597:(crawling water beetles)
1496:Sikhotealinia zhiltzovae
1301:10.1163/187631210X537385
638:from the Late Triassic (
628:found in Late Triassic (
2720:(forest stream beetles)
2581:(synchroa bark beetles)
2541:(dead log bark beetles)
2533:(fire-coloured beetles)
2422:(ant-like leaf beetles)
2156:(silken fungus beetles)
2090:(well polished beetles)
2064:(minute fungus beetles)
1883:Acanthocnemus nigricans
1223:Journal of Paleontology
905:Mesaros, Gabor (2013).
340:Jałoszyński et al. 2020
3231:(round fungus beetles)
2565:(false flower beetles)
2549:(wedge-shaped beetles)
2507:(false blister beetle)
2491:(hairy fungus beetles)
2446:(conifer bark beetles)
2343:(leaf-rolling weevils)
2004:(bark-gnawing beetles)
1764:(wounded-tree beetles)
1630:(false ground beetles)
1552:(trout-stream beetles)
615:preservation potential
578:Qvarnström et al. 2021
566:Qvarnström et al. 2021
3471:Paleobiology Database
3143:(false clown beetles)
2965:(plate-thigh beetles)
2847:(false click beetles)
2752:(water-penny beetles)
2401:(ship-timber beetles)
2380:(pine flower weevils)
2228:(root-eating beetles)
2048:(minute bark beetles)
1959:Phloiophilus edwardsi
1486:(reticulated beetles)
1005:Systematic Entomology
947:Systematic Entomology
920:10.5937/bnhmb1306071m
3393:Fauna Europaea (new)
3021:(false stag beetles)
3006:Belohina inexpectata
2868:(net-winged beetles)
2831:(rare click beetles)
2744:(travertine beetles)
2736:(minute mud beetles)
2301:(dry-fungus beetles)
2136:Agapytho foveicollis
1893:(false skin beetles)
1756:(Jacobson's beetles)
1476:Crowsoniella relicta
1142:(15): 3374–3381.e5.
644:Cow Branch Formation
3019:Diphyllostomatidae
2351:(primitive weevils)
2164:(flat bark beetles)
1914:(checkered beetles)
1901:(fruitworm beetles)
1589:(whirligig beetles)
1235:2012JPal...86..116C
1148:2021CBio...31E3374Q
1096:2023FrEaS..10.1573L
959:2020SysEn..45..175F
583:Unplaced in family
445:Fikáček et al. 2022
16:Suborder of beetles
2884:(glowworm beetles)
2589:(darkling beetles)
2436:Archeocrypticidae
2293:(palmetto beetles)
2040:(dry bark beetles)
1827:(longhorn beetles)
1017:10.1111/syen.12571
967:10.1111/syen.12386
686:and extinct genus
664:, dating from the
275:malpighian tubules
3486:
3485:
3294:Taxon identifiers
3285:
3284:
3272:
3271:
3268:
3267:
3264:
3263:
3256:
3248:
3247:(carrion beetles)
3240:
3232:
3219:
3193:
3175:
3144:
3136:
3110:
3109:
3102:
3094:
3086:
3078:
3070:
3062:
3054:
3046:
3038:
3030:
3022:
3009:
2979:
2978:
2966:
2958:
2928:
2903:
2885:
2877:
2869:
2861:
2848:
2840:
2832:
2824:
2823:(soldier beetles)
2816:
2813:Brachypsectridae
2808:
2787:
2779:
2753:
2745:
2737:
2729:
2721:
2713:
2705:
2692:
2684:
2676:
2650:
2624:
2623:
2616:
2598:
2590:
2582:
2574:
2571:Stenotrachelidae
2566:
2558:
2550:
2542:
2534:
2521:
2508:
2500:
2492:
2484:
2476:
2475:(blister beetles)
2468:
2460:
2447:
2439:
2431:
2423:
2402:
2381:
2373:
2360:
2352:
2344:
2336:
2326:
2302:
2294:
2286:
2258:
2250:
2242:
2229:
2221:
2209:
2201:
2183:
2165:
2157:
2139:
2099:
2091:
2078:
2065:
2057:
2049:
2041:
2023:Akalyptoischiidae
2005:
1962:
1945:
1933:
1915:
1902:
1894:
1886:
1836:
1828:
1802:
1801:
1794:
1773:
1765:
1757:
1744:
1736:
1692:
1691:
1669:
1643:
1642:
1639:
1638:
1631:
1623:
1615:
1598:
1590:
1582:
1574:
1566:
1553:
1517:
1516:
1499:
1487:
1479:
1202:978-0-12-804223-6
849:Walter de Gruyter
595:
579:
567:
555:
545:
535:
522:
512:
502:
492:
482:
468:
457:
446:
436:
423:
409:
399:
389:
379:
369:
356:
341:
330:
317:
304:
235:
234:
200:
3526:
3504:Insect suborders
3479:
3478:
3466:
3465:
3453:
3452:
3450:NHMSYS0001719091
3440:
3439:
3427:
3426:
3414:
3413:
3401:
3400:
3388:
3387:
3375:
3374:
3362:
3361:
3349:
3348:
3336:
3335:
3334:
3321:
3320:
3319:
3289:
3254:
3246:
3238:
3230:
3217:
3191:
3173:
3142:
3134:
3121:
3116:Staphyliniformia
3100:
3092:
3084:
3076:
3068:
3060:
3052:
3044:
3036:
3028:
3020:
3003:
2990:
2964:
2955:Declinia relicta
2952:
2922:
2901:
2895:Sinopyrophoridae
2890:Rhagophthalmidae
2883:
2875:
2867:
2859:
2846:
2838:
2830:
2822:
2814:
2806:
2805:Artematopodidae
2785:
2777:
2751:
2743:
2735:
2727:
2719:
2712:(riffle beetles)
2711:
2703:
2691:(turtle beetles)
2690:
2682:
2674:
2648:
2635:
2614:
2596:
2588:
2580:
2572:
2564:
2556:
2548:
2540:
2532:
2519:
2506:
2498:
2490:
2482:
2474:
2466:
2458:
2445:
2437:
2429:
2421:
2400:
2379:
2371:
2358:
2350:
2342:
2335:(fungus weevils)
2334:
2325:
2324:
2320:
2307:Tasmosalpingidae
2300:
2292:
2284:
2256:
2248:
2240:
2227:
2215:
2207:
2199:
2181:
2163:
2155:
2133:
2097:
2089:
2076:
2063:
2055:
2047:
2039:
2003:
1956:
1939:
1931:
1913:
1900:
1892:
1880:
1879:Acanthocnemidae
1834:
1826:
1813:
1792:
1771:
1763:
1755:
1742:
1734:
1721:
1712:
1698:
1667:
1649:
1629:
1621:
1609:
1596:
1588:
1580:
1572:
1565:(ground beetles)
1564:
1551:
1537:
1523:
1493:
1485:
1473:
1472:Crowsoniellidae
1455:
1394:
1387:
1380:
1371:
1362:Data related to
1361:
1354:
1333:
1311:
1305:
1304:
1280:
1274:
1273:
1261:
1255:
1254:
1243:10.1666/11-050.1
1218:
1212:
1211:
1210:
1209:
1176:
1170:
1169:
1159:
1127:
1118:
1117:
1107:
1075:
1066:
1065:
1055:
1046:
1037:
1036:
996:
987:
986:
938:
925:
924:
922:
902:
896:
895:
893:
892:
881:
875:
869:
863:
862:
836:
819:
816:
775:
752:
733:
704:and the extinct
666:Early Cretaceous
658:Yixian Formation
593:
577:
565:
553:
543:
530:
520:
510:
500:
490:
477:
466:
455:
444:
431:
418:
407:
397:
387:
377:
364:
351:
339:
325:
312:
299:
195:
133:
132:
104:
94:
31:
27:Temporal range:
21:
3534:
3533:
3529:
3528:
3527:
3525:
3524:
3523:
3519:Beetles of Asia
3489:
3488:
3487:
3482:
3474:
3469:
3461:
3456:
3448:
3443:
3435:
3430:
3422:
3417:
3409:
3404:
3396:
3391:
3383:
3378:
3370:
3365:
3357:
3352:
3344:
3339:
3330:
3329:
3324:
3315:
3314:
3309:
3296:
3286:
3281:
3260:
3202:
3153:
3135:(clown beetles)
3106:
3077:(betsy beetles)
2985:Scarabaeiformia
2975:
2932:
2907:
2839:(click beetles)
2815:(Texas beetles)
2791:
2762:
2758:Ptilodactylidae
2683:(cedar beetles)
2659:
2620:
2603:Trictenotomidae
2489:Mycetophagidae
2406:
2385:
2322:
2321:
2319:
2311:
2263:Phloeostichidae
2214:Lamingtoniidae
2206:Laemophloeidae
2154:Cryptophagidae
2118:
2009:
1992:Thanerocleridae
1955:Phloiophilidae
1942:Metaxina ornata
1907:Chaetosomatidae
1865:
1798:
1777:
1735:(auger beetles)
1716:Bostrichiformia
1706:
1688:
1684:Torridincolidae
1668:(skiff beetles)
1666:Hydroscaphidae
1657:
1635:
1628:Trachypachidae
1573:(tiger beetles)
1531:
1513:
1463:
1449:
1405:
1398:
1345:
1342:
1337:
1336:
1312:
1308:
1282:
1281:
1277:
1263:
1262:
1258:
1220:
1219:
1215:
1207:
1205:
1203:
1178:
1177:
1173:
1136:Current Biology
1129:
1128:
1121:
1077:
1076:
1069:
1053:
1048:
1047:
1040:
998:
997:
990:
940:
939:
928:
904:
903:
899:
890:
888:
883:
882:
878:
870:
866:
859:
847:. Vol. 1.
838:
837:
833:
828:
823:
822:
817:
813:
808:
796:
791:
790:
789:
786:
776:
767:
753:
744:
734:
722:
611:
602:
554:Reichardt, 1973
501:Reichardt, 1976
475:Torridincolidae
456:Kirejtshuk 2009
388:Reichardt, 1973
290:
270:
226:
224:Torridincolidae
222:
218:
214:
194:
127:
117:Torridincolidae
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
3532:
3530:
3522:
3521:
3516:
3511:
3506:
3501:
3491:
3490:
3484:
3483:
3481:
3480:
3467:
3454:
3441:
3428:
3415:
3402:
3389:
3380:Fauna Europaea
3376:
3363:
3350:
3337:
3322:
3306:
3304:
3298:
3297:
3292:
3283:
3282:
3277:
3274:
3273:
3270:
3269:
3266:
3265:
3262:
3261:
3259:
3258:
3255:(rove beetles)
3253:Staphylinidae
3250:
3242:
3234:
3226:
3221:
3212:
3210:
3208:Staphylinoidea
3204:
3203:
3201:
3200:
3195:
3190:Hydrophilidae
3187:
3182:
3177:
3169:
3163:
3161:
3159:Hydrophiloidea
3155:
3154:
3152:
3151:
3146:
3138:
3129:
3127:
3118:
3112:
3111:
3108:
3107:
3105:
3104:
3101:(hide beetles)
3096:
3088:
3085:(rain beetles)
3080:
3072:
3064:
3061:(stag beetles)
3056:
3048:
3040:
3032:
3024:
3016:
3014:Bolboceratidae
3011:
2998:
2996:
2987:
2981:
2980:
2977:
2976:
2974:
2973:
2968:
2960:
2948:
2942:
2940:
2934:
2933:
2931:
2930:
2921:Rhinorhipidae
2917:
2915:
2913:Rhinorhipoidea
2909:
2908:
2906:
2905:
2897:
2892:
2887:
2879:
2871:
2863:
2855:
2850:
2842:
2834:
2826:
2818:
2810:
2801:
2799:
2793:
2792:
2790:
2789:
2781:
2772:
2770:
2764:
2763:
2761:
2760:
2755:
2747:
2739:
2731:
2726:Heteroceridae
2723:
2715:
2707:
2699:
2694:
2689:Chelonariidae
2686:
2681:Callirhipidae
2678:
2675:(pill beetles)
2669:
2667:
2661:
2660:
2658:
2657:
2652:
2643:
2641:
2632:
2626:
2625:
2622:
2621:
2619:
2618:
2610:
2605:
2600:
2592:
2587:Tenebrionidae
2584:
2576:
2568:
2560:
2552:
2544:
2536:
2528:
2523:
2515:
2510:
2502:
2494:
2486:
2478:
2470:
2462:
2454:
2449:
2441:
2433:
2425:
2416:
2414:
2412:Tenebrionoidea
2408:
2407:
2405:
2404:
2395:
2393:
2387:
2386:
2384:
2383:
2375:
2370:Curculionidae
2367:
2362:
2354:
2346:
2338:
2329:
2327:
2317:Curculionoidea
2313:
2312:
2310:
2309:
2304:
2296:
2288:
2280:
2278:Protocucujidae
2275:
2270:
2265:
2260:
2252:
2244:
2236:
2231:
2223:
2211:
2203:
2195:
2190:
2185:
2177:
2172:
2170:Cybocephalidae
2167:
2159:
2151:
2146:
2141:
2128:
2126:
2120:
2119:
2117:
2116:
2111:
2106:
2101:
2093:
2085:
2080:
2072:
2070:Discolomatidae
2067:
2059:
2054:Coccinellidae
2051:
2043:
2038:Bothrideridae
2035:
2030:
2025:
2019:
2017:
2015:Coccinelloidea
2011:
2010:
2008:
2007:
2002:Trogossitidae
1999:
1994:
1989:
1984:
1979:
1974:
1969:
1964:
1952:
1947:
1935:
1927:
1922:
1920:Lophocateridae
1917:
1909:
1904:
1896:
1888:
1875:
1873:
1867:
1866:
1864:
1863:
1858:
1853:
1848:
1843:
1838:
1835:(leaf beetles)
1833:Chrysomelidae
1830:
1821:
1819:
1817:Chrysomeloidea
1810:
1804:
1803:
1800:
1799:
1797:
1796:
1787:
1785:
1779:
1778:
1776:
1775:
1767:
1762:Nosodendridae
1759:
1754:Jacobsoniidae
1751:
1746:
1743:(skin beetles)
1738:
1729:
1727:
1718:
1708:
1707:
1701:
1694:
1693:
1690:
1689:
1687:
1686:
1681:
1676:
1671:
1662:
1659:
1658:
1652:
1645:
1644:
1641:
1640:
1637:
1636:
1634:
1633:
1625:
1617:
1612:Meru phyllisae
1605:
1600:
1592:
1584:
1576:
1568:
1560:
1555:
1546:
1544:
1533:
1532:
1526:
1519:
1518:
1515:
1514:
1512:
1511:
1506:
1504:Micromalthidae
1501:
1489:
1481:
1468:
1465:
1464:
1458:
1451:
1450:
1448:
1447:
1441:
1435:
1429:
1423:
1417:
1410:
1407:
1406:
1399:
1397:
1396:
1389:
1382:
1374:
1368:
1367:
1366:at Wikispecies
1355:
1341:
1340:External links
1338:
1335:
1334:
1306:
1295:(4): 329–338.
1275:
1256:
1213:
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1171:
1119:
1067:
1038:
1011:(2): 233–249.
988:
953:(1): 175–187.
926:
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857:
830:
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821:
820:
810:
809:
807:
804:
803:
802:
795:
792:
788:
787:
777:
770:
768:
764:Hydroscaphidae
754:
747:
745:
735:
728:
725:
724:
723:
721:
718:
610:
607:
601:
598:
597:
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580:
559:
558:
557:
556:
546:
536:
532:Endrödy-Younga
523:
513:
503:
493:
471:
470:
469:
458:
447:
437:
412:
411:
410:
400:
390:
380:
362:Hydroscaphidae
349:Sphaeriusoidea
345:
344:
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342:
331:
289:
286:
269:
266:
260:, and feed on
233:
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212:Hydroscaphidae
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38:
33:
32:
30:Carnian–Recent
26:
15:
13:
10:
9:
6:
4:
3:
2:
3531:
3520:
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3515:
3512:
3510:
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3390:
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3312:
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3299:
3295:
3290:
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3275:
3257:
3251:
3249:
3243:
3241:
3235:
3233:
3227:
3225:
3222:
3220:
3214:
3213:
3211:
3209:
3205:
3199:
3196:
3194:
3188:
3186:
3183:
3181:
3178:
3176:
3170:
3168:
3165:
3164:
3162:
3160:
3156:
3150:
3147:
3145:
3141:Sphaeritidae
3139:
3137:
3131:
3130:
3128:
3126:
3122:
3119:
3117:
3113:
3103:
3097:
3095:
3091:Scarabaeidae
3089:
3087:
3081:
3079:
3073:
3071:
3065:
3063:
3057:
3055:
3049:
3047:
3041:
3039:
3033:
3031:
3029:(dor beetles)
3025:
3023:
3017:
3015:
3012:
3010:
3007:
3000:
2999:
2997:
2995:
2994:Scarabaeoidea
2991:
2988:
2986:
2982:
2972:
2969:
2967:
2961:
2959:
2956:
2949:
2947:
2944:
2943:
2941:
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2935:
2929:
2926:
2919:
2918:
2916:
2914:
2910:
2904:
2898:
2896:
2893:
2891:
2888:
2886:
2880:
2878:
2872:
2870:
2864:
2862:
2856:
2854:
2851:
2849:
2843:
2841:
2835:
2833:
2829:Cerophytidae
2827:
2825:
2819:
2817:
2811:
2809:
2803:
2802:
2800:
2798:
2794:
2788:
2784:Rhipiceridae
2782:
2780:
2774:
2773:
2771:
2769:
2765:
2759:
2756:
2754:
2748:
2746:
2740:
2738:
2732:
2730:
2724:
2722:
2718:Eulichadidae
2716:
2714:
2708:
2706:
2700:
2698:
2697:Cneoglossidae
2695:
2693:
2687:
2685:
2679:
2677:
2671:
2670:
2668:
2666:
2662:
2656:
2655:Schizopodidae
2653:
2651:
2645:
2644:
2642:
2640:
2636:
2633:
2631:
2630:Elateriformia
2627:
2617:
2611:
2609:
2606:
2604:
2601:
2599:
2595:Tetratomidae
2593:
2591:
2585:
2583:
2577:
2575:
2569:
2567:
2561:
2559:
2553:
2551:
2547:Ripiphoridae
2545:
2543:
2537:
2535:
2531:Pyrochroidae
2529:
2527:
2526:Pterogeniidae
2524:
2522:
2516:
2514:
2511:
2509:
2503:
2501:
2495:
2493:
2487:
2485:
2479:
2477:
2471:
2469:
2465:Melandryidae
2463:
2461:
2455:
2453:
2452:Chalcodryidae
2450:
2448:
2442:
2440:
2434:
2432:
2426:
2424:
2418:
2417:
2415:
2413:
2409:
2403:
2397:
2396:
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2378:Nemonychidae
2376:
2374:
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2273:Propalticidae
2271:
2269:
2268:Priasilphidae
2266:
2264:
2261:
2259:
2253:
2251:
2247:Passandridae
2245:
2243:
2241:(sap beetles)
2237:
2235:
2232:
2230:
2224:
2222:
2219:
2212:
2210:
2204:
2202:
2196:
2194:
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2189:
2186:
2184:
2178:
2176:
2173:
2171:
2168:
2166:
2160:
2158:
2152:
2150:
2149:Cavognathidae
2147:
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2137:
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2127:
2125:
2121:
2115:
2112:
2110:
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2100:
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2083:Eupsilobiidae
2081:
2079:
2075:Endomychidae
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2062:Corylophidae
2060:
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2018:
2016:
2012:
2006:
2000:
1998:
1995:
1993:
1990:
1988:
1985:
1983:
1980:
1978:
1977:Protopeltidae
1975:
1973:
1972:Prionoceridae
1970:
1968:
1965:
1963:
1960:
1953:
1951:
1948:
1946:
1943:
1936:
1934:
1928:
1926:
1925:Mauroniscidae
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1918:
1916:
1910:
1908:
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1903:
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1846:Megalopodidae
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1825:Cerambycidae
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1822:
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1818:
1814:
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1791:Derodontidae
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1783:Derodontoidea
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1768:
1766:
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1749:Endecatomidae
1747:
1745:
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1733:Bostrichidae
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1730:
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1725:Bostrichoidea
1722:
1719:
1717:
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1705:
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761:
759:
755:Specimens of
751:
746:
742:
740:
732:
727:
719:
717:
715:
714:
713:Crowsonaerius
709:
708:
703:
702:
697:
696:
691:
690:
685:
684:
679:
678:Burmese amber
675:
671:
667:
663:
659:
655:
654:
649:
645:
641:
637:
636:
631:
627:
623:
621:
616:
609:Fossil record
608:
606:
599:
592:
591:
586:
585:
584:
576:
575:
570:
569:
568:
564:
552:
551:
547:
544:Steffan, 1964
542:
541:
537:
533:
529:
528:
524:
521:Steffan, 1973
519:
518:
514:
509:
508:
504:
499:
498:
494:
489:
488:
484:
483:
480:
476:
472:
465:
464:
463:Crowsonaerius
459:
454:
453:
448:
443:
442:
438:
434:
430:
429:
425:
424:
421:
417:
416:Sphaeriusidae
413:
406:
405:
401:
396:
395:
391:
386:
385:
381:
378:LeConte, 1874
376:
375:
371:
370:
367:
363:
359:
358:
357:
354:
350:
338:
337:
332:
328:
324:
323:
319:
318:
315:
311:
307:
306:
305:
302:
298:
293:
287:
285:
283:
278:
276:
267:
265:
263:
259:
255:
251:
247:
243:
239:
231:
230:
225:
221:
220:Sphaeriusidae
217:
213:
208:
203:
198:
193:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
153:
150:
147:
146:
143:
140:
137:
136:
131:
126:
122:
118:
114:
112:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
3301:
3180:Helophoridae
3172:Georissidae
3167:Epimetopidae
3083:Pleocomidae
3067:Ochodaeidae
3051:Hybosoridae
3035:Glaphyridae
3027:Geotrupidae
3005:
3002:Belohinidae
2963:Eucinetidae
2954:
2951:Decliniidae
2924:
2882:Phengodidae
2821:Cantharidae
2776:Dascillidae
2768:Dascilloidea
2750:Psephenidae
2742:Lutrochidae
2734:Limnichidae
2647:Buprestidae
2639:Buprestoidea
2579:Synchroidae
2563:Scraptiidae
2555:Salpingidae
2518:Prostomidae
2505:Oedemeridae
2481:Mordellidae
2399:Lymexylidae
2391:Lymexyloidea
2341:Attelabidae
2333:Anthribidae
2291:Smicripidae
2255:Phalacridae
2239:Nitidulidae
2234:Myraboliidae
2226:Monotomidae
2217:
2198:Kateretidae
2175:Cyclaxyridae
2135:
2132:Agapythidae
2096:Latridiidae
2046:Cerylonidae
2033:Anamorphidae
1967:Phycosecidae
1958:
1941:
1938:Metaxinidae
1891:Biphyllidae
1882:
1851:Orsodacnidae
1808:Cucujiformia
1741:Dermestidae
1654:
1611:
1550:Amphizoidae
1495:
1475:
1461:Archostemata
1445:Holometabola
1443:Superorder:
1437:Infraclass:
1350:
1322:
1318:
1314:
1309:
1292:
1288:
1278:
1269:
1265:
1259:
1229:(1): 116–9.
1226:
1222:
1216:
1206:, retrieved
1184:
1174:
1139:
1135:
1087:
1083:
1061:
1057:
1008:
1004:
950:
946:
913:(6): 71–74.
910:
900:
889:. Retrieved
879:
874:, p. 43
867:
844:
840:
834:
814:
779:
778:Specimen of
756:
737:
736:Specimen of
711:
705:
699:
693:
687:
681:
651:
633:
618:
612:
603:
600:Distribution
588:
582:
572:
560:
548:
540:Torridincola
538:
525:
517:Incoltorrida
515:
505:
495:
485:
461:
450:
439:
426:
402:
392:
382:
372:
347:Superfamily
346:
334:
320:
297:Lepiceroidea
295:Superfamily
294:
291:
279:
271:
246:Archostemata
237:
236:
210:
191:
109:
18:
3419:iNaturalist
3326:Wikispecies
3224:Hydraenidae
3198:Spercheidae
3185:Hydrochidae
3149:Synteliidae
3133:Histeridae
3125:Histeroidea
3075:Passalidae
3043:Glaresidae
2900:Throscidae
2860:(fireflies)
2858:Lampyridae
2845:Eucnemidae
2837:Elateridae
2797:Elateroidea
2613:Zopheridae
2497:Mycteridae
2428:Anthicidae
2299:Sphindidae
2283:Silvanidae
2193:Hobartiidae
2180:Erotylidae
2109:Mycetaeidae
2104:Murmidiidae
2088:Euxestidae
1982:Rentoniidae
1856:Oxypeltidae
1841:Disteniidae
1674:Lepiceridae
1603:Hygrobiidae
1595:Haliplidae
1579:Dytiscidae
1558:Aspidytidae
1347:"Myxophaga"
1315:Lepiceratus
885:"Myxophaga"
758:Hydroscapha
707:Burmasporum
689:Lepiceratus
662:Jehol Biota
653:Hydroscapha
648:rove beetle
635:Leehermania
590:Leehermania
563:Triamyxidae
452:Burmasporum
374:Hydroscapha
336:Lepiceratus
327:Motschulsky
310:Lepiceridae
268:Description
258:semiaquatic
229:Triamyxidae
216:Lepiceridae
3493:Categories
3245:Silphidae
3237:Ptiliidae
3229:Leiodidae
3216:Agyrtidae
3059:Lucanidae
2938:Scirtoidea
2874:Omethidae
2853:Jurasaidae
2702:Dryopidae
2673:Byrrhidae
2665:Byrrhoidea
2357:Brentidae
2162:Cucujidae
2144:Boganiidae
2124:Cucujoidea
1997:Thymalidae
1987:Rhadalidae
1930:Melyridae
1899:Byturidae
1861:Vesperidae
1770:Ptiniidae
1620:Noteridae
1587:Gyrinidae
1563:Carabidae
1492:Jurodidae
1484:Cupedidae
1431:Subclass:
1421:Arthropoda
1402:Coleoptera
1272:: 155–164.
1208:2021-07-26
1090:: 901573.
891:2017-05-26
826:References
783:inaequalis
701:Bezesporum
674:Cenomanian
487:Claudiella
441:Bezesporum
242:Coleoptera
188:Suborder:
182:Coleoptera
162:Arthropoda
3499:Myxophaga
3346:Myxophaga
3332:Myxophaga
3302:Myxophaga
3099:Trogidae
3093:(scarabs)
2971:Scirtidae
2946:Clambidae
2539:Pythidae
2473:Meloidae
2420:Aderidae
2323:(weevils)
2188:Helotidae
2114:Teredidae
2028:Alexiidae
1912:Cleridae
1871:Cleroidea
1704:Polyphaga
1702:Suborder
1655:Myxophaga
1653:Suborder
1608:Meruidae
1527:Suborder
1509:Ommatidae
1459:Suborder
1433:Pterygota
1413:Kingdom:
1364:Myxophaga
1251:140170420
1114:2296-6463
1064:: 99–102.
1033:254440169
1025:0307-6970
983:203407211
975:0307-6970
781:Lepicerus
739:Sphaerius
695:Sphaerius
683:Lepicerus
626:coprolite
428:Sphaerius
384:Scaphydra
322:Lepicerus
238:Myxophaga
205:Families
192:Myxophaga
148:Kingdom:
142:Eukaryota
113:kurosawai
24:Myxophaga
3354:BugGuide
3311:Wikidata
2866:Lycidae
2710:Elmidae
2608:Ulodidae
2444:Boridae
2349:Belidae
1950:Peltidae
1543:families
1529:Adephaga
1439:Neoptera
1419:Phylum:
1415:Animalia
1404:families
1166:34197727
794:See also
620:Triamyxa
574:Triamyxa
561:Family â€
527:Satonius
420:Erichson
404:Confossa
353:Erichson
288:Taxonomy
158:Phylum:
152:Animalia
138:Domain:
111:Satonius
3411:4577555
3372:2650686
3317:Q133005
2457:Ciidae
2365:Caridae
1427:Insecta
1425:Class:
1400:Extant
1231:Bibcode
1144:Bibcode
1092:Bibcode
1058:Denisia
955:Bibcode
741:minutus
720:Gallery
676:) aged
660:in the
630:Carnian
497:Delevea
479:Steffan
473:Family
414:Family
366:LeConte
360:Family
308:Family
254:aquatic
250:beetles
197:Crowson
178:Order:
172:Insecta
168:Class:
3437:112736
3424:340472
1541:Extant
1325:(13).
1249:
1199:
1164:
1112:
1031:
1023:
981:
973:
855:
760:natans
670:Aptian
640:Norian
534:, 1997
481:, 1964
435:, 1838
422:, 1845
368:, 1874
355:, 1845
329:, 1855
316:, 1936
314:Hinton
303:, 1936
301:Hinton
282:galeae
244:after
199:, 1955
3476:69191
3463:63907
3385:10998
3359:40604
1247:S2CID
1054:(PDF)
1029:S2CID
979:S2CID
806:Notes
507:Iapir
433:Waltl
262:algae
3458:NCBI
3432:ITIS
3406:GBIF
1197:ISBN
1162:PMID
1110:ISSN
1021:ISSN
971:ISSN
853:ISBN
710:and
698:and
394:Yara
256:and
35:Preęž’
3445:NBN
3367:EoL
3341:AFD
1327:doi
1297:doi
1270:108
1239:doi
1189:doi
1152:doi
1100:doi
1013:doi
963:doi
915:doi
550:Ytu
3495::
3473::
3460::
3447::
3434::
3421::
3408::
3395::
3382::
3369::
3356::
3343::
3328::
3313::
1349:.
1323:18
1321:.
1293:41
1291:.
1287:.
1268:.
1245:.
1237:.
1227:86
1225:.
1195:,
1183:,
1160:.
1150:.
1140:31
1138:.
1134:.
1122:^
1108:.
1098:.
1088:10
1086:.
1082:.
1070:^
1062:26
1060:.
1056:.
1041:^
1027:.
1019:.
1009:48
1007:.
1003:.
991:^
977:.
969:.
961:.
951:45
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851:.
843:.
642:)
264:.
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85:Pg
3008:)
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2957:)
2953:(
2927:)
2923:(
2220:)
2216:(
2138:)
2134:(
1961:)
1957:(
1944:)
1940:(
1885:)
1881:(
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1494:(
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1474:(
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1329::
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1299::
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1241::
1233::
1191::
1168:.
1154::
1146::
1116:.
1102::
1094::
1035:.
1015::
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965::
957::
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917::
894:.
861:.
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762:(
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