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2091:
146:
124:
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1133:
552:
The phylogenetic relationships of the
Machimosauridae were analyzed in a comprehensive analysis of teleosauroid relationships. The results of the analysis are shown below:
476:. The family was largely expanded in 2020 when the systematics of Teleosauroidea were re-reviewed. Members of this family generally were larger than the teleosaurids.
2135:
1250:
1043:
reaching up to 6.6 m (22 ft) long. Machimosaurins were the largest teleosauroids, with body lengths up to 7.1 m (23 ft) (
1023:
Machimosaurids grew in body size and diversified feeding strategies throughout their evolutionary history. Primitive machimosaurids, such as
2164:
1193:"Big-headed marine crocodyliforms and why we must be cautious when using extant species as body length proxies for long-extinct relatives"
1031:, were longirostrine (long-snouted) generalists with body lengths less than 5 m (16 ft) long. Machimosaurines originated in the
2044:
492:
2128:
484:
Machimosauridae is a diverse group of teleosauroids, phylogenetically defined as "the largest clade within
Teleosauroidea containing
2159:
1155:"The oldest durophagous teleosauroid (Crocodylomorpha, Thalattosuchia) from the lower Bathonian of central High Atlas, Morocco"
145:
2121:
1243:
1083:"The phylogenetics of Teleosauroidea (Crocodylomorpha, Thalattosuchia) and implications for their ecology and evolution"
597:
645:
721:
2154:
2070:
2049:
1192:
839:
815:
754:
669:
573:
132:
1236:
533:
798:
498:
1338:
502:." The members of the Machimosauridae share several unique characters among teleosauroids, which are:
1438:
1191:
Young, Mark T.; Rabi, Márton; Bell, Mark A.; Foffa, Davide; Steel, Lorna; Sachs, Sven; Peyer, Karin.
2090:
1572:
904:
863:
771:
486:
314:
140:
1689:
1565:
1214:
1122:
1104:
887:
621:
304:
1767:
1757:
1631:
1579:
1204:
1166:
1112:
1094:
703:
539:
526:
382:
324:
43:
512:
the premaxillary anterior and anterolateral margins are not sub-vertical and do not extend
2018:
1821:
1624:
1604:
1307:
1039:
and simultaneously underwent a decrease in snout length but an increase in body size with
1036:
682:
465:
372:
239:
226:
123:
2105:
1908:
1805:
1794:
1787:
1780:
1642:
1586:
1462:
1424:
1398:
1376:
1350:
1259:
1117:
784:
449:
446:
442:
334:
265:
252:
39:
2148:
1946:
1894:
1828:
1738:
1710:
1665:
1658:
1530:
1507:
1480:
1469:
1449:
460:
420:
410:
81:
2025:
2011:
2004:
1994:
1955:
1901:
1856:
1731:
1724:
1617:
1412:
1323:
1298:
548:
the cultriform process of the basisphenoid is exposed and bifurcates the pterygoids
439:
362:
213:
529:
is sub-rectangular and slightly interdigitating (most noticeably near the midline)
2098:
1880:
1843:
1814:
1651:
1593:
1523:
1496:
1489:
454:
400:
344:
56:
1137:
1919:
1887:
1867:
1289:
519:
101:
66:
17:
1218:
1153:
Stéphane Jouve; Bastien
Mennecart; Julien Douteau; Nour-Eddine Jalil (2016).
1108:
2101:
1928:
1717:
1703:
1537:
1032:
513:
469:
200:
157:
106:
50:
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1962:
1935:
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445:
crocodyliforms. The family was first identified in 2016, when fossils of
177:
96:
91:
76:
71:
61:
1745:
1171:
1154:
1132:
1099:
523:
473:
187:
111:
86:
1277:
167:
1082:
1081:
Johnson, Michela M.; Young, Mark T.; Brusatte, Stephen L. (2020).
545:
a mostly horizontal pterygoid with a distinct posterolateral angle
507:
1228:
1209:
1047:), and were durophagous hunters, meaning they fed on hard prey.
1270:
1232:
1136:
Text was copied from this source, which is available under a
2109:
1138:
Creative
Commons Attribution 4.0 International License
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1992:
1945:
1918:
1866:
1842:
1804:
1766:
1755:
1687:
1641:
1603:
1551:
1506:
1479:
1448:
1422:
1396:
1349:
1322:
542:is excluded from the orbit posteroventral margin
452:, including an indeterminate close relative of
2129:
1244:
1076:
1074:
1072:
1070:
1068:
1066:
1064:
1062:
1060:
8:
1148:
1146:
2136:
2122:
1989:
1872:
1848:
1772:
1763:
1695:
1609:
1557:
1454:
1430:
1404:
1393:
1328:
1319:
1267:
1251:
1237:
1229:
122:
31:
1208:
1170:
1116:
1098:
1056:
7:
2087:
2085:
1186:
1184:
1182:
879:
855:
831:
790:
746:
695:
688:
661:
637:
613:
589:
565:
555:
493:Plagiophthalmosuchus gracilirostris
532:no anterolateral expansion of the
25:
2089:
2066:
2065:
2048:
2043:
1131:
144:
1:
2108:. You can help Knowledge by
598:Clovesuurdameredeor stephani
2165:Prehistoric archosaur stubs
646:Seldsienean megistorhynchus
506:dorsally oriented external
2181:
2084:
1197:Palaeontologia Electronica
722:Proexochokefalos bouchardi
464:, were described from the
27:Extinct family of reptiles
2061:
2041:
1988:
1875:
1851:
1775:
1698:
1612:
1560:
1457:
1433:
1407:
1392:
1373:
1331:
1318:
1266:
901:
884:
877:
860:
853:
836:
829:
816:Yvridiosuchus boutilieri
812:
795:
788:
768:
751:
744:
717:
704:Proexochokefalos hebertii
700:
693:
686:
666:
659:
642:
635:
618:
611:
594:
587:
570:
563:
297:
292:
141:Scientific classification
139:
130:
121:
34:
840:Machimosaurus buffetauti
755:Neosteneosaurus edwardsi
670:Deslongchampsina larteti
622:Charitomenosuchus leedsi
574:Macrospondylus bollensis
438:is an extinct family of
133:Macrospondylus bollensis
534:supratemporal fenestrae
2160:Pseudosuchian families
799:Lemmysuchus obtusidens
499:Teleosaurus cadomensis
2097:This article about a
1439:Plagiophthalmosuchus
1573:Clovesuurdameredeor
905:Machimosaurus hugii
864:Machimosaurus mosae
772:Andrianavoay baroni
487:Machimosaurus hugii
315:Clovesuurdameredeor
1172:10.1111/pala.12262
1100:10.7717/peerj.9808
2117:
2116:
2079:
2078:
2057:
2056:
2039:
2038:
2035:
2034:
1984:
1983:
1980:
1979:
1976:
1975:
1972:
1971:
1838:
1837:
1690:Metriorhynchoidea
1683:
1682:
1679:
1678:
1675:
1674:
1566:Charitomenosuchus
1547:
1546:
1388:
1387:
1384:
1383:
1369:
1368:
1045:Machimosaurus rex
1029:Charitomenosuchus
1015:
1014:
1006:
1005:
997:
996:
988:
987:
979:
978:
970:
969:
961:
960:
952:
951:
943:
942:
934:
933:
925:
924:
916:
915:
888:Machimosaurus rex
733:
732:
433:
432:
305:Charitomenosuchus
288:
16:(Redirected from
2172:
2155:Thalattosuchians
2138:
2131:
2124:
2093:
2086:
2069:
2068:
2052:
2047:
1990:
1873:
1849:
1773:
1768:Metriorhynchinae
1764:
1758:Metriorhynchidae
1696:
1632:Proexochokefalos
1610:
1580:Deslongchampsina
1558:
1455:
1431:
1405:
1394:
1329:
1320:
1313:
1312:
1268:
1253:
1246:
1239:
1230:
1223:
1222:
1212:
1188:
1177:
1176:
1174:
1150:
1141:
1135:
1130:
1120:
1102:
1078:
880:
856:
832:
791:
747:
696:
689:
662:
638:
614:
590:
566:
556:
450:thalattosuchians
418:
408:
398:
390:
383:Proexochokefalos
380:
370:
360:
352:
342:
332:
325:Deslongchampsina
322:
312:
302:
282:
277:
264:
251:
149:
148:
126:
116:
53:
44:Early Cretaceous
38:Temporal range:
32:
21:
2180:
2179:
2175:
2174:
2173:
2171:
2170:
2169:
2145:
2144:
2143:
2142:
2082:
2080:
2075:
2053:
2031:
2019:Rhamphocephalus
1968:
1941:
1914:
1862:
1834:
1822:Maledictosuchus
1800:
1760:
1751:
1692:
1671:
1637:
1625:Neosteneosaurus
1605:Machimosaurinae
1599:
1553:Machimosauridae
1543:
1502:
1475:
1444:
1427:
1418:
1401:
1380:
1365:
1345:
1314:
1308:Crocodylomorpha
1273:
1272:
1262:
1257:
1227:
1226:
1190:
1189:
1180:
1152:
1151:
1144:
1080:
1079:
1058:
1053:
1041:Neosteneosaurus
1037:Middle Jurassic
1021:
1016:
1007:
998:
989:
980:
971:
962:
953:
944:
935:
926:
917:
734:
683:Machimosaurinae
560:Machimosauridae
482:
466:Middle Jurassic
443:thalattosuchian
436:Machimosauridae
373:Neosteneosaurus
354:Machimosaurinae
281:
279:Machimosauridae
275:
262:
249:
240:Crocodyliformes
227:Crocodylomorpha
143:
117:
115:
114:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
48:
47:
36:
35:Machimosauridae
28:
23:
22:
15:
12:
11:
5:
2178:
2176:
2168:
2167:
2162:
2157:
2147:
2146:
2141:
2140:
2133:
2126:
2118:
2115:
2114:
2094:
2077:
2076:
2074:
2073:
2062:
2059:
2058:
2055:
2054:
2042:
2040:
2037:
2036:
2033:
2032:
2030:
2029:
2022:
2015:
2008:
2000:
1998:
1986:
1985:
1982:
1981:
1978:
1977:
1974:
1973:
1970:
1969:
1967:
1966:
1959:
1951:
1949:
1943:
1942:
1940:
1939:
1932:
1924:
1922:
1916:
1915:
1913:
1912:
1909:Tyrannoneustes
1905:
1898:
1891:
1884:
1876:
1870:
1864:
1863:
1861:
1860:
1852:
1846:
1840:
1839:
1836:
1835:
1833:
1832:
1825:
1818:
1810:
1808:
1806:Rhacheosaurini
1802:
1801:
1799:
1798:
1795:Thalattosuchus
1791:
1788:Metriorhynchus
1784:
1781:Gracilineustes
1776:
1770:
1761:
1756:
1753:
1752:
1750:
1749:
1742:
1735:
1728:
1721:
1714:
1707:
1699:
1693:
1688:
1685:
1684:
1681:
1680:
1677:
1676:
1673:
1672:
1670:
1669:
1662:
1655:
1647:
1645:
1643:Machimosaurini
1639:
1638:
1636:
1635:
1628:
1621:
1613:
1607:
1601:
1600:
1598:
1597:
1590:
1587:Macrospondylus
1583:
1576:
1569:
1561:
1555:
1549:
1548:
1545:
1544:
1542:
1541:
1534:
1527:
1520:
1512:
1510:
1504:
1503:
1501:
1500:
1493:
1485:
1483:
1477:
1476:
1474:
1473:
1466:
1463:Indosinosuchus
1458:
1452:
1446:
1445:
1443:
1442:
1434:
1428:
1425:Teleosauroidea
1423:
1420:
1419:
1417:
1416:
1408:
1402:
1399:Thalattosuchia
1397:
1390:
1389:
1386:
1385:
1382:
1381:
1377:Thalattosuchia
1374:
1371:
1370:
1367:
1366:
1364:
1363:
1362:
1361:
1355:
1353:
1351:Thalattosuchia
1347:
1346:
1344:
1343:
1342:
1341:
1332:
1326:
1316:
1315:
1311:
1310:
1301:
1292:
1286:
1280:
1271:
1264:
1263:
1260:Thalattosuchia
1258:
1256:
1255:
1248:
1241:
1233:
1225:
1224:
1178:
1165:(6): 863–876.
1142:
1055:
1054:
1052:
1049:
1025:Macrospondylus
1020:
1017:
1013:
1012:
1009:
1008:
1004:
1003:
1000:
999:
995:
994:
991:
990:
986:
985:
982:
981:
977:
976:
973:
972:
968:
967:
964:
963:
959:
958:
955:
954:
950:
949:
946:
945:
941:
940:
937:
936:
932:
931:
928:
927:
923:
922:
919:
918:
914:
913:
910:
909:
900:
897:
896:
893:
892:
883:
878:
876:
873:
872:
869:
868:
859:
854:
852:
849:
848:
845:
844:
835:
830:
828:
825:
824:
821:
820:
811:
808:
807:
804:
803:
794:
789:
787:
785:Machimosaurini
781:
780:
777:
776:
767:
764:
763:
760:
759:
750:
745:
743:
740:
739:
736:
735:
731:
730:
727:
726:
716:
713:
712:
709:
708:
699:
694:
692:
687:
685:
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678:
675:
674:
665:
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658:
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654:
651:
650:
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634:
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588:
586:
583:
582:
579:
578:
569:
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562:
554:
550:
549:
546:
543:
536:
530:
516:
510:
481:
480:Classification
478:
431:
430:
429:
428:
427:
426:
425:
424:
414:
404:
392:Machimosaurini
386:
376:
366:
348:
338:
335:Macrospondylus
328:
318:
308:
295:
294:
290:
289:
273:
269:
268:
266:Teleosauroidea
260:
256:
255:
253:Thalattosuchia
247:
243:
242:
237:
230:
229:
224:
217:
216:
211:
204:
203:
198:
191:
190:
185:
181:
180:
175:
171:
170:
165:
161:
160:
155:
151:
150:
137:
136:
128:
127:
119:
118:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
54:
40:Early Jurassic
37:
26:
24:
18:Machimosaurini
14:
13:
10:
9:
6:
4:
3:
2:
2177:
2166:
2163:
2161:
2158:
2156:
2153:
2152:
2150:
2139:
2134:
2132:
2127:
2125:
2120:
2119:
2113:
2111:
2107:
2103:
2100:
2095:
2092:
2088:
2083:
2072:
2064:
2063:
2060:
2051:
2046:
2028:
2027:
2023:
2021:
2020:
2016:
2014:
2013:
2009:
2007:
2006:
2002:
2001:
1999:
1997:
1996:
1991:
1987:
1965:
1964:
1960:
1958:
1957:
1953:
1952:
1950:
1948:
1947:Plesiosuchina
1944:
1938:
1937:
1933:
1931:
1930:
1926:
1925:
1923:
1921:
1917:
1911:
1910:
1906:
1904:
1903:
1899:
1897:
1896:
1895:Purranisaurus
1892:
1890:
1889:
1885:
1883:
1882:
1878:
1877:
1874:
1871:
1869:
1865:
1859:
1858:
1854:
1853:
1850:
1847:
1845:
1841:
1831:
1830:
1829:Rhacheosaurus
1826:
1824:
1823:
1819:
1817:
1816:
1812:
1811:
1809:
1807:
1803:
1797:
1796:
1792:
1790:
1789:
1785:
1783:
1782:
1778:
1777:
1774:
1771:
1769:
1765:
1762:
1759:
1754:
1748:
1747:
1743:
1741:
1740:
1739:Teleidosaurus
1736:
1734:
1733:
1729:
1727:
1726:
1722:
1720:
1719:
1715:
1713:
1712:
1711:Magyarosuchus
1708:
1706:
1705:
1701:
1700:
1697:
1694:
1691:
1686:
1668:
1667:
1666:Yvridiosuchus
1663:
1661:
1660:
1659:Machimosaurus
1656:
1654:
1653:
1649:
1648:
1646:
1644:
1640:
1634:
1633:
1629:
1627:
1626:
1622:
1620:
1619:
1615:
1614:
1611:
1608:
1606:
1602:
1596:
1595:
1591:
1589:
1588:
1584:
1582:
1581:
1577:
1575:
1574:
1570:
1568:
1567:
1563:
1562:
1559:
1556:
1554:
1550:
1540:
1539:
1535:
1533:
1532:
1531:Mycterosuchus
1528:
1526:
1525:
1521:
1519:
1518:
1514:
1513:
1511:
1509:
1508:Aeolodontinae
1505:
1499:
1498:
1494:
1492:
1491:
1487:
1486:
1484:
1482:
1481:Teleosaurinae
1478:
1472:
1471:
1470:Mystriosaurus
1467:
1465:
1464:
1460:
1459:
1456:
1453:
1451:
1450:Teleosauridae
1447:
1441:
1440:
1436:
1435:
1432:
1429:
1426:
1421:
1415:
1414:
1410:
1409:
1406:
1403:
1400:
1395:
1391:
1379:
1378:
1372:
1359:
1358:
1357:
1356:
1354:
1352:
1348:
1340:
1336:
1335:
1334:
1333:
1330:
1327:
1325:
1321:
1317:
1309:
1305:
1302:
1300:
1296:
1293:
1291:
1287:
1285:
1281:
1279:
1275:
1274:
1269:
1265:
1261:
1254:
1249:
1247:
1242:
1240:
1235:
1234:
1231:
1220:
1216:
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461:Machimosaurus
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423:
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415:
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49:183–130
45:
41:
33:
30:
19:
2110:expanding it
2096:
2081:
2026:Steneosaurus
2024:
2017:
2012:Neustosaurus
2010:
2005:Enaliosuchus
2003:
1995:Nomina dubia
1993:
1961:
1956:Plesiosuchus
1954:
1934:
1927:
1907:
1902:Torvoneustes
1900:
1893:
1886:
1879:
1857:Neptunidraco
1855:
1827:
1820:
1813:
1793:
1786:
1779:
1744:
1737:
1732:Pelagosaurus
1730:
1725:Peipehsuchus
1723:
1716:
1709:
1702:
1664:
1657:
1650:
1630:
1623:
1618:Andrianavoay
1616:
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1515:
1495:
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1437:
1413:Turnersuchus
1411:
1375:
1339:Pseudosuchia
1324:Pseudosuchia
1303:
1299:Pseudosuchia
1294:
1210:10.26879/648
1200:
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363:Andrianavoay
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233:
220:
214:Pseudosuchia
207:
194:
131:
29:
2099:prehistoric
1881:Aggiosaurus
1844:Geosaurinae
1815:Cricosaurus
1652:Lemmysuchus
1594:Seldsienean
1524:Bathysuchus
1497:Teleosaurus
1490:Platysuchus
1203:(3): 1–14.
540:postorbital
455:Lemmysuchus
401:Lemmysuchus
345:Seldsienean
201:Archosauria
2149:Categories
1920:Geosaurina
1888:Dakosaurus
1868:Geosaurini
1360:see below↓
1290:Sauropsida
1051:References
520:premaxilla
490:, but not
293:Subgroups
246:Suborder:
2102:archosaur
1929:Geosaurus
1718:Opisuchus
1704:Eoneustes
1538:Sericodon
1276:Kingdom:
1219:1094-8074
1109:2167-8359
1093:: e9808.
1033:Bathonian
1019:Evolution
514:ventrally
470:Bathonian
164:Kingdom:
158:Eukaryota
2071:Category
1963:Suchodus
1936:Ieldraan
1517:Aeolodon
1284:Chordata
1282:Phylum:
1278:Animalia
1127:33083104
272:Family:
188:Reptilia
178:Chordata
174:Phylum:
168:Animalia
154:Domain:
1746:Zoneait
1288:Class:
1118:7548081
524:maxilla
474:Morocco
184:Class:
1217:
1125:
1115:
1107:
527:suture
287:, 2016
285:et al.
283:Jouve
2104:is a
1304:Clade
1295:Clade
1087:PeerJ
508:nares
472:) of
234:Clade
221:Clade
208:Clade
195:Clade
2106:stub
1337:see
1215:ISSN
1123:PMID
1105:ISSN
1027:and
719:cf.
538:the
518:the
496:and
458:and
57:PreꞒ
1205:doi
1167:doi
1113:PMC
1095:doi
2151::
1306::
1297::
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1201:19
1199:.
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1181:^
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1059:^
236::
223::
210::
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107:Pg
51:Ma
46:,
2137:e
2130:t
2123:v
2112:.
1252:e
1245:t
1238:v
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522:-
468:(
417:†
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263:†
250:†
112:N
102:K
97:J
92:T
87:P
82:C
77:D
72:S
67:O
62:Ꞓ
42:-
20:)
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