601:
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147:
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1339:
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129:
2075:; Barrett, Paul M.; Coria, Rodolfo A.; Le Loueff, Jean; Xu Xing; Zhao Xijin; Sahni, Ashok; Gomani, Elizabeth M.P.; Noto, Christopher N. (2004). "Dinosaur distribution". In Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (eds.).
1533:
was recovered in a polytomy with halszkaraptorines, unenlagiines and microraptorines outside the subfamily. However, other more recent analyses still support the
Halszkaraptorine placement of
526:
as the result of a size squeeze. Like birds, troodontids and dromaeosaurids were not small throughout their evolutionary history, and showed size increases among several different lineages.
522:
lineages, suggest that small size was not an innovation of early birds, but a common trait of early paravians; small size would have preceded flight and would not have been a special avian
530:
has a combination of characteristics found among basal troodontids and birds, but lacks some that are present in more derived dromaeosaurids. A study in 2017 found that
3159:
1819:
Cau, A.; Beyrand, V.; Voeten, D. F. A. E.; Fernandez, V.; Tafforeau, P.; Stein, K.; Barsbold, R.; Tsogtbaatar, K.; Currie, P. J.; Godefroit, P. (2017).
551:
below is based on the phylogenetic analysis conducted in 2017 by Cau et al. using updated data from the
Theropod Working Group in their description of
2171:
1945:"Early Cretaceous troodontine troodontid (Dinosauria: Theropoda) from the Ohyamashimo Formation of Japan reveals the early evolution of Troodontinae"
3083:
3174:
2125:
2092:
2056:
1792:
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was a small dromaeosaurid, measuring 50–70 cm (20–28 in) long and weighing 400–700 g (14–25 oz). The holotype specimen,
1876:"New Dromaeosaurid Dinosaur (Theropoda, Dromaeosauridae) from New Mexico and Biodiversity of Dromaeosaurids at the end of the Cretaceous"
3179:
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1707:
1698:
are known to be strong swimmers, this suggests that
Mahakala may too have been a strong swimmer like the other members of its family.
600:
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371:. Despite its late appearance, it is among the most basal dromaeosaurids. Its small size, and the small size of other basal
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and derived dromaeosaurids, but unlike basal troodontids and other dromaeosaurids, the middle (third)
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Jerzykiewicz, Tom (1997). "Djadokhta
Formation". In Currie, Phillip J.; Padian, Kevin (eds.).
2021:
1982:
1964:
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1907:
1848:
1798:
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1821:"Synchrotron scanning reveals amphibious ecomorphology in a new clade of bird-like dinosaurs"
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2013:
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to be the most basal known dromaeosaurid. Their results, along with the small size of other
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version. It had a typical dromaeosaurid form of the second toe, with an expanded claw.
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HTML version of "A Basal
Dromaeosaurid and Size Evolution Preceding Avian Flight"
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The paleoenvironment of the
Djadokhta Formation is interpreted as having a
921:
1732:
Turner, A.H.; Pol, D.; Clarke, J.A.; Erickson, G.M.; Norell, M.A. (2007).
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Kubota, Katsuhiro; Kobayashi, Yoshitsugu; Ikeda, Tadahiro (2024-07-25).
1844:
470:, it was close to adulthood. This genus can be distinguished from other
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2001:
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84:
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Jasinski, Steven E.; Sullivan, Robert M.; Dodson, Peter (2020-03-26).
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performed by Turner and colleagues, who described the specimen, found
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was not compressed, suggesting that the uncompressed version was the
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2116:(2nd ed.). Berkeley: University of California Press. pp.
2083:(2nd ed.). Berkeley: University of California Press. pp.
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1569:, and moisture was seasonal. Animals present included terrestrial
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1734:"A Basal Dromaeosaurid and Size Evolution Preceding Avian Flight"
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100/1033, consists of a partial skeleton including skull bones,
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375:, suggests that small size appeared before flight capability in
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are the most diverse dinosaurs, including fellow dromaeosaurid
363:, and its skeleton shows features that are also found in early
452:. Although this individual was small, comparable in size to
2002:"A Unified Framework for Predatory Dinosaur Macroevolution"
474:(dromaeosaurids, troodontids, and birds) by details of the
398:
Artist's reconstruction of
Mahakala as a halszkaraptorine
3127:
426:
Size compared to a human, reconstructed as a generic
283:
3045:
2859:
2773:
2738:
2676:
2604:
2493:
2451:
2388:
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2108:Weishampel, Davi, ed. (2004). "Dromaeosauridae".
351:. It is based on a partial skeleton found in the
2006:Bollettino della Società Paleontologica Italiana
2165:
8:
136:Diagram of known elements from the holotype
1787:. Princeton University Press. p. 150.
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1565:and was sometimes affected by dust and
336:-age (about 80 million years ago)
2047:. San Diego: Academic Press. pp.
1784:The Princeton Field Guide to Dinosaurs
1593:were not present. The majority of the
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7:
1653:; other dinosaurs present included
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1708:Timeline of dromaeosaurid research
444:, limb bones, and portions of the
383:, one of eight protector deities (
25:
3160:Late Cretaceous dinosaurs of Asia
1597:was small to medium-sized. Small
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411:2009 skeletal reconstruction of
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1687:would have been a small active
534:was a member of the enigmatic,
1585:, and a variety of dinosaurs;
1:
3175:Fossil taxa described in 2007
1683:. Like other dromaeosaurids,
1541:Paleoecology and paleobiology
486:, and tail vertebrae. Like
27:Extinct species of dinosaur
3196:
1961:10.1038/s41598-024-66815-2
1903:10.1038/s41598-020-61480-7
3180:Taxa named by Mark Norell
3006:
2974:
2685:
2399:
2384:
2365:
2315:
2288:
2261:
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2187:
2043:Encyclopedia of Dinosaurs
1781:Paul, Gregory S. (2016).
1561:landscape was drained by
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379:. The genus is named for
282:
275:
142:Scientific classification
140:
135:
126:
34:
315:[mɑːhɑːˈkɑːˈlɑː]
1759:10.1126/science.1144066
1557:settings. The semiarid
430:
419:
399:
3110:Paleobiology Database
779:Unenlagia comahuensis
518:found at the base of
508:phylogenetic analysis
425:
410:
397:
2073:Weishampel, David B.
2018:10.4435/BSPI.2024.08
2000:Cau, Andrea (2024).
1563:intermittent streams
318:from Sanskrit) is a
2740:Saurornitholestinae
1894:2020NatSR..10.5105J
1845:10.1038/nature24679
1837:2017Natur.552..395C
1750:2007Sci...317.1378T
1744:(5843): 1378–1381.
796:Unenlagia paynemili
341:Djadokhta Formation
18:Mahakala (dinosaur)
1949:Scientific Reports
1881:Scientific Reports
431:
420:
400:
373:deinonychosaurians
288:Mahakala omnogovae
3170:Halszkaraptorines
3125:
3124:
3097:Open Tree of Life
3039:Taxon identifiers
3030:
3029:
3002:
3001:
2972:
2971:
2968:
2967:
2937:Saurornitholestes
2828:Saurornitholestes
2764:Saurornitholestes
2453:Halszkaraptorinae
2380:
2379:
2376:
2375:
2361:
2360:
2357:
2356:
2353:
2352:
2127:978-0-520-24209-8
2094:978-0-520-24209-8
2058:978-0-12-226810-6
1831:(7685): 395–399.
1794:978-1-78684-190-2
1696:Halszkaraptorinae
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1031:
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815:
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806:
672:
671:
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662:
582:Halszkaraptorinae
542:Halszkaraptorinae
302:
301:
271:
244:Halszkaraptorinae
16:(Redirected from
3187:
3142:
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3133:
3118:
3117:
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3104:
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3091:
3079:
3078:
3066:
3065:
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3020:
3019:
2997:
2991:
2986:
2980:
2861:Velociraptorinae
2807:Dromaeosauroides
2683:
2667:Zhongjianosaurus
2645:Sinornithosaurus
2397:
2386:
2313:
2286:
2259:
2250:
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2174:
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1668:, the hadrosaur
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1326:
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1229:Velociraptorinae
1191:
1169:
1154:
1140:
1125:
1115:
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1070:
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1005:
998:Sinornithosaurus
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389:Tibetan Buddhism
338:Upper Cretaceous
324:halszkaraptorine
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40:Temporal range:
32:
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3165:Djadochta fauna
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2139:External links
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2112:The Dinosauria
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2079:The Dinosauria
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1393:
1389:
1388:
1385:
1384:
1375:
1372:
1371:
1368:
1367:
1358:
1353:
1351:
1348:
1347:
1344:
1343:
1329:
1324:
1322:
1319:
1318:
1315:
1314:
1310:
1309:
1306:
1305:
1296:
1293:
1292:
1289:
1288:
1274:
1269:
1267:
1262:
1260:
1257:
1256:
1253:
1252:
1238:
1233:
1231:
1225:
1224:
1221:
1220:
1216:
1215:
1212:
1211:
1207:
1206:
1203:
1202:
1198:
1197:
1194:
1193:
1179:
1176:
1175:
1172:
1171:
1157:
1152:
1150:
1147:
1146:
1143:
1142:
1128:
1123:
1121:
1113:
1111:
1106:
1104:
1101:
1100:
1097:
1096:
1087:
1082:
1080:
1077:
1076:
1073:
1072:
1058:
1053:
1051:
1043:
1041:
1038:
1037:
1034:
1033:
1029:
1028:
1025:
1024:
1015:
1012:
1011:
1008:
1007:
993:
990:
989:
986:
985:
971:
968:
967:
964:
963:
954:
951:
950:
947:
946:
934:
931:
930:
927:
926:
912:
907:
905:
897:
895:
890:
888:
885:
884:
881:
880:
866:
861:
859:
856:
855:
852:
851:
842:
837:
835:
832:
831:
828:
827:
823:
822:
819:
818:
814:
813:
810:
809:
805:
804:
801:
800:
791:
788:
787:
784:
783:
774:
771:
770:
767:
766:
752:
749:
748:
745:
744:
735:
730:
728:
725:
724:
721:
720:
706:
701:
699:
691:
689:
684:
682:
679:
678:
675:
674:
670:
669:
666:
665:
661:
660:
657:
656:
642:
639:
638:
635:
634:
620:
615:
613:
610:
609:
606:
605:
591:
586:
584:
576:
574:
569:
567:
559:
503:
502:Classification
500:
428:deinonychosaur
417:deinonychosaur
404:
401:
300:
299:
292:
280:
279:
273:
272:
251:
247:
246:
238:
234:
233:
225:
221:
220:
215:
208:
207:
202:
195:
194:
189:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
138:
137:
133:
132:
124:
123:
115:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
52:
39:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
3192:
3181:
3178:
3176:
3173:
3171:
3168:
3166:
3163:
3161:
3158:
3157:
3155:
3145:
3135:
3131:
3116:
3111:
3107:
3103:
3098:
3094:
3090:
3085:
3081:
3077:
3072:
3068:
3063:
3057:
3053:
3052:
3050:
3048:
3044:
3040:
3035:
3023:
3015:
3013:
3009:
3008:
3005:
2996:
2992:
2990:
2985:
2979:
2961:
2960:
2956:
2954:
2953:
2949:
2947:
2946:
2942:
2939:
2938:
2934:
2931:
2930:
2926:
2923:
2922:
2918:
2916:
2915:
2911:
2909:
2908:
2904:
2902:
2901:
2900:Kansaignathus
2897:
2894:
2893:
2889:
2886:
2885:
2881:
2879:
2878:
2874:
2872:
2871:
2867:
2866:
2864:
2862:
2858:
2852:
2851:
2847:
2845:
2844:
2840:
2838:
2837:
2833:
2830:
2829:
2825:
2823:
2822:
2818:
2816:
2815:
2814:Dromaeosaurus
2811:
2809:
2808:
2804:
2801:
2800:
2796:
2793:
2792:
2788:
2786:
2785:
2781:
2780:
2778:
2776:
2772:
2766:
2765:
2761:
2758:
2757:
2753:
2751:
2750:
2746:
2745:
2743:
2741:
2737:
2730:
2729:
2725:
2723:
2722:
2718:
2715:
2714:
2710:
2708:
2707:
2706:Dineobellator
2703:
2701:
2700:
2696:
2693:
2692:
2688:
2687:
2684:
2681:
2679:
2675:
2669:
2668:
2664:
2662:
2661:
2657:
2654:
2653:
2649:
2647:
2646:
2642:
2640:
2639:
2635:
2632:
2631:
2630:Hesperonychus
2627:
2625:
2624:
2623:Graciliraptor
2620:
2618:
2617:
2616:Changyuraptor
2613:
2612:
2610:
2607:
2606:Microraptoria
2603:
2597:
2596:
2592:
2589:
2588:
2584:
2581:
2580:
2576:
2574:
2573:
2569:
2566:
2565:
2561:
2558:
2557:
2553:
2551:
2550:
2546:
2543:
2542:
2541:Ornithodesmus
2538:
2536:
2535:
2534:Neuquenraptor
2531:
2529:
2528:
2524:
2521:
2520:
2516:
2514:
2513:
2509:
2507:
2506:
2502:
2501:
2499:
2496:
2492:
2486:
2485:
2481:
2479:
2478:
2474:
2472:
2471:
2467:
2465:
2464:
2463:Halszkaraptor
2460:
2459:
2457:
2454:
2450:
2444:
2443:
2439:
2436:
2435:
2431:
2429:
2428:
2424:
2421:
2420:
2416:
2414:
2413:
2409:
2407:
2406:
2402:
2401:
2398:
2395:
2392:
2387:
2383:
2371:
2370:
2364:
2343:
2342:
2341:
2340:
2338:
2336:
2332:
2324:
2320:
2319:
2318:
2317:
2314:
2311:
2309:
2305:
2297:
2293:
2292:
2291:
2290:
2287:
2284:
2282:
2278:
2270:
2266:
2265:
2264:
2263:
2260:
2257:
2255:
2251:
2247:
2239:
2235:
2232:
2230:
2226:
2223:
2221:
2217:
2214:
2212:
2208:
2206:
2202:
2200:
2196:
2195:
2190:
2186:
2182:
2175:
2170:
2168:
2163:
2161:
2156:
2155:
2152:
2146:
2143:
2142:
2138:
2129:
2123:
2119:
2114:
2113:
2104:
2101:
2096:
2090:
2086:
2081:
2080:
2074:
2068:
2065:
2060:
2054:
2050:
2045:
2044:
2035:
2032:
2027:
2023:
2019:
2015:
2011:
2007:
2003:
1996:
1993:
1988:
1984:
1979:
1974:
1970:
1966:
1962:
1958:
1954:
1950:
1946:
1939:
1936:
1931:
1927:
1922:
1917:
1913:
1909:
1904:
1899:
1895:
1891:
1887:
1883:
1882:
1877:
1870:
1867:
1862:
1858:
1854:
1850:
1846:
1842:
1838:
1834:
1830:
1826:
1822:
1815:
1813:
1809:
1804:
1800:
1796:
1790:
1786:
1785:
1777:
1774:
1769:
1765:
1760:
1755:
1751:
1747:
1743:
1739:
1735:
1728:
1726:
1724:
1720:
1713:
1709:
1706:
1705:
1701:
1699:
1697:
1693:
1690:
1686:
1682:
1681:
1677:
1673:
1672:
1667:
1666:
1665:Udanoceratops
1661:
1660:
1659:Protoceratops
1656:
1652:
1651:
1646:
1645:
1641:
1637:
1636:
1631:
1630:
1625:
1624:
1620:
1616:
1615:
1610:
1609:
1604:
1600:
1596:
1592:
1588:
1584:
1580:
1576:
1572:
1568:
1564:
1560:
1556:
1552:
1548:
1540:
1538:
1536:
1532:
1522:
1521:
1513:
1512:
1504:
1503:
1495:
1494:
1486:
1485:
1477:
1476:
1468:
1467:
1459:
1458:
1450:
1449:
1441:
1440:
1432:
1431:
1423:
1422:
1414:
1413:
1405:
1404:
1396:
1395:
1387:
1386:
1383:
1382:
1381:
1374:
1373:
1370:
1369:
1366:
1365:
1364:
1357:
1356:
1350:
1349:
1346:
1345:
1342:
1340:
1336:
1335:
1328:
1327:
1321:
1320:
1317:
1316:
1308:
1307:
1304:
1303:
1302:
1295:
1294:
1291:
1290:
1287:
1285:
1281:
1280:
1273:
1272:
1266:
1265:
1259:
1258:
1255:
1254:
1251:
1249:
1245:
1244:
1237:
1236:
1230:
1227:
1226:
1223:
1222:
1214:
1213:
1205:
1204:
1196:
1195:
1192:
1190:
1186:
1185:
1184:Dromaeosaurus
1178:
1177:
1174:
1173:
1170:
1168:
1164:
1163:
1156:
1155:
1149:
1148:
1145:
1144:
1141:
1139:
1135:
1134:
1127:
1126:
1120:
1117:
1116:
1110:
1109:
1103:
1102:
1099:
1098:
1095:
1094:
1093:
1086:
1085:
1079:
1078:
1075:
1074:
1071:
1069:
1065:
1064:
1057:
1056:
1050:
1047:
1046:
1040:
1039:
1036:
1035:
1027:
1026:
1023:
1022:
1021:
1020:Hesperonychus
1014:
1013:
1010:
1009:
1006:
1004:
1000:
999:
992:
991:
988:
987:
984:
982:
978:
977:
970:
969:
966:
965:
962:
961:
960:
959:Graciliraptor
953:
952:
949:
948:
945:
943:
939:
933:
932:
929:
928:
925:
923:
919:
918:
917:Changyuraptor
911:
910:
904:
903:Microraptoria
901:
900:
894:
893:
887:
886:
883:
882:
879:
877:
873:
872:
865:
864:
858:
857:
854:
853:
850:
849:
848:
841:
840:
834:
833:
830:
829:
821:
820:
812:
811:
803:
802:
799:
798:
797:
790:
789:
786:
785:
782:
781:
780:
773:
772:
769:
768:
765:
763:
759:
758:
757:Neuquenraptor
751:
750:
747:
746:
743:
742:
741:
734:
733:
727:
726:
723:
722:
719:
717:
713:
712:
705:
704:
698:
695:
694:
688:
687:
681:
680:
677:
676:
668:
667:
659:
658:
655:
653:
649:
648:
641:
640:
637:
636:
633:
631:
627:
626:
619:
618:
612:
611:
608:
607:
604:
602:
598:
597:
596:Halszkaraptor
590:
589:
583:
580:
579:
573:
572:
566:
563:
562:
558:
556:
555:
554:Halszkaraptor
550:
545:
543:
540:
537:
533:
529:
525:
521:
517:
513:
509:
501:
499:
497:
493:
489:
488:Archaeopteryx
485:
481:
477:
473:
469:
468:
463:
462:
457:
456:
455:Archaeopteryx
451:
447:
443:
439:
435:
429:
424:
418:
415:as a generic
414:
409:
402:
396:
392:
390:
386:
382:
378:
374:
370:
366:
362:
361:dromaeosaurid
358:
354:
350:
346:
342:
339:
335:
331:
328:
325:
321:
316:
308:
307:
296:
290:
289:
281:
278:
274:
270:
266:
260:
259:
252:
249:
248:
245:
239:
236:
235:
232:
226:
223:
222:
219:
216:
213:
210:
209:
206:
203:
200:
197:
196:
193:
190:
187:
184:
183:
180:
177:
174:
173:
170:
167:
164:
163:
160:
157:
154:
153:
148:
143:
139:
134:
130:
125:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
50:
43:
37:
33:
30:
19:
3046:
2981:
2959:Velociraptor
2957:
2950:
2943:
2935:
2927:
2919:
2912:
2905:
2898:
2890:
2882:
2875:
2870:Acheroraptor
2868:
2848:
2841:
2834:
2826:
2819:
2812:
2805:
2797:
2791:Dakotaraptor
2789:
2784:Achillobator
2782:
2762:
2754:
2749:Atrociraptor
2747:
2728:Zhenyuanlong
2726:
2719:
2713:Tianyuraptor
2711:
2704:
2697:
2689:
2665:
2658:
2652:Tianyuraptor
2650:
2643:
2636:
2628:
2621:
2614:
2593:
2585:
2577:
2570:
2562:
2554:
2547:
2539:
2532:
2525:
2519:Dakotaraptor
2517:
2512:Buitreraptor
2510:
2505:Austroraptor
2503:
2495:Unenlagiinae
2482:
2476:
2475:
2468:
2461:
2442:Zhenyuanlong
2440:
2432:
2427:Tianyuraptor
2425:
2417:
2410:
2403:
2367:
2233:
2224:
2215:
2111:
2103:
2078:
2067:
2042:
2034:
2009:
2005:
1995:
1952:
1948:
1938:
1885:
1879:
1869:
1828:
1824:
1783:
1776:
1741:
1737:
1684:
1680:Pinacosaurus
1678:
1676:ankylosaurid
1671:Plesiohadros
1669:
1663:
1657:
1655:ceratopsians
1648:
1642:
1633:
1627:
1621:
1619:oviraptorids
1612:
1608:Byronosaurus
1606:
1603:Velociraptor
1602:
1575:crocodilians
1544:
1534:
1530:
1528:
1378:
1377:
1361:
1360:
1334:Velociraptor
1332:
1331:
1299:
1298:
1277:
1276:
1241:
1240:
1182:
1181:
1160:
1159:
1133:Achillobator
1131:
1130:
1092:Tianyuraptor
1090:
1089:
1061:
1060:
1018:
1017:
996:
995:
974:
973:
957:
956:
936:
915:
914:
871:Zhenyuanlong
869:
868:
845:
844:
794:
793:
777:
776:
755:
754:
740:Buitreraptor
738:
737:
711:Austroraptor
709:
708:
697:Unenlagiinae
645:
644:
624:
623:
622:
594:
593:
552:
546:
531:
527:
524:autapomorphy
511:
505:
487:
465:
459:
453:
433:
432:
412:
359:was a small
356:
305:
304:
303:
294:
287:
286:
277:Type species
264:
257:
256:
211:
198:
185:
35:
29:
2914:Linheraptor
2892:Deinonychus
2843:Yurgovuchia
2799:Deinonychus
2756:Bambiraptor
2721:Vectiraptor
2699:Bambiraptor
2638:Microraptor
2549:Pamparaptor
2484:Natovenator
2323:Maniraptora
2308:Maniraptora
2012:(1): 1–19.
1888:(1): 5105.
1380:Linheraptor
1279:Deinonychus
1063:Bambiraptor
976:Microraptor
461:Caudipteryx
403:Description
385:dharmapalas
365:troodontids
353:Gobi Desert
237:Subfamily:
3154:Categories
3010:See also:
2884:Boreonykus
2836:Utahraptor
2587:Variraptor
2556:Pyroraptor
2434:Variraptor
2412:Pyroraptor
2344:see below↓
2220:Dinosauria
2211:Sauropsida
1714:References
1689:predaceous
1674:, and the
1567:sandstorms
1162:Utahraptor
492:metatarsal
205:Saurischia
192:Dinosauria
3144:Dinosaurs
2877:Adasaurus
2572:Unenlagia
2564:Rahonavis
2470:Hulsanpes
2296:Theropoda
2281:Theropoda
2229:Theropoda
2197:Kingdom:
2026:0375-7633
1969:2045-2322
1955:: 16392.
1912:2045-2322
1803:985402380
1692:carnivore
1635:Oviraptor
1551:sand dune
1529:In 2020,
1243:Adasaurus
647:Hulsanpes
549:cladogram
539:subfamily
516:theropods
480:thighbone
472:paravians
442:vertebrae
334:Campanian
332:from the
218:Theropoda
165:Kingdom:
159:Eukaryota
3056:Wikidata
3047:Mahakala
3022:Category
3012:Timeline
2929:Nuthetes
2850:Zapsalis
2821:Itemirus
2595:Ypupiara
2477:Mahakala
2405:Daurlong
2205:Chordata
2203:Phylum:
2199:Animalia
1987:39054320
1978:11272788
1930:32218481
1853:29211712
1768:17823350
1702:See also
1685:Mahakala
1650:Shuvuuia
1623:Citipati
1555:alluvial
1547:semiarid
1535:Mahakala
1531:Mahakala
625:Mahakala
532:Mahakala
528:Mahakala
520:paravian
512:Mahakala
434:Mahakala
413:Mahakala
381:Mahakala
369:avialans
357:Mahakala
349:Mongolia
345:Ömnögovi
330:dinosaur
327:theropod
306:Mahakala
258:Mahakala
224:Family:
179:Chordata
175:Phylum:
169:Animalia
155:Domain:
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2238:Paraves
2209:Class:
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1890:Bibcode
1861:4471941
1833:Bibcode
1746:Bibcode
1738:Science
1583:mammals
1579:lizards
1571:turtles
1363:Tsaagan
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293:Turner
263:Turner
250:Genus:
116:↓
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847:Shanag
464:, and
446:pelvis
297:, 2007
295:et al.
265:et al.
2234:Clade
2225:Clade
2216:Clade
1857:S2CID
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1595:fauna
1589:like
536:basal
496:basal
484:ilium
387:) in
377:birds
320:genus
212:Clade
199:Clade
186:Clade
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