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Majungasaurus

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1747: 6335: 8467: 1436: 6328: 4832: 5657: 5643: 1212: 1383: 7919: 7123: 7905: 5636: 2225: 1540: 7912: 4838: 1054:, like other majungasaurines, lacked the characteristic pits and grooves where claws and tendons would normally attach, and its finger bones were fused together, indicating that the hand was immobile. In 2012, a better specimen was described, showing that the lower arm was robust, though short, and that the hand contained four metatarsals and four, probably inflexible and very reduced, fingers, possibly with no claws. The minimum phalanx formula was 1-2-1-1. 6342: 1952: 1293: 154: 5650: 586: 8480: 7141: 864: 133: 7129: 1349: 7148: 8474: 757: 7899: 962: 7136: 649: 2211: 4239: 1663: 1271: 753:, some of which were attached to an isolated premaxilla that was described in 1996. The following seven expeditions would turn up tens of thousands of fossils, many of which belonged to species new to science. The Mahajanga Basin Project claims credit for quintupling the known diversity of fossil taxa in the region. 1565:, and perhaps other abelisaurids, specialized on hunting sauropods. Adaptations to strengthen the head and neck for a bite-and-hold type of attack might have been very useful against sauropods, which would have been tremendously powerful animals. This hypothesis may also be supported by the hindlegs of 847:. Although the monograph is comprehensive, the editors noted that it describes only material recovered from 1993 through 2001. A significant quantity of specimens, some very complete, were excavated in 2003 and 2005 and await preparation and description in future publications. The dentary was made the 1919:
different points, forming a solid bony mass. There is no sign of any other vertebrae after the fifth in the series, indicating that the tail ended there prematurely. From the size of the last vertebrae, scientists judged that about ten vertebrae were lost. One explanation for this pathology is severe
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too fragmentary to confidently assign to the same species as the skull. Further fieldwork over the next decade turned up a series of less complete skulls, as well as dozens of partial skeletons of individuals ranging from juveniles to adults. Project members also collected hundreds of isolated bones
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had an even broader snout than other abelisaurids, and other aspects of its anatomy may also support the bite-and-hold hypothesis. The neck was strengthened, with robust vertebrae, interlocking ribs and ossified tendons, as well as reinforced muscle attachment sites on the vertebrae and the back of
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The most serious pathology discovered was in a series of five large tail vertebrae. The first two vertebrae showed only minor abnormalities with the exception of a large groove that extended along the left side of both bones. However, the next three vertebrae were completely fused together at many
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and other abelisaurids indicate different predatory habits than other theropods. Whereas most theropods were characterized by long, low skulls of narrow width, abelisaurid skulls were taller and wider, and often shorter in length as well. The narrow skulls of other theropods were well equipped to
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through these channels at the beginning of the wet season, burying the carcasses of organisms killed during the preceding dry season and providing for their exceptional preservation as fossils. Sea levels in the area were rising throughout the Maastrichtian, and would continue to do so into the
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specimens observed amongst those recovered indicates most well preserved individuals generally lack observable pathologies, while a few select individuals were shown to have possessed multiple pathologies, a general pattern also noted in other large, nonavian theropods. Such patterns may be the
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line, to which birds belong, occurred very early in the history of theropods. The avian respiratory system, present in both lines, must therefore have evolved before the split, and well before the evolution of birds themselves. This provides further evidence of the dinosaurian
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is rotated so that its lateral canal is parallel to the ground, the entire skull is nearly horizontal. This contrasts with many other theropods, where the head was more strongly downturned when in the alert position. The lateral canal is also significantly longer in
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would not have moved as fast as other similar-sized theropods, it would have had no trouble keeping up with slow-moving sauropods. The robust hindlimb bones suggest very powerful legs, and their shorter length would have lowered the animal's center of gravity. Thus
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was a medium-sized theropod that typically reached 5.6–7 m (18–23 ft) in length and weighed 750–1,100 kg (1,650–2,430 lb). Fragmentary remains of larger individuals indicate that some adults could have been similar in size to its relative
1516:. This may have been an adaptation to prevent the fracture of the lower jaw when holding onto a struggling prey animal. The front teeth of the upper jaw were more robust than the rest, to provide an anchor point for the bite, while the low crown height of 1535:
indicates that the theropod replaced its teeth anywhere from 2 to 13 times faster than other theropods, replacing the entire set within a span of two months. Gnawing on bone may have been a significant reason for such rapid tooth replacement.
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result of a snowball effect where one injury or an infection increased the likelihood of additional maladies and injuries due to functional impairment or compromised immune systems in individuals once an initial injury had occurred.
1861:, this was the first time an abelisauroid had been examined in this manner. No wounds were found on any skull elements, in contrast to tyrannosaurids where sometimes gruesome facial bites were common. One of the specimens was a 1978:) seems to have become taller and more robust; additionally, the skull bones became more fused and the eye sockets became proportionally smaller. This indicates a shift in dietary preferences between juveniles and adults. 891:
is exceptionally well known compared to most theropods and generally similar to that of other abelisaurids. Like other abelisaurid skulls, its length was proportionally short for its height, although not as short as in
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Tortosa, Thierry; Eric Buffetaut; Nicolas Vialle; Yves Dutour; Eric Turini; Gilles Cheylan (2013). "A new abelisaurid dinosaur from the Late Cretaceous of southern France: Palaeobiogeographical implications".
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had teeth curved on the front edge but straighter on the back (cutting) edge. This structure may have served to prevent slicing, and instead holding the teeth in place when biting. Examination of the teeth of
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Depéret, Charles (1896). "Note sur les Dinosauriens Sauropodes et Théropodes du Crétacé supérieur de Madagascar" [Note on the Dinosaurs Sauropods and Theropods of the Upper Cretaceous of Madagascar].
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with a short snout. Although the forelimbs are not completely known, they were very short, while the hind limbs were longer and very stocky. It can be distinguished from other abelisaurids by its wider
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Lavocat, René (1955). "Sur une portion de mandibule de Théropode provenant du Crétacé supérieur de Madagascar" [On a portion of Theropod's mandible from the Upper Cretaceous of Madagascar].
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Depéret, Charles; Savornin, Justin (1928). "La faune de Reptiles et de Poissons albiens de Timimoun (Sahara algérien)" [The fauna of Reptiles and Albian Fish of Timimoun (Algerian Sahara)].
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took twenty years to reach maturity, which may have been a result of the harsh environment in which it lived. However, other abelisaurids have also been found to have comparably slow growth rates.
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vertebrae imply the presence of these air sacs. These air sacs may have allowed for a basic form of avian-style 'flow-through ventilation,' where air flow through the lungs is one-way, so that
916:, which were extremely thick and fused together, with a low central ridge running along the half of the bone closest to the nostrils. A distinctive dome-like horn protruded from the fused 1725:, besides providing an understanding of its respiratory biology, also has larger-scale implications for evolutionary biology. The split between the ceratosaur line, which led to 2914:
Agnolin, Federico L.; Chiarelli, Pablo (2009). "The position of the claws in Noasauridae (Dinosauria: Abelisauroidea) and its implications for abelisauroid manus evolution".
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in the same region where the original material was found. The teeth matched those first described by Depéret, but the strongly curved jaw bone was very different from both
522:, the very rough texture and thickened bone on the top of its snout, and the single rounded horn on the roof of its skull, which was originally mistaken for the dome of a 4036:
Forster, Catherine; Sampson, Scott D.; Chiappe, Luis M.; Krause, David W. (1998). "The theropod ancestry of birds: new evidence from the Late Cretaceous of Madagascar".
2584:"Case 3487: Megalosaurus crenatissimus Depéret, 1896 (currently Majungasaurus crenatissimus; Dinosauria, Theropoda): proposed replacement of the holotype by a neotype" 1938:
suggest that the multitude of injuries that occurred were obtained over the course of the lives of the individuals studied. Furthermore, the small number of injured
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as the only non-avian theropod with confirmed cannibalistic tendencies, although there is some evidence that cannibalism may have occurred in other species as well.
1892:(benign bone tumor) was deemed unlikely. Another specimen, a small caudal (tail) vertebra, was also found to have an abnormal growth, this time on the top of its 1050:
had very short forelimbs with four extremely reduced digits, first reported with only two very short external fingers and no claws. The hand and finger bones of
1962:, being known from many well-preserved specimens of different ages, is well studied in regards to its growth and development. Throughout ontogeny, the skull of 639:("most"), in reference to the numerous serrations on both front and rear edges of the teeth. Depéret later reassigned the species to the North American genus 8713: 2622:"Opinion 2269 (Case 3487): Megalosaurus crenatissimus Depéret, 1896 (currently Majungasaurus crenatissimus; Dinosauria, Theropoda): designation of a neotype" 1606:
have been discovered bearing tooth marks identical to those found on sauropod bones from the same localities. These marks have the same spacing as teeth in
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is the only large theropod known from the area, the simplest explanation is that it was feeding on other members of its own species. Suggestions that the
2061: 1849:. Scientists examined the remains of at least 21 individuals and discovered four with noticeable pathologies. While pathology had been studied in large 672: 478: 2302:; Sampson, Scott D.; Carrano, Matthew T.; O'Connor, Patrick M. (2007). "Overview of the history of discovery, taxonomy, phylogeny, and biogeography of 2382:
Sues, Hans-Dieter; Taquet, Phillipe (1979). "A pachycephalosaurid dinosaur from Madagascar and a Laurasia−Gondwanaland connection in the Cretaceous".
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is perhaps most distinctive for its skull ornamentation, including the swollen and fused nasals and the frontal horn. Other ceratosaurs, including
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O'Connor, Patrick M.; Claessens, Leon P.A.M. (2006). "Basic avian pulmonary design and flow-through ventilation in non-avian theropod dinosaurs".
6334: 4252: 8738: 8733: 6327: 8466: 547:. It appears to be most closely related to abelisaurids from India rather than South America or continental Africa, a fact that has important 4130:
Sampson, Scott D.; Carrano, Matthew T.; Forster, Catherine A. (2001). "A bizarre predatory dinosaur from the Late Cretaceous of Madagascar".
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was markedly wider than in other abelisaurids. All abelisaurids had a rough, sculptured texture on the outside faces of the skull bones, and
3501: 1077:(ankle bones) were fused together, and the feet bore three functional digits, with a smaller first digit that did not contact the ground. 839:. In this volume, the dentary described by Lavocat was re-evaluated and determined to be diagnostic for this species. Therefore, the name 2434: 2037:
teeth have also been found in the underlying Masorobe Member and the overlying Miadana Member. While these sediments have not been dated
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Sampson, S. D. (May 15, 1998). "Predatory Dinosaur Remains from Madagascar: Implications for the Cretaceous Biogeography of Gondwana".
3686: 3050: 1927:(infection) of the last remaining vertebrae. Alternatively, the infection may have come first and led to the end of the tail becoming 826: 5656: 4079:
Makovicky, Peter J.; ApesteguĂ­a, Sebastian; AgnolĂ­n, Federico L. (2005). "The earliest dromaeosaurid theropod from South America".
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in northwestern Madagascar. Most of these, including all of the most complete material, came from the Anembalemba Member, although
1115:. Abelisaurids are known for their tall skulls with blunt snouts, extensive sculpturing on the outer surfaces of the facial bones ( 7122: 3993:
Forster, Catherine A.; Chiappe, Luis M.; Krause, David W.; Sampson, Scott D. (1996). "The first Cretaceous bird from Madagascar".
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Rogers, Raymond R. (2005). "Fine-grained debris flows and extraordinary vertebrate burials in the Late Cretaceous of Madagascar".
1896:, which projects upwards from the vertebrae, allowing muscle attachment. Similar growths from the neural spine have been found in 8718: 812:
teeth. Taken together, these remains represent nearly all the bones of the skeleton, although most of the forelimbs, most of the
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The genus is one of the first abelisaurs to be discovered, first being found in 1896 (although it was thought to be a species of
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would have weakened the structure and probably precluded its use in direct physical combat, although the horn may have served a
7904: 2238: 4831: 2751: 900:(frontmost upper jaw bone), which made the tip of the snout very blunt, was also typical of the family. However, the skull of 8632: 3823: 2151: 5635: 3223:; Smith, Paula M.; Thomason, Jeffrey J.; Upchurch, Paul (2001). "Cranial design and function in a large theropod dinosaur". 671:
in northwestern Madagascar were recovered by French collectors over the next 100 years, many of which were deposited in the
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Most of the pathologies occurred on the vertebrae. For example, a dorsal (back) vertebra from a juvenile animal showed an
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8678) recovered from the Maevarano Formation. Most of these vertebrae and some of the ribs contained cavities (pneumatic
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PR 2100). On top of this skull was a dome-shaped swelling nearly identical to the one described by Sues and Taquet as
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between certain bones that allowed a high degree of flexibility in the lower jaw, although not to the extent seen in
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Gutherz, Samuel B.; Groenke, Joseph R.; Sertich, Joseph J. W.; Burch, Sara H.; O’Connor, Patrick M. (July 2, 2020).
8479: 3871:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar". In Sampson, Scott D.; Krause, David W. (eds.). 3614:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar". In Sampson, Scott D.; Krause, David W. (eds.). 3000:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar". In Sampson, Scott D.; Krause, David W. (eds.). 2845:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar". In Sampson, Scott D.; Krause, David W. (eds.). 2803:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar". In Sampson, Scott D.; Krause, David W. (eds.). 2306:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar". In Sampson, Scott D.; Krause, David W. (eds.). 1811: 1683: 1557:
was the largest predator in its environment, while the only known large herbivores at the time were sauropods like
738: 7147: 1698:. In birds, the neck vertebrae and ribs are hollowed out by the cervical air sac, the upper back vertebrae by the 1435: 2187:
was by far the largest carnivore and probably the dominant predator on land, although large crocodylomorphs like
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from mainland Africa. This leaves open the possibility of separate clades of abelisaurids in western and eastern
3706: 3157: 8723: 4382: 4243: 3303: 1489: 7128: 3971: 3630: 3521: 3016: 2861: 2819: 8473: 3887: 2777: 1569:, which were short and stocky, as opposed to the longer and more slender legs of most other theropods. While 1411:, although their exact function within that context is unknown. The hollow cavity inside the frontal horn of 4287: 2949:
Burch, S.H.; Carrano, M.T. (2012). "An articulated pectoral girdle and forelimb of the abelisaurid theropod
2548: 2435:"The premaxilla of Majungasaurus (Dinosauria: Theropoda), with implications for Gondwanan paleobiogeography" 2322: 1897: 1769:
structure. Overall, the brain was very small relative to body size, but otherwise similar to many other non-
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the skull. These muscles would have been able to hold the head steady despite the struggles of its prey.
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sediments near the village of Berivotra, in Mahajanga Province. Among these scientists was paleontologist
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is usually parallel to the ground when the animal holds its head in an alert posture. When the skull of
574: 307: 3956:"Late Cretaceous terrestrial vertebrates from Madagascar: implications for Latin American biogeography" 7898: 1931:
and falling off. This is the first example of tail truncation known in a non-avian theropod dinosaur.
8641: 8546: 8431: 6542: 6510: 4249: 4193: 4139: 4088: 4045: 4002: 3918: 3753: 3559: 3406: 3338: 3291: 3279: 3232: 3189: 2962: 2672: 2485: 2446: 2391: 1912: 1908: 1116: 932:(CT scanning) of the skull shows that both the nasal structure and the frontal horn contained hollow 921: 741:
began the Mahajanga Basin Project, a series of expeditions to examine the fossils and geology of the
1803:, which also had a small flocculus, did not rely on quick head movements to sight and capture prey. 1650:
will then proceed to cannibalize the remains of their rivals, which may suggest similar behavior in
539:) and being named in 1955. Known from several well-preserved skulls and abundant skeletal material, 6866: 6653: 4184:; Forster, Catherine A. (2001). "The last of the dinosaur titans: a new sauropod from Madagascar". 3109: 2230: 2124: 2103: 2072: 1907:, probably resulting from the ossification of a ligament running either between the neural spines ( 1754: 1500:, where the nasal bones were fused and thickened for strength. On the other hand, the lower jaw of 1465:, most theropods may have used a strategy similar in some ways to that of long- and narrow-snouted 1458: 1382: 1120: 929: 783:
Fieldwork in 1996 turned up a spectacularly complete theropod skull preserved in exquisite detail (
688: 544: 7135: 1539: 8537: 8338: 6842: 6702: 6558: 6498: 5933: 5843: 5726: 4590: 4280: 4217: 4163: 4112: 4018: 3975: 3891: 3867:; Rasoamiaramanana, Armand H.; Rahantarisoa, Lydia (2007). "Paleoenvironment and Paleoecology of 3710: 3634: 3583: 3525: 3430: 3325:
D'Emic, M.D.; O'Connor, P.M.; Pascucci, T.R.; Gavras, J.N.; Mardakhayava, E.; Lund, E.K. (2019).
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on top of the skull as well. In life, these structures would have been covered with some sort of
832: 668: 663:.MAJ 1), the right dentary of a subadult individual – MusĂ©um national d'histoire naturelle, Paris 336: 148: 3533: 749:
of Stony Brook. The first expedition turned up hundreds of theropod teeth identical to those of
597: 327: 8685: 8590: 1951: 1598:, discoveries published in 2007 detail finds in Madagascar that indicate the presence of other 1586:
bones confirm that it at least fed on these sauropods, whether or not it actually killed them.
1292: 8667: 8572: 8220: 8137: 8111: 7950: 7937: 7815: 7356: 7308: 6140: 5490: 4569: 4209: 4155: 4104: 4061: 3665: 3657: 3575: 3479: 3422: 3366: 3248: 3117: 3077: 2715: 2705: 2552: 2533:"Majungasaurus crenatissimus (Theropoda:Abelisauridae) from the late Cretaceous of Madagascar" 2501: 2462: 1424: 1416: 1097: 992: 836: 617: 467: 8672: 8577: 8324: 8081: 7991: 7780: 7696: 7558: 7477: 7449: 7409: 7290: 7260: 6778: 6531: 6395: 6252: 6152: 6001: 5908: 5866: 5433: 4743: 4201: 4147: 4096: 4053: 4010: 3967: 3926: 3883: 3798: 3761: 3702: 3626: 3567: 3517: 3469: 3461: 3414: 3356: 3346: 3299: 3240: 3197: 3153: 3101: 3067: 3059: 3012: 2970: 2923: 2896: 2857: 2815: 2773: 2680: 2633: 2595: 2544: 2493: 2454: 2399: 2318: 2216: 1971: 1186: 1093: 1004: 896:. The skulls of large individuals measured 60–70 centimeters (24–28 in) long. The tall 821: 795:
was redescribed as an abelisaurid rather than a pachycephalosaur in 1998. Although the name
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cavities, perhaps to reduce weight. The teeth were typical of abelisaurids in having short
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and most other theropods, which were curved on both the front and back, abelisaurids like
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Like other abelisaurids, the hindlimbs were stocky and short compared to body length. The
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with comparisons to allosaur pathology in the Cleveland-Lloyd Dinosaur Quarry collection"
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Grillo, O.N.; Delcourt, R. (2016). "Allometry and body length of abelisauroid theropods:
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teeth are found up until the very end of the Maastrichtian, when all non-avian dinosaurs
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described a dome-shaped skull fragment (MNHN.MAJ 4) as a new genus of pachycephalosaur (
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teeth prevented them from breaking off during a struggle. Finally, unlike the teeth of
1509: 1496:, creating the characteristic rough texture of the bones. This is particularly true of 1348: 1105: 730:) in 1979. This was the first report of a pachycephalosaur in the Southern Hemisphere. 704: 594: 496: 237: 8702: 8394: 8240: 8201: 8180: 8059: 8052: 7979: 7773: 7746: 7604: 7520: 7498: 7491: 7484: 7427: 7344: 7207: 7179: 7166: 7090: 6967: 6807: 6641: 6572: 6458: 6445: 6190: 6168: 6161: 6125: 6098: 6050: 5974: 5710: 5501: 5475: 5391: 5372: 5302: 5281: 5260: 5245: 5231: 5190: 5156: 5149: 5068: 4920: 4807: 4800: 4727: 4697: 4686: 4576: 4547: 4540: 4466: 4167: 3895: 3448:
Nesbitt, Sterling J.; Turner, Alan H.; Erickson, Gregory M.; Norell, Mark A. (2006).
3165: 3137: 2869: 2785: 2171: 2161: 2050: 2017:, most within 50 kilometers (31 mi) to the southeast of the provincial capital, 1924: 1862: 1858: 1770: 1643: 1614:
teeth, and contain smaller notches consistent with the serrations on those teeth. As
1480:, with short and broad snouts, that bite once and hold on until the prey is subdued. 1407:
itself bore crests on the head. These structures are likely to have played a role in
1326: 1128: 937: 556: 250: 83: 45: 4116: 3714: 3638: 3529: 3311: 3024: 2982: 2935: 2827: 2645: 2607: 2564: 2513: 2330: 1872:(bony growth) on its underside. The growth probably resulted from the conversion of 8366: 8304: 8274: 8194: 8187: 8165: 8157: 8037: 8029: 8009: 7724: 7635: 7573: 7463: 7434: 7386: 7215: 7200: 6975: 6907: 6835: 6814: 6793: 6609: 6565: 6373: 6360: 6290: 6243: 6118: 6008: 5960: 5953: 5890: 5859: 5852: 5733: 5560: 5412: 5309: 5211: 5204: 5099: 5083: 5019: 4970: 4902: 4679: 4562: 4555: 4526: 4489: 4458: 4221: 4022: 3979: 3653: 3587: 3434: 3275: 3260: 3105: 2900: 2458: 2430: 2411: 2176: 2053: 1986: 1985:
indicates that it was among the slowest-growing theropods. Based on studies of the
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Inferences about behavior can also be drawn from examination of the inner ear. The
1714:-rich air inhaled from outside the body is never mixed with exhaled air laden with 1445: 1403: 1158:. The most recent analysis, using the most complete information, instead recovered 1112: 1016: 987:, the only other abelisaurid genera for which complete skeletal material is known. 917: 768: 756: 641: 604:
in 1896. These included two teeth, a claw, and some vertebrae discovered along the
569: 548: 535: 477:, making it one of the last-known non-avian dinosaurs that went extinct during the 346: 4057: 3955: 3766: 3741: 3144:
Bonaparte 1985 (Theropoda: Abelisauridae) from the Late Cretaceous of Patagonia".
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Burch, S.; Carrano, M. (2008). "Abelisaurid forelimb evolution: New evidence from
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less than 20 million years earlier. It was drifting northwards but still 10 to 15
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to bone during development, but the cause of the ossification was not determined.
1865:(toe bone) of the foot, which had apparently been broken and subsequently healed. 1457:
withstand the vertical stress of a powerful bite, but not as good at withstanding
8522: 3351: 3201: 2974: 2532: 8626: 8531: 8420: 8401: 8331: 8281: 8267: 8209: 8150: 8095: 7972: 7766: 7642: 7611: 7527: 7513: 7245: 7193: 7061: 6983: 6946: 6828: 6800: 6753: 6673: 6551: 6408: 6259: 6232: 6132: 6079: 6015: 5994: 5967: 5718: 5531: 5524: 5510: 5336: 5183: 5174: 5076: 5047: 4977: 4942: 4935: 4735: 4713: 4706: 4665: 4657: 4633: 4618: 4597: 4505: 4497: 3954:; Sampson, Scott D.; Buckley, Gregory A.; Rogers, Raymond R. (August 23, 2006). 3041: 2120: 2107: 2096: 1695: 1623: 1265: 1196: 1135: 1101: 880: 453: 58: 961: 816:
and the tip of the tail are still unknown. This fieldwork culminated in a 2007
8359: 8288: 8233: 7854: 7653: 7470: 7456: 7441: 7393: 7327: 7253: 7075: 7068: 7054: 6960: 6893: 6821: 6785: 6738: 6680: 6666: 6633: 6625: 6618: 6599: 6422: 6387: 6175: 5776: 5567: 5517: 5454: 5425: 5380: 5358: 5351: 5343: 5267: 5163: 5122: 5115: 5107: 5091: 5011: 4882: 4758: 4611: 4474: 4333: 3783:"Ontogenetic changes in the craniomandibular skeleton of abelisaurid dinosaur 2927: 2556: 2206: 2014: 1967: 1854: 1799: 1786: 1774: 1522: 1476:, may instead have been adapted for a feeding strategy more similar to modern 1343: 1309: 1180: 1170: 1164: 1109: 1070: 983: 970: 909: 897: 601: 471: 463: 211: 103: 68: 8617: 3116:(Second ed.). Berkeley: University of California Press. pp. 47–70. 2719: 991:
was bipedal, with a long tail to balance out the head and torso, putting the
573:, and is also one of the few dinosaurs for which there is direct evidence of 8380: 8373: 8073: 8066: 7739: 7731: 7662: 7315: 7033: 6579: 6483: 6218: 5803: 5688: 5675: 5602: 5538: 5461: 5440: 5197: 5060: 4868: 4855: 4773: 4625: 4444: 4431: 4394: 4303: 2165: 2018: 1885: 1869: 1850: 1842: 1766: 1730: 1639: 1462: 1155: 1143: 1139: 1074: 817: 608:
by a French army officer and deposited in the collection of what is now the
560: 440: 224: 165: 108: 52: 4213: 4159: 4108: 3803: 3782: 3579: 3483: 3465: 3426: 3370: 3252: 3081: 3063: 1022:
attached to the cervical ribs, giving them a forked appearance, as seen in
648: 4238: 4065: 2637: 2599: 2505: 2001: 1687: 8611: 8516: 7964: 7372: 7365: 7337: 7230: 6730: 6111: 5447: 5274: 5035: 4672: 4351: 4327: 2080: 1928: 1877: 1619: 1231:. A simplified version showing the taxa within the group is shown below. 1205: 1000: 564: 514: 459: 456: 198: 185: 98: 93: 78: 73: 63: 4100: 3571: 3418: 1833:, indicating a greater sensitivity to side-to-side motions of the head. 1662: 1270: 543:
has recently become one of the best-studied theropod dinosaurs from the
8659: 8564: 7505: 7267: 6997: 6900: 6225: 5546: 2084: 1889: 1154:
from South America, while others were unable to firmly place it in the
1124: 1035: 949: 945: 925: 848: 652: 620:
containing any number of unrelated large theropods, as the new species
482: 113: 88: 3824:"Bone histology reveals unusual life history in the theropod dinosaur 3625:. Vol. 27. Journal of Vertebrate Paleontology. pp. 180–184. 3011:. Vol. 27. Journal of Vertebrate Paleontology. pp. 163–179. 2856:. Vol. 27. Journal of Vertebrate Paleontology. pp. 127–162. 2814:. Vol. 27. Journal of Vertebrate Paleontology. pp. 103–126. 1718:. This method of respiration, while complicated, is highly efficient. 1646:
sometimes kill each other when competing for access to carcasses. The
8173: 7861: 5288: 4321: 4205: 4151: 4014: 3930: 3450:"Prey choice and cannibalistic behaviour in the theropod Coelophysis" 3244: 2772:. Vol. 27. Journal of Vertebrate Paleontology. pp. 32–102. 2699: 2403: 2156: 2088: 1711: 1703: 1647: 1142:
relationships) within the family Abelisauridae are confused. Several
1019: 1008: 954: 813: 633: 175: 8493: 3882:. Vol. 27. Journal of Vertebrate Paleontology. pp. 21–31. 3140:; Chiappe, Luis M.; Dingus, Lowell (2002). "A new close relative of 2887:(Abelisauridae: Theropoda) from the late Cretaceous of Madagascar". 2317:. Vol. 27. Journal of Vertebrate Paleontology. pp. 1–20. 1038:(upper arm bone) was short and curved, closely resembling those of 7004: 6745: 6687: 3502:"A reevaluation of cooperative pack hunting and gregariousness in 2756:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar" 2100: 2000: 1950: 1762: 1745: 1661: 1642:
their carcasses. However, some researchers have noted that modern
1538: 1513: 1477: 1466: 1434: 1381: 1210: 1089: 1058: 960: 862: 755: 676: 647: 625: 584: 527: 519: 511: 449: 3876:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar 3781:
O'Connor, Patrick; Ratsimbaholison, Nirina; Felice, Ryan (2016).
3619:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar 3327:"Evolution of high tooth replacement rates in theropod dinosaurs" 3005:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar 2850:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar 2808:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar 2765:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar 2311:(Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar 2106:. Strong geological evidence suggests the occurrence of periodic 1508:(opening) on each side, as seen in other ceratosaurs, as well as 8646: 8551: 8437: 7047: 3687:"Multiple injury and infection in a sub-adult theropod dinosaur 2841:
O'Connor, Patrick M. (2007). "The postcranial axial skeleton of
2143: 2139: 1699: 1691: 1493: 1488:
Abelisaurid skulls were also strengthened in many areas by bone
1027: 996: 852: 784: 776: 684: 660: 8497: 7935: 7164: 6358: 5673: 4853: 4429: 4314: 4276: 3656:(2005). "Theropod paleopathology: state-of-the-art review". In 1034:
had long depressions along the sides for weight reduction. The
1026:. All of these features resulted in a very strong and muscular 3046:"New dinosaurs link southern landmasses in the Mid-Cretaceous" 1790: 1706:(hip) vertebrae by the abdominal air sac. Similar features in 1012: 3610:
Farke, Andrew A.; O'Connor, Patrick M. (2007). "Pathology in
3100:
Tykoski, Ronald B.; Rowe, Timothy (2004). "Ceratosauria". In
2799:
Smith, Joshua B. (2007). "Dental morphology and variation in
1469:, with the delivery of many bites weakening the prey animal. 1007:) to reduce their weight, they were robust, with exaggerated 713: 708: 412: 27:
Abelisaurid theropod dinosaur from the Late Cretaceous period
4272: 2582:; O'Connor, Patrick M.; Sampson, Scott D. (September 2009). 1934:
The small number of specimens preserved with pathologies in
1215:
Skeletal restoration showing known elements of two specimens
952:
of the lower jaw, more than in any other abelisaurid except
3707:
10.1671/0272-4634(2002)022[0076:MIAIIA]2.0.CO;2
3664:. Bloomington: Indiana University Press. pp. 351–365. 3158:
10.1671/0272-4634(2002)022[0460:ANCROC]2.0.CO;2
2543:(Supplement 2). Society of Vertebrate Paleontology: 1–184. 1761:
skull (FMNH PR 2100) allowed a rough reconstruction of its
418: 403: 397: 385: 3304:
10.1666/0094-8373(2002)028<0527:DAMPC>2.0.CO;2
2996:
Carrano, Matthew T. (2007). "The appendicular skeleton of
1773:
theropods, with a very conservative form closer to modern
1419:
purpose. While there is variation in the ornamentation of
1131:), and stocky hindlimb proportions, among other features. 391: 3972:
10.3417/0026-6493(2006)93[178:LCTVFM]2.0.CO;2
3631:
10.1671/0272-4634(2007)27[180:PIMCTA]2.0.CO;2
3522:
10.3374/0079-032X(2007)48[103:AROCPH]2.0.CO;2
3017:
10.1671/0272-4634(2007)27[163:TASOMC]2.0.CO;2
2862:
10.1671/0272-4634(2007)27[127:TPASOM]2.0.CO;2
2820:
10.1671/0272-4634(2007)27[103:DMAVIM]2.0.CO;2
1452:
Scientists have suggested that the unique skull shape of
824:
on all aspects of the animal's biology, published in the
400: 394: 3888:
10.1671/0272-4634(2007)27[21:PAPOMC]2.0.CO;2
2778:
10.1671/0272-4634(2007)27[32:CAOMCT]2.0.CO;2
2029:
have been recovered from the Maevarano Formation in the
1594:
Although sauropods may have been the prey of choice for
908:
was no exception. This was carried to an extreme on the
3545: 3543: 2549:
10.1671/0272-4634(2007)27[1:OOTHOD]2.0.CO;2
2323:
10.1671/0272-4634(2007)27[1:OOTHOD]2.0.CO;2
699:. Based on this dentary, Lavocat created the new genus 600:
described the first theropod remains from northwestern
884:, possibly exceeding 8 m (26 ft) in length. 703:, using an older spelling of Mahajanga as well as the 2056:, which lasted from 70 to 66 Ma (million years ago). 1627:
was a cannibal have been recently disproven, leaving
1174:
from India, but excluding South American genera like
530:
in both upper and lower jaws than most abelisaurids.
421: 409: 388: 3495: 3493: 2745: 2743: 2741: 2739: 2737: 2735: 2733: 2731: 2729: 415: 8601: 8506: 8419: 8314: 8257: 8219: 8136: 8109: 8019: 7989: 7948: 7846: 7814: 7756: 7706: 7679: 7652: 7621: 7594: 7583: 7542: 7407: 7355: 7326: 7288: 7277: 7177: 7089: 7023: 6945: 6876: 6852: 6764: 6712: 6652: 6608: 6597: 6541: 6509: 6456: 6406: 6371: 6289: 6242: 6189: 6151: 6097: 6048: 5984: 5943: 5919: 5888: 5842: 5812: 5801: 5686: 5601: 5500: 5411: 5390: 5326: 5229: 5173: 5139: 5046: 4991: 4960: 4900: 4866: 4792: 4772: 4696: 4643: 4524: 4442: 4393: 4366: 3605: 3603: 3601: 3599: 3597: 2138:, fossil taxa recovered from the Maevarano include 1923:resulting in the loss of the tail tip, followed by 1682:
based on a superbly preserved series of vertebrae (
406: 382: 3095: 3093: 3091: 1690:) that may have resulted from the infiltration of 831:. The papers are in English, although each has an 3942: 3940: 3858: 3856: 3854: 3852: 3850: 3848: 3510:Bulletin of the Peabody Museum of Natural History 3384: 3382: 3380: 3036: 3034: 2761:. In Sampson, Scott D.; Krause, David W. (eds.). 3393:(2007). "Cannibalism in the Madagascan dinosaur 2365:Bulletin du MusĂ©um National d'Histoire Naturelle 2294: 2292: 2290: 2288: 2286: 2284: 2282: 2280: 2278: 2750:Sampson, Scott D.; Witmer, Lawrence M. (2007). 2160:and several other birds, the possibly flighted 1793:with movements of the head. This suggests that 1104:, abelisaurids are included in the superfamily 3280:"Dinosaurian and mammalian predators compared" 2067:Then as now, Madagascar was an island, having 612:. DepĂ©ret referred these fossils to the genus 4288: 3500:Roach, Brian T.; Brinkman, Daniel T. (2007). 2199:might have competed with it closer to water. 2146:, lizards, snakes, seven distinct species of 1810:within the inner ear aid in balance, and the 8: 3878:. Society of Vertebrate Paleontology Memoir 3621:. Society of Vertebrate Paleontology Memoir 3007:. Society of Vertebrate Paleontology Memoir 2852:. Society of Vertebrate Paleontology Memoir 2810:. Society of Vertebrate Paleontology Memoir 2768:. Society of Vertebrate Paleontology Memoir 2313:. Society of Vertebrate Paleontology Memoir 2049:suggest that they were deposited during the 1065:was even stockier than that of its relative 2346:Bulletin de la SociĂ©tĂ© GĂ©ologique de France 2265:Bulletin de la SociĂ©tĂ© GĂ©ologique de France 2087:of the time was semi-arid, with pronounced 1757:, also known as CT scanning, of a complete 735:State University of New York at Stony Brook 430: 8494: 8225: 8142: 8117: 8001: 7997: 7956: 7945: 7932: 7843: 7811: 7627: 7591: 7550: 7419: 7415: 7300: 7296: 7285: 7185: 7174: 7161: 6952: 6942: 6858: 6770: 6658: 6605: 6515: 6468: 6464: 6414: 6379: 6368: 6355: 6286: 6195: 6103: 6056: 5925: 5900: 5896: 5824: 5820: 5809: 5694: 5683: 5670: 5598: 5417: 5408: 5237: 5052: 5003: 4999: 4912: 4908: 4874: 4863: 4850: 4649: 4532: 4450: 4439: 4426: 4372: 4363: 4311: 4295: 4281: 4273: 2704:. Princeton University Press. p. 85. 2119:may have roamed coastal environments like 1789:that helps to coordinate movements of the 1088:is classified as a member of the theropod 1069:, with a prominent crest on the knee. The 131: 31: 3828:from the Latest Cretaceous of Madagascar" 3802: 3765: 3473: 3360: 3350: 3071: 2953:from the Late Cretaceous of Madagascar". 1721:The recognition of pneumatic foramina in 803:, the authors judged the type dentary of 1108:, which is in turn a subdivision of the 3960:Annals of the Missouri Botanical Garden 3822:; O'Connor, Patrick M. (October 2016). 3787:from the Late Cretaceous of Madagascar" 3506:and other non-avian theropod dinosaurs" 2626:The Bulletin of Zoological Nomenclature 2588:The Bulletin of Zoological Nomenclature 2254: 1638:actively hunted their own kind or only 1003:had numerous cavities and excavations ( 3863:Rogers, Raymond R.; Krause, David W.; 3389:Rogers, Raymond R.; Krause, David W.; 2701:The Princeton Field Guide to Dinosaurs 1777:than to birds. One difference between 1750:Head and neck of specimen FMNH PR 2836 1423:individuals, there is no evidence for 995:over the hips. Although the cervical ( 851:specimen after a 2009 petition to the 1578:may have sacrificed speed for power. 1092:Abelisauridae, which is considered a 479:Cretaceous–Paleogene extinction event 7: 1989:in several bones, it was found that 1781:and other theropods was its smaller 1461:(twisting). In comparison to modern 1134:As with many dinosaur families, the 673:MusĂ©um National d'Histoire Naturelle 8714:Late Cretaceous dinosaurs of Africa 2021:(marked with a red dot on the map). 1666:Comparison between the air sacs of 1244: 1234: 589:Original material described in 1896 3835:Journal of Vertebrate Paleontology 3695:Journal of Vertebrate Paleontology 3146:Journal of Vertebrate Paleontology 3051:Proceedings of the Royal Society B 2955:Journal of Vertebrate Paleontology 2889:Journal of Vertebrate Paleontology 2537:Journal of Vertebrate Paleontology 2439:Journal of Vertebrate Paleontology 1674:Scientists have reconstructed the 827:Society of Vertebrate Paleontology 667:Numerous fragmentary remains from 25: 3044:; Wilson, JA; Conrad, JL (2007). 2083:than it is today. The prevailing 1602:in their diet. Numerous bones of 1561:. Scientists have suggested that 1150:shares a close relationship with 1030:. Uniquely, the cervical ribs of 944:bore seventeen teeth in both the 8478: 8472: 8465: 7917: 7910: 7903: 7897: 7146: 7139: 7134: 7127: 7121: 6340: 6333: 6326: 5655: 5648: 5641: 5634: 4836: 4830: 4237: 2433:; Forster, Catherine A. (1996). 2223: 2209: 1347: 1291: 1269: 624:. This name is derived from the 610:UniversitĂ© Claude Bernard Lyon 1 378: 152: 2239:Timeline of ceratosaur research 2127:represents the contemporaneous 2009:fossils have been found in the 1046:. Also like related dinosaurs, 1015:that interlocked for strength. 843:was replaced by the older name 495:, a name which is considered a 491:. This dinosaur is also called 2901:10.1080/02724634.2008.10010459 2459:10.1080/02724634.1996.10011350 2152:five or six species of mammals 1610:jaws, are of the same size as 1081:Classification and systematics 683:described a theropod dentary ( 645:, another poorly known taxon. 481:. The genus contains a single 1: 8734:Fossil taxa described in 1955 4058:10.1126/science.279.5358.1915 3791:Acta Palaeontologica Polonica 3767:10.1016/j.cretres.2020.104553 2685:10.1016/j.cretres.2016.09.001 2498:10.1126/science.280.5366.1048 2091:in temperature and rainfall. 1742:Brain and inner ear structure 1127:) forelimbs (convergent with 733:In 1993, scientists from the 3352:10.1371/journal.pone.0224734 3202:10.1016/j.annpal.2013.10.003 2975:10.1080/02724634.2012.622027 2916:PalĂ€ontologische Zeitschrift 2763:Majungasaurus crenatissimus 1797:and other abelisaurids like 1146:studies have indicated that 714: 687:.MAJ 1) with teeth from the 8633:Majungasaurus crenatissimus 8603:Majungasaurus crenatissimus 3874:Majungasaurus crenatissimus 3869:Majungasaurus crenatissimus 3826:Majungasaurus crenatissimus 3785:Majungasaurus crenatissimus 3617:Majungasaurus crenatissimus 3612:Majungasaurus crenatissimus 3003:Majungasaurus crenatissimus 2998:Majungasaurus crenatissimus 2951:Majungasaurus crenatissimus 2885:Majungasaurus crenatissimus 2848:Majungasaurus crenatissimus 2843:Majungasaurus crenatissimus 2806:Majungasaurus crenatissimus 2801:Majungasaurus crenatissimus 2754:Majungasaurus crenatissimus 2309:Majungasaurus crenatissimus 2304:Majungasaurus crenatissimus 1981:Research by Michael D'Emic 977:closely resembles those of 870:compared in size to a human 797:Majungasaurus crenatissimus 488:Majungasaurus crenatissimus 320:Majungasaurus crenatissimus 18:Majungasaurus crenatissimus 8755: 8739:Taxa named by RenĂ© Lavocat 3685:Hanna, Rebecca R. (2002). 1812:lateral semicircular canal 1386:Skull cast of FMNH PR 2100 739:University of Antananarivo 709: 616:, which at the time was a 8462: 8228: 8145: 8120: 8004: 8000: 7959: 7944: 7931: 7895: 7842: 7810: 7630: 7553: 7422: 7418: 7303: 7299: 7188: 7173: 7160: 7119: 6955: 6941: 6861: 6773: 6661: 6518: 6471: 6467: 6417: 6382: 6367: 6354: 6323: 6285: 6198: 6106: 6059: 5928: 5903: 5899: 5827: 5823: 5697: 5682: 5669: 5631: 5597: 5420: 5240: 5055: 5006: 5002: 4915: 4911: 4877: 4862: 4849: 4827: 4652: 4535: 4453: 4438: 4425: 4375: 4362: 4310: 4263:mount erected in 2006 at 3662:The Carnivorous Dinosaurs 3274:Van Valkenburgh, Blaire; 2928:10.1007/s12542-009-0044-2 2752:"Craniofacial anatomy of 2698:Paul, Gregory S. (2016). 2123:as well. The neighboring 1702:, and the lower back and 1472:Abelisaurids, especially 1409:intraspecific competition 1340: 1323: 1306: 1284: 1262: 1249: 1242: 948:of the upper jaw and the 506:Like other abelisaurids, 342: 335: 313: 306: 149:Scientific classification 147: 139: 130: 34: 3950:; O'Connor, Patrick M.; 3182:Annales de PalĂ©ontologie 2632:(1): 89–90. March 2011. 1987:lines of arrested growth 1966:(more specifically, the 1841:A 2007 report described 1100:. Along with the family 771:horn misidentified as a 8719:Dinosaurs of Madagascar 3504:Deinonychus antirrhopus 1955:Restoration of an adult 1911:) or along their tops ( 1888:were ruled out, and an 1219:A cladogram by Tortosa 362:Sues & Taquet, 1979 4265:Stony Brook University 4182:Curry Rogers, Kristina 3952:Rogers, Kristina Curry 3865:Curry Rogers, Kristina 3820:Curry Rogers, Kristina 3804:10.4202/app.00132.2014 3466:10.1098/rsbl.2006.0524 3391:Curry Rogers, Kristina 3219:; Horner, Celeste C.; 3064:10.1098/rspb.2004.2692 2661:Pycnonemosaurus nevesi 2465:on September 27, 2007. 2244:Evolution of dinosaurs 2022: 1956: 1751: 1671: 1551: 1449: 1387: 1251:Pourcieux abelisaurid 1216: 966: 871: 808:and thousands of shed 780: 664: 590: 142:Stony Brook University 8681:Paleobiology Database 8586:Paleobiology Database 6854:Carcharodontosaurinae 6766:Carcharodontosauridae 4255:May 20, 2006, at the 3721:on September 27, 2007 2638:10.21805/bzn.v68i1.a7 2600:10.21805/bzn.v66i3.a7 2578:Carrano, Matthew T.; 2179:sauropods, including 2004: 1954: 1749: 1665: 1654:and other theropods. 1543:Mounted skeletons of 1542: 1438: 1385: 1214: 1011:attachment sites and 964: 866: 759: 651: 588: 299:M. crenatissimus 8729:Maastrichtian genera 7847:Dubious coelurosaurs 6543:Metriacanthosaurinae 6511:Metriacanthosauridae 4793:Dubious neotheropods 4246:at Wikimedia Commons 3395:Majungatholus atopus 3102:Weishampel, David B. 2095:inhabited a coastal 1913:supraspinal ligament 1909:interspinal ligament 1227:in a new subfamily, 1121:carcharodontosaurids 1061:(lower leg bone) of 820:consisting of seven 801:Majungatholus atopus 789:Majungatholus atopus 762:Majungatholus atopus 728:Majungatholus atopus 718:(meaning "lizard"). 632:("notched") and the 581:Discovery and naming 563:, mainly preying on 470:, at the end of the 358:Majungatholus atopus 6867:Carcharodontosaurus 6654:Carcharodontosauria 4198:2001Natur.412..530C 4144:2001Natur.409..504S 4101:10.1038/nature03996 4093:2005Natur.437.1007M 4087:(7061): 1007–1011. 4050:1998Sci...279.1915F 4044:(5358): 1915–1919. 4007:1996Natur.382..532F 3923:2005Geo....33..297R 3758:2020CrRes.11504553G 3746:Cretaceous Research 3689:Allosaurus fragilis 3652:Rothschild, Bruce; 3572:10.1038/nature03716 3564:2005Natur.436..253O 3536:on August 10, 2007. 3419:10.1038/nature01532 3411:2003Natur.422..515R 3343:2019PLoSO..1424734D 3296:2002Pbio...28..527V 3237:2001Natur.409.1033R 3231:(6823): 1033–1037. 3194:2014AnPal.100...63T 3142:Carnotaurus sastrei 3058:(1546): 1325–1330. 2967:2012JVPal..32....1B 2677:2017CrRes..69...71G 2665:Cretaceous Research 2527:Sampson, Scott D.; 2490:1998Sci...280.1048S 2484:(5366): 1048–1051. 2451:1996JVPal..16..601S 2425:Sampson, Scott D.; 2396:1979Natur.279..633S 2231:Paleontology portal 2125:Berivotra Formation 2073:Indian subcontinent 1947:Ontogeny and growth 1808:semicircular canals 1755:Computed tomography 1463:mammalian predators 1439:Restoration of two 1378:Skull ornamentation 930:Computed tomography 924:, possibly made of 689:Maevarano Formation 545:Southern Hemisphere 526:. It also had more 8339:Archaeornithomimus 6843:Veterupristisaurus 6703:Veterupristisaurus 6559:Metriacanthosaurus 6499:Piatnitzkysauridae 5934:Eustreptospondylus 5844:Piatnitzkysauridae 5727:Chuandongocoelurus 4591:Notatesseraeraptor 2663:is the new king". 2099:cut by many sandy 2079:more southerly in 2031:Mahajanga Province 2023: 2011:Mahajanga Province 1957: 1882:Hypervitaminosis A 1785:, a region of the 1752: 1696:lungs and air sacs 1676:respiratory system 1672: 1658:Respiratory system 1552: 1450: 1388: 1217: 967: 872: 781: 669:Mahajanga Province 665: 591: 140:Mounted Skeleton, 8696: 8695: 8668:Open Tree of Life 8573:Open Tree of Life 8500:Taxon identifiers 8491: 8490: 8487: 8486: 8460: 8459: 8456: 8455: 8452: 8451: 8448: 8447: 8415: 8414: 8411: 8410: 8221:Macrocheiriformes 8138:Ornithomimosauria 8112:Maniraptoriformes 8105: 8104: 7951:Maniraptoromorpha 7938:Maniraptoromorpha 7927: 7926: 7893: 7892: 7889: 7888: 7885: 7884: 7838: 7837: 7834: 7833: 7816:Maniraptoromorpha 7806: 7805: 7802: 7801: 7798: 7797: 7794: 7793: 7790: 7789: 7538: 7537: 7403: 7402: 7357:Proceratosauridae 7309:Chingkankousaurus 7156: 7155: 7117: 7116: 7113: 7112: 7109: 7108: 7085: 7084: 6937: 6936: 6933: 6932: 6929: 6928: 6925: 6924: 6921: 6920: 6917: 6916: 6593: 6592: 6589: 6588: 6350: 6349: 6321: 6320: 6317: 6316: 6313: 6312: 6309: 6308: 6281: 6280: 6277: 6276: 6273: 6272: 6269: 6268: 6185: 6184: 6141:Vallibonavenatrix 6044: 6043: 6040: 6039: 5884: 5883: 5665: 5664: 5629: 5628: 5625: 5624: 5621: 5620: 5593: 5592: 5589: 5588: 5585: 5584: 5581: 5580: 5577: 5576: 5225: 5224: 5221: 5220: 4987: 4986: 4845: 4844: 4825: 4824: 4821: 4820: 4817: 4816: 4768: 4767: 4421: 4420: 4417: 4416: 4413: 4412: 4242:Media related to 4192:(6846): 530–534. 4138:(6819): 504–506. 4001:(6591): 532–534. 3818:D'Emic, Michael; 3671:978-0-253-34539-4 3558:(7048): 253–256. 3405:(6931): 515–518. 3215:Rayfield, Emily; 3138:Coria, Rodolfo A. 3123:978-0-520-24209-8 3110:OsmĂłlska, Halszka 2711:978-1-78684-190-2 2390:(5714): 633–635. 2025:All specimens of 1823:than in its more 1670:and a modern bird 1634:It is unknown if 1425:sexual dimorphism 1369: 1368: 1360: 1359: 1123:), very reduced ( 1098:Linnaean taxonomy 993:center of gravity 822:scientific papers 760:Type specimen of 618:wastebasket taxon 468:million years ago 369: 368: 363: 354: 287: 16:(Redirected from 8746: 8689: 8688: 8676: 8675: 8663: 8662: 8650: 8649: 8637: 8636: 8635: 8622: 8621: 8620: 8594: 8593: 8581: 8580: 8568: 8567: 8555: 8554: 8542: 8541: 8540: 8527: 8526: 8525: 8495: 8482: 8476: 8469: 8325:Aepyornithomimus 8226: 8143: 8118: 8082:Sinocalliopteryx 8002: 7998: 7992:Neocoelurosauria 7957: 7946: 7933: 7921: 7914: 7907: 7901: 7844: 7812: 7781:Zhuchengtyrannus 7697:Thanatotheristes 7628: 7592: 7559:Appalachiosaurus 7551: 7420: 7416: 7410:Pantyrannosauria 7301: 7297: 7291:Tyrannosauroidea 7286: 7261:Xinjiangovenator 7186: 7175: 7162: 7150: 7143: 7138: 7131: 7125: 6953: 6943: 6859: 6779:Acrocanthosaurus 6771: 6659: 6606: 6532:Yangchuanosaurus 6516: 6469: 6465: 6415: 6396:Lourinhanosaurus 6380: 6369: 6356: 6344: 6337: 6330: 6287: 6253:Sigilmassasaurus 6196: 6153:Ceratosuchopsini 6104: 6057: 6002:Dubreuillosaurus 5926: 5909:Streptospondylus 5901: 5897: 5867:Piatnitzkysaurus 5825: 5821: 5810: 5695: 5684: 5671: 5659: 5652: 5645: 5638: 5599: 5434:Ekrixinatosaurus 5418: 5409: 5238: 5053: 5004: 5000: 4913: 4909: 4875: 4864: 4851: 4840: 4834: 4744:Procompsognathus 4650: 4533: 4451: 4440: 4427: 4373: 4364: 4357: 4356: 4312: 4297: 4290: 4283: 4274: 4241: 4226: 4225: 4206:10.1038/35087566 4178: 4172: 4171: 4152:10.1038/35054046 4127: 4121: 4120: 4076: 4070: 4069: 4033: 4027: 4026: 4015:10.1038/382532a0 3990: 3984: 3983: 3948:Krause, David W. 3944: 3935: 3934: 3931:10.1130/G21036.1 3906: 3900: 3899: 3860: 3843: 3842: 3832: 3815: 3809: 3808: 3806: 3778: 3772: 3771: 3769: 3737: 3731: 3730: 3728: 3726: 3717:. Archived from 3682: 3676: 3675: 3654:Tanke, Darren H. 3649: 3643: 3642: 3607: 3592: 3591: 3547: 3538: 3537: 3532:. Archived from 3497: 3488: 3487: 3477: 3445: 3439: 3438: 3386: 3375: 3374: 3364: 3354: 3337:(11): e0224734. 3322: 3316: 3315: 3276:Molnar, Ralph E. 3271: 3265: 3264: 3245:10.1038/35059070 3217:Norman, David B. 3212: 3206: 3205: 3176: 3170: 3169: 3134: 3128: 3127: 3097: 3086: 3085: 3075: 3038: 3029: 3028: 2993: 2987: 2986: 2946: 2940: 2939: 2911: 2905: 2904: 2880: 2874: 2873: 2838: 2832: 2831: 2796: 2790: 2789: 2760: 2747: 2724: 2723: 2695: 2689: 2688: 2656: 2650: 2649: 2618: 2612: 2611: 2580:Krause, David W. 2575: 2569: 2568: 2529:Krause, David W. 2524: 2518: 2517: 2473: 2467: 2466: 2461:. Archived from 2427:Krause, David W. 2422: 2416: 2415: 2404:10.1038/279633a0 2379: 2373: 2372: 2360: 2354: 2353: 2341: 2335: 2334: 2300:Krause, David W. 2296: 2273: 2272: 2259: 2233: 2228: 2227: 2226: 2219: 2217:Dinosaurs portal 2214: 2213: 2212: 2169:, the noasaurid 2041:, evidence from 1845:in the bones of 1504:sported a large 1351: 1295: 1273: 1245: 1235: 1187:Ekrixinatosaurus 1162:in a clade with 965:Life restoration 773:pachycephalosaur 720:Hans-Dieter Sues 717: 712: 711: 657:M. crenatissimus 622:M. crenatissimus 524:pachycephalosaur 447: 444: 438: 435: 432: 428: 427: 424: 423: 420: 417: 414: 411: 408: 405: 402: 399: 396: 393: 390: 387: 384: 361: 352: 322: 318: 282: 275: 262: 249: 236: 157: 156: 135: 125: 55: 40:Temporal range: 32: 21: 8754: 8753: 8749: 8748: 8747: 8745: 8744: 8743: 8724:Maevarano fauna 8699: 8698: 8697: 8692: 8684: 8679: 8671: 8666: 8658: 8653: 8645: 8640: 8631: 8630: 8625: 8616: 8615: 8610: 8597: 8589: 8584: 8576: 8571: 8563: 8558: 8550: 8545: 8536: 8535: 8530: 8521: 8520: 8515: 8502: 8492: 8483: 8470: 8444: 8407: 8388:Sinornithomimus 8310: 8253: 8215: 8132: 8126:Compsognathidae 8114: 8101: 8089:Sinosauropteryx 8021:Compsognathidae 8015: 7994: 7985: 7953: 7940: 7923: 7922: 7908: 7881: 7876:Shanyangosaurus 7869:Phaedrolosaurus 7830: 7786: 7752: 7702: 7681:Daspletosaurini 7675: 7648: 7623:Tyrannosaurinae 7617: 7596:Albertosaurinae 7588: 7586:Tyrannosauridae 7579: 7547: 7545:Eutyrannosauria 7534: 7412: 7399: 7380:Proceratosaurus 7351: 7322: 7293: 7282: 7273: 7239:Richardoestesia 7223:Chilantaisaurus 7182: 7169: 7152: 7151: 7132: 7105: 7081: 7041:Australovenator 7019: 7013:Chilantaisaurus 6991:Phuwiangvenator 6913: 6878:Giganotosaurini 6872: 6848: 6760: 6723:Chilantaisaurus 6708: 6648: 6602: 6585: 6537: 6505: 6461: 6452: 6430:Monolophosaurus 6411: 6402: 6376: 6363: 6346: 6345: 6331: 6305: 6265: 6238: 6204:Camarillasaurus 6181: 6147: 6093: 6087:Ostafrikasaurus 6065:Camarillasaurus 6053: 6036: 5986:Afrovenatorinae 5980: 5939: 5915: 5893: 5880: 5838: 5817: 5815:Megalosauroidea 5806: 5797: 5792:Vectaerovenator 5785:Szechuanosaurus 5763:Monolophosaurus 5749:Kaijiangosaurus 5742:Cryolophosaurus 5703:Calamospondylus 5691: 5678: 5661: 5660: 5653: 5646: 5639: 5617: 5573: 5554:Pycnonemosaurus 5496: 5407: 5401:Majungasaurinae 5386: 5328:Majungasaurinae 5322: 5317:Xenotarsosaurus 5253:Dryptosauroides 5234: 5217: 5169: 5141:Elaphrosaurinae 5135: 5042: 5028:Ornithomimoides 4996: 4983: 4956: 4905: 4896: 4871: 4858: 4841: 4835: 4813: 4788: 4764: 4692: 4639: 4605:Shuangbaisaurus 4529: 4520: 4447: 4434: 4409: 4389: 4383:Avemetatarsalia 4368:Avemetatarsalia 4358: 4343:Avemetatarsalia 4317: 4316: 4306: 4301: 4257:Wayback Machine 4234: 4229: 4180: 4179: 4175: 4129: 4128: 4124: 4078: 4077: 4073: 4035: 4034: 4030: 3992: 3991: 3987: 3946: 3945: 3938: 3908: 3907: 3903: 3862: 3861: 3846: 3830: 3817: 3816: 3812: 3780: 3779: 3775: 3739: 3738: 3734: 3724: 3722: 3684: 3683: 3679: 3672: 3651: 3650: 3646: 3609: 3608: 3595: 3549: 3548: 3541: 3499: 3498: 3491: 3454:Biology Letters 3447: 3446: 3442: 3388: 3387: 3378: 3324: 3323: 3319: 3273: 3272: 3268: 3221:Horner, John R. 3214: 3213: 3209: 3178: 3177: 3173: 3136: 3135: 3131: 3124: 3099: 3098: 3089: 3042:Sereno, Paul C. 3040: 3039: 3032: 2995: 2994: 2990: 2948: 2947: 2943: 2913: 2912: 2908: 2882: 2881: 2877: 2840: 2839: 2835: 2798: 2797: 2793: 2758: 2749: 2748: 2727: 2712: 2697: 2696: 2692: 2658: 2657: 2653: 2620: 2619: 2615: 2577: 2576: 2572: 2531:, eds. (2007). 2526: 2525: 2521: 2475: 2474: 2470: 2424: 2423: 2419: 2381: 2380: 2376: 2362: 2361: 2357: 2343: 2342: 2338: 2298: 2297: 2276: 2261: 2260: 2256: 2252: 2229: 2224: 2222: 2215: 2210: 2208: 2205: 2190:Mahajangasuchus 2148:crocodylomorphs 2113:Paleocene Epoch 2043:biostratigraphy 2039:radiometrically 1999: 1949: 1921:physical trauma 1853:theropods like 1839: 1744: 1736:origin of birds 1660: 1644:Komodo monitors 1592: 1582:tooth marks on 1510:synovial joints 1433: 1380: 1375: 1370: 1361: 1239:Majungasaurinae 1229:Majungasaurinae 1083: 861: 799:was older than 747:David W. Krause 743:Late Cretaceous 724:Philippe Taquet 606:Betsiboka River 598:Charles DepĂ©ret 583: 549:biogeographical 445: 439: 436: 433: 381: 377: 360: 351: 331: 324: 316: 315: 302: 281: 273: 264:Majungasaurinae 260: 247: 234: 151: 126: 124: 123: 122: 121: 116: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 50: 49: 42:Late Cretaceous 38: 28: 23: 22: 15: 12: 11: 5: 8752: 8750: 8742: 8741: 8736: 8731: 8726: 8721: 8716: 8711: 8701: 8700: 8694: 8693: 8691: 8690: 8677: 8664: 8651: 8638: 8623: 8607: 8605: 8599: 8598: 8596: 8595: 8582: 8569: 8556: 8543: 8528: 8512: 8510: 8504: 8503: 8498: 8489: 8488: 8485: 8484: 8464: 8463: 8461: 8458: 8457: 8454: 8453: 8450: 8449: 8446: 8445: 8443: 8442: 8441: 8440: 8434: 8425: 8423: 8417: 8416: 8413: 8412: 8409: 8408: 8406: 8405: 8398: 8391: 8384: 8377: 8370: 8363: 8356: 8353:Dromiceiomimus 8349: 8342: 8335: 8328: 8320: 8318: 8316:Ornithomimidae 8312: 8311: 8309: 8308: 8301: 8297:Paraxenisaurus 8293: 8285: 8278: 8271: 8263: 8261: 8259:Deinocheiridae 8255: 8254: 8252: 8251: 8248:Shenzhousaurus 8244: 8237: 8229: 8223: 8217: 8216: 8214: 8213: 8206: 8198: 8191: 8184: 8177: 8170: 8162: 8154: 8146: 8140: 8134: 8133: 8131: 8130: 8121: 8115: 8110: 8107: 8106: 8103: 8102: 8100: 8099: 8092: 8085: 8078: 8070: 8063: 8056: 8049: 8045:Beipiaognathus 8041: 8034: 8025: 8023: 8017: 8016: 8014: 8013: 8005: 7995: 7990: 7987: 7986: 7984: 7983: 7976: 7969: 7960: 7954: 7949: 7942: 7941: 7936: 7929: 7928: 7925: 7924: 7916: 7902: 7896: 7894: 7891: 7890: 7887: 7886: 7883: 7882: 7880: 7879: 7872: 7865: 7858: 7850: 7848: 7840: 7839: 7836: 7835: 7832: 7831: 7829: 7828: 7827: 7826: 7820: 7818: 7808: 7807: 7804: 7803: 7800: 7799: 7796: 7795: 7792: 7791: 7788: 7787: 7785: 7784: 7777: 7770: 7762: 7760: 7758:Tyrannosaurini 7754: 7753: 7751: 7750: 7743: 7736: 7728: 7721: 7717:Bistahieversor 7712: 7710: 7708:Teratophoneini 7704: 7703: 7701: 7700: 7693: 7690:Daspletosaurus 7685: 7683: 7677: 7676: 7674: 7673: 7670:Qianzhousaurus 7666: 7658: 7656: 7650: 7649: 7647: 7646: 7639: 7631: 7625: 7619: 7618: 7616: 7615: 7608: 7600: 7598: 7589: 7584: 7581: 7580: 7578: 7577: 7570: 7566:Bistahieversor 7562: 7554: 7548: 7543: 7540: 7539: 7536: 7535: 7533: 7532: 7524: 7517: 7510: 7502: 7495: 7488: 7481: 7474: 7467: 7460: 7453: 7446: 7438: 7431: 7423: 7413: 7408: 7405: 7404: 7401: 7400: 7398: 7397: 7390: 7383: 7376: 7369: 7361: 7359: 7353: 7352: 7350: 7349: 7341: 7333: 7331: 7324: 7323: 7321: 7320: 7312: 7304: 7294: 7289: 7283: 7280:Tyrannoraptora 7278: 7275: 7274: 7272: 7271: 7264: 7257: 7250: 7242: 7235: 7227: 7219: 7212: 7204: 7197: 7189: 7183: 7178: 7171: 7170: 7165: 7158: 7157: 7154: 7153: 7145: 7126: 7120: 7118: 7115: 7114: 7111: 7110: 7107: 7106: 7104: 7103: 7102: 7101: 7095: 7093: 7087: 7086: 7083: 7082: 7080: 7079: 7072: 7065: 7058: 7051: 7044: 7037: 7029: 7027: 7025:Megaraptoridae 7021: 7020: 7018: 7017: 7009: 7001: 6994: 6987: 6980: 6972: 6964: 6956: 6950: 6939: 6938: 6935: 6934: 6931: 6930: 6927: 6926: 6923: 6922: 6919: 6918: 6915: 6914: 6912: 6911: 6904: 6897: 6890: 6887:Giganotosaurus 6882: 6880: 6874: 6873: 6871: 6870: 6862: 6856: 6850: 6849: 6847: 6846: 6839: 6832: 6825: 6818: 6811: 6804: 6797: 6790: 6782: 6774: 6768: 6762: 6761: 6759: 6758: 6750: 6742: 6735: 6727: 6718: 6716: 6714:Neovenatoridae 6710: 6709: 6707: 6706: 6699: 6696:Ulughbegsaurus 6692: 6684: 6677: 6670: 6662: 6656: 6650: 6649: 6647: 6646: 6638: 6630: 6622: 6614: 6612: 6603: 6598: 6595: 6594: 6591: 6590: 6587: 6586: 6584: 6583: 6576: 6569: 6562: 6555: 6547: 6545: 6539: 6538: 6536: 6535: 6528: 6524:Xuanhanosaurus 6519: 6513: 6507: 6506: 6504: 6503: 6495: 6491:Xuanhanosaurus 6487: 6480: 6477:Asfaltovenator 6472: 6462: 6457: 6454: 6453: 6451: 6450: 6442: 6438:Megalosauridae 6434: 6426: 6418: 6412: 6407: 6404: 6403: 6401: 6400: 6392: 6383: 6377: 6372: 6365: 6364: 6359: 6352: 6351: 6348: 6347: 6339: 6325: 6324: 6322: 6319: 6318: 6315: 6314: 6311: 6310: 6307: 6306: 6304: 6303: 6302: 6301: 6295: 6293: 6283: 6282: 6279: 6278: 6275: 6274: 6271: 6270: 6267: 6266: 6264: 6263: 6256: 6248: 6246: 6240: 6239: 6237: 6236: 6229: 6222: 6215: 6212:Ichthyovenator 6208: 6199: 6193: 6187: 6186: 6183: 6182: 6180: 6179: 6172: 6165: 6157: 6155: 6149: 6148: 6146: 6145: 6137: 6129: 6122: 6115: 6107: 6101: 6095: 6094: 6092: 6091: 6083: 6076: 6073:Cristatusaurus 6069: 6060: 6054: 6049: 6046: 6045: 6042: 6041: 6038: 6037: 6035: 6034: 6030:Poekilopleuron 6026: 6023:Piveteausaurus 6019: 6012: 6005: 5998: 5990: 5988: 5982: 5981: 5979: 5978: 5971: 5964: 5957: 5949: 5947: 5945:Megalosaurinae 5941: 5940: 5938: 5937: 5929: 5923: 5921:Megalosauridae 5917: 5916: 5914: 5913: 5904: 5894: 5889: 5886: 5885: 5882: 5881: 5879: 5878: 5874:Xuanhanosaurus 5870: 5863: 5856: 5848: 5846: 5840: 5839: 5837: 5836: 5833:Yunyangosaurus 5828: 5818: 5813: 5807: 5802: 5799: 5798: 5796: 5795: 5788: 5781: 5773: 5770:Pandoravenator 5766: 5759: 5756:Kayentavenator 5752: 5745: 5738: 5730: 5723: 5715: 5707: 5698: 5692: 5687: 5680: 5679: 5674: 5667: 5666: 5663: 5662: 5654: 5647: 5640: 5633: 5632: 5630: 5627: 5626: 5623: 5622: 5619: 5618: 5616: 5615: 5614: 5613: 5607: 5605: 5595: 5594: 5591: 5590: 5587: 5586: 5583: 5582: 5579: 5578: 5575: 5574: 5572: 5571: 5564: 5557: 5550: 5543: 5535: 5528: 5521: 5514: 5506: 5504: 5498: 5497: 5495: 5494: 5487: 5484:Skorpiovenator 5480: 5472: 5465: 5458: 5451: 5444: 5437: 5430: 5421: 5415: 5406: 5405: 5396: 5394: 5388: 5387: 5385: 5384: 5377: 5369: 5362: 5355: 5348: 5340: 5332: 5330: 5324: 5323: 5321: 5320: 5313: 5306: 5299: 5296:Spectrovenator 5292: 5285: 5278: 5271: 5264: 5257: 5249: 5241: 5235: 5230: 5227: 5226: 5223: 5222: 5219: 5218: 5216: 5215: 5208: 5201: 5194: 5187: 5179: 5177: 5171: 5170: 5168: 5167: 5160: 5153: 5145: 5143: 5137: 5136: 5134: 5133: 5130:Spinostropheus 5126: 5119: 5112: 5104: 5096: 5088: 5080: 5073: 5065: 5056: 5050: 5044: 5043: 5041: 5040: 5032: 5024: 5016: 5007: 4997: 4994:Abelisauroidea 4992: 4989: 4988: 4985: 4984: 4982: 4981: 4974: 4966: 4964: 4962:Ceratosauridae 4958: 4957: 4955: 4954: 4951:Saltriovenator 4947: 4939: 4932: 4928:Fosterovenator 4924: 4916: 4906: 4901: 4898: 4897: 4895: 4894: 4890:Lophostropheus 4886: 4878: 4872: 4867: 4860: 4859: 4854: 4847: 4846: 4843: 4842: 4829: 4828: 4826: 4823: 4822: 4819: 4818: 4815: 4814: 4812: 4811: 4804: 4796: 4794: 4790: 4789: 4787: 4786: 4785: 4784: 4778: 4776: 4770: 4769: 4766: 4765: 4763: 4762: 4755: 4751:Pterospondylus 4747: 4740: 4732: 4724: 4721:Lucianovenator 4717: 4710: 4702: 4700: 4694: 4693: 4691: 4690: 4683: 4676: 4669: 4662: 4653: 4647: 4645:Coelophysoidea 4641: 4640: 4638: 4637: 4630: 4622: 4615: 4608: 4601: 4594: 4587: 4584:Lophostropheus 4580: 4573: 4566: 4559: 4552: 4544: 4536: 4530: 4525: 4522: 4521: 4519: 4518: 4510: 4502: 4494: 4486: 4483:Erythrovenator 4479: 4471: 4463: 4454: 4448: 4443: 4436: 4435: 4430: 4423: 4422: 4419: 4418: 4415: 4414: 4411: 4410: 4408: 4407: 4406: 4405: 4399: 4397: 4391: 4390: 4388: 4387: 4386: 4385: 4376: 4370: 4360: 4359: 4355: 4354: 4345: 4336: 4330: 4324: 4315: 4308: 4307: 4302: 4300: 4299: 4292: 4285: 4277: 4269: 4268: 4247: 4233: 4232:External links 4230: 4228: 4227: 4173: 4122: 4071: 4028: 3985: 3966:(2): 178–208. 3936: 3917:(4): 297–300. 3901: 3844: 3810: 3773: 3732: 3677: 3670: 3644: 3593: 3539: 3516:(1): 103–138. 3489: 3460:(4): 611–614. 3440: 3376: 3317: 3290:(4): 527–543. 3266: 3207: 3171: 3152:(2): 460–465. 3129: 3122: 3114:The Dinosauria 3087: 3030: 2988: 2941: 2922:(2): 293–300. 2906: 2875: 2833: 2791: 2725: 2710: 2690: 2651: 2613: 2594:(3): 261–264. 2570: 2519: 2468: 2445:(4): 601–605. 2417: 2374: 2355: 2336: 2274: 2253: 2251: 2248: 2247: 2246: 2241: 2235: 2234: 2220: 2204: 2201: 2196:Trematochampsa 2062:became extinct 2047:paleomagnetism 1998: 1995: 1948: 1945: 1859:tyrannosaurids 1838: 1835: 1771:coelurosaurian 1743: 1740: 1716:carbon dioxide 1659: 1656: 1591: 1588: 1432: 1429: 1379: 1376: 1374: 1371: 1367: 1366: 1363: 1362: 1358: 1357: 1354: 1353: 1339: 1336: 1335: 1332: 1331: 1322: 1319: 1318: 1315: 1314: 1305: 1302: 1301: 1298: 1297: 1283: 1280: 1279: 1276: 1275: 1261: 1258: 1257: 1254: 1253: 1248: 1243: 1241: 1233: 1129:tyrannosaurids 1106:Abelisauroidea 1082: 1079: 860: 857: 595:paleontologist 582: 579: 551:implications. 497:junior synonym 466:from 70 to 66 462:that lived in 367: 366: 365: 364: 355: 340: 339: 333: 332: 325: 311: 310: 304: 303: 295: 293: 289: 288: 271: 267: 266: 258: 254: 253: 245: 241: 240: 238:Abelisauroidea 232: 228: 227: 222: 215: 214: 209: 202: 201: 196: 189: 188: 183: 179: 178: 173: 169: 168: 163: 159: 158: 145: 144: 137: 136: 128: 127: 119: 118: 117: 112: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 56: 39: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 8751: 8740: 8737: 8735: 8732: 8730: 8727: 8725: 8722: 8720: 8717: 8715: 8712: 8710: 8707: 8706: 8704: 8687: 8682: 8678: 8674: 8669: 8665: 8661: 8656: 8652: 8648: 8643: 8639: 8634: 8628: 8624: 8619: 8613: 8609: 8608: 8606: 8604: 8600: 8592: 8587: 8583: 8579: 8574: 8570: 8566: 8561: 8557: 8553: 8548: 8544: 8539: 8538:Majungasaurus 8533: 8529: 8524: 8518: 8514: 8513: 8511: 8509: 8508:Majungasaurus 8505: 8501: 8496: 8481: 8477: 8475: 8468: 8439: 8435: 8433: 8429: 8428: 8427: 8426: 8424: 8422: 8418: 8404: 8403: 8399: 8397: 8396: 8395:Struthiomimus 8392: 8390: 8389: 8385: 8383: 8382: 8378: 8376: 8375: 8371: 8369: 8368: 8364: 8362: 8361: 8357: 8355: 8354: 8350: 8348: 8347: 8346:"Coelosaurus" 8343: 8341: 8340: 8336: 8334: 8333: 8329: 8327: 8326: 8322: 8321: 8319: 8317: 8313: 8307: 8306: 8302: 8299: 8298: 8294: 8291: 8290: 8286: 8284: 8283: 8279: 8277: 8276: 8272: 8270: 8269: 8265: 8264: 8262: 8260: 8256: 8250: 8249: 8245: 8243: 8242: 8241:Pelecanimimus 8238: 8236: 8235: 8231: 8230: 8227: 8224: 8222: 8218: 8212: 8211: 8207: 8204: 8203: 8202:Thecocoelurus 8199: 8197: 8196: 8192: 8190: 8189: 8185: 8183: 8182: 8181:Kinnareemimus 8178: 8176: 8175: 8171: 8168: 8167: 8163: 8160: 8159: 8155: 8153: 8152: 8148: 8147: 8144: 8141: 8139: 8135: 8128: 8127: 8123: 8122: 8119: 8116: 8113: 8108: 8098: 8097: 8093: 8091: 8090: 8086: 8084: 8083: 8079: 8076: 8075: 8071: 8069: 8068: 8064: 8062: 8061: 8060:Huaxiagnathus 8057: 8055: 8054: 8053:Compsognathus 8050: 8047: 8046: 8042: 8040: 8039: 8035: 8032: 8031: 8027: 8026: 8024: 8022: 8018: 8012: 8011: 8007: 8006: 8003: 7999: 7996: 7993: 7988: 7982: 7981: 7980:Ornitholestes 7977: 7975: 7974: 7970: 7967: 7966: 7962: 7961: 7958: 7955: 7952: 7947: 7943: 7939: 7934: 7930: 7920: 7915: 7913: 7906: 7900: 7878: 7877: 7873: 7871: 7870: 7866: 7864: 7863: 7859: 7857: 7856: 7852: 7851: 7849: 7845: 7841: 7824: 7823: 7822: 7821: 7819: 7817: 7813: 7809: 7783: 7782: 7778: 7776: 7775: 7774:Tyrannosaurus 7771: 7769: 7768: 7764: 7763: 7761: 7759: 7755: 7749: 7748: 7747:Teratophoneus 7744: 7742: 7741: 7737: 7734: 7733: 7729: 7727: 7726: 7722: 7719: 7718: 7714: 7713: 7711: 7709: 7705: 7699: 7698: 7694: 7692: 7691: 7687: 7686: 7684: 7682: 7678: 7672: 7671: 7667: 7665: 7664: 7660: 7659: 7657: 7655: 7651: 7645: 7644: 7640: 7638: 7637: 7633: 7632: 7629: 7626: 7624: 7620: 7614: 7613: 7609: 7607: 7606: 7605:Albertosaurus 7602: 7601: 7599: 7597: 7593: 7590: 7587: 7582: 7576: 7575: 7571: 7568: 7567: 7563: 7561: 7560: 7556: 7555: 7552: 7549: 7546: 7541: 7530: 7529: 7525: 7523: 7522: 7521:Xiongguanlong 7518: 7516: 7515: 7511: 7508: 7507: 7503: 7501: 7500: 7499:Suskityrannus 7496: 7494: 7493: 7492:Stokesosaurus 7489: 7487: 7486: 7485:Santanaraptor 7482: 7480: 7479: 7475: 7473: 7472: 7468: 7466: 7465: 7461: 7459: 7458: 7454: 7452: 7451: 7447: 7444: 7443: 7439: 7437: 7436: 7432: 7430: 7429: 7428:Alectrosaurus 7425: 7424: 7421: 7417: 7414: 7411: 7406: 7396: 7395: 7391: 7389: 7388: 7384: 7382: 7381: 7377: 7375: 7374: 7370: 7368: 7367: 7363: 7362: 7360: 7358: 7354: 7347: 7346: 7345:Tanycolagreus 7342: 7340: 7339: 7335: 7334: 7332: 7329: 7325: 7318: 7317: 7313: 7311: 7310: 7306: 7305: 7302: 7298: 7295: 7292: 7287: 7284: 7281: 7276: 7270: 7269: 7265: 7263: 7262: 7258: 7256: 7255: 7251: 7248: 7247: 7243: 7241: 7240: 7236: 7233: 7232: 7228: 7225: 7224: 7220: 7218: 7217: 7213: 7210: 7209: 7208:Bahariasaurus 7205: 7203: 7202: 7198: 7196: 7195: 7191: 7190: 7187: 7184: 7181: 7180:Coelurosauria 7176: 7172: 7168: 7167:Coelurosauria 7163: 7159: 7149: 7144: 7142: 7137: 7130: 7124: 7099: 7098: 7097: 7096: 7094: 7092: 7091:Coelurosauria 7088: 7078: 7077: 7073: 7071: 7070: 7066: 7064: 7063: 7059: 7057: 7056: 7052: 7050: 7049: 7045: 7043: 7042: 7038: 7036: 7035: 7031: 7030: 7028: 7026: 7022: 7016: 7014: 7010: 7008: 7006: 7002: 7000: 6999: 6995: 6993: 6992: 6988: 6986: 6985: 6981: 6978: 6977: 6973: 6970: 6969: 6968:Bahariasaurus 6965: 6963: 6962: 6958: 6957: 6954: 6951: 6948: 6944: 6940: 6910: 6909: 6905: 6903: 6902: 6898: 6896: 6895: 6891: 6889: 6888: 6884: 6883: 6881: 6879: 6875: 6869: 6868: 6864: 6863: 6860: 6857: 6855: 6851: 6845: 6844: 6840: 6838: 6837: 6833: 6831: 6830: 6826: 6824: 6823: 6819: 6817: 6816: 6812: 6810: 6809: 6808:Kelmayisaurus 6805: 6803: 6802: 6798: 6796: 6795: 6791: 6788: 6787: 6783: 6781: 6780: 6776: 6775: 6772: 6769: 6767: 6763: 6756: 6755: 6751: 6748: 6747: 6743: 6741: 6740: 6736: 6733: 6732: 6728: 6725: 6724: 6720: 6719: 6717: 6715: 6711: 6705: 6704: 6700: 6698: 6697: 6693: 6690: 6689: 6685: 6683: 6682: 6678: 6676: 6675: 6671: 6669: 6668: 6664: 6663: 6660: 6657: 6655: 6651: 6644: 6643: 6642:Saurophaganax 6639: 6636: 6635: 6631: 6628: 6627: 6623: 6621: 6620: 6616: 6615: 6613: 6611: 6607: 6604: 6601: 6596: 6582: 6581: 6577: 6575: 6574: 6573:Siamotyrannus 6570: 6568: 6567: 6563: 6561: 6560: 6556: 6554: 6553: 6549: 6548: 6546: 6544: 6540: 6534: 6533: 6529: 6526: 6525: 6521: 6520: 6517: 6514: 6512: 6508: 6501: 6500: 6496: 6493: 6492: 6488: 6486: 6485: 6481: 6479: 6478: 6474: 6473: 6470: 6466: 6463: 6460: 6459:Allosauroidea 6455: 6448: 6447: 6446:Spinosauridae 6443: 6440: 6439: 6435: 6432: 6431: 6427: 6425: 6424: 6420: 6419: 6416: 6413: 6410: 6405: 6398: 6397: 6393: 6390: 6389: 6385: 6384: 6381: 6378: 6375: 6370: 6366: 6362: 6357: 6353: 6343: 6338: 6336: 6329: 6299: 6298: 6297: 6296: 6294: 6292: 6288: 6284: 6262: 6261: 6257: 6255: 6254: 6250: 6249: 6247: 6245: 6241: 6235: 6234: 6230: 6228: 6227: 6223: 6221: 6220: 6216: 6214: 6213: 6209: 6206: 6205: 6201: 6200: 6197: 6194: 6192: 6191:Spinosaurinae 6188: 6178: 6177: 6173: 6171: 6170: 6169:Riparovenator 6166: 6164: 6163: 6162:Ceratosuchops 6159: 6158: 6156: 6154: 6150: 6143: 6142: 6138: 6135: 6134: 6130: 6128: 6127: 6126:Riojavenatrix 6123: 6121: 6120: 6116: 6114: 6113: 6109: 6108: 6105: 6102: 6100: 6099:Baryonychinae 6096: 6089: 6088: 6084: 6082: 6081: 6077: 6075: 6074: 6070: 6067: 6066: 6062: 6061: 6058: 6055: 6052: 6051:Spinosauridae 6047: 6032: 6031: 6027: 6025: 6024: 6020: 6018: 6017: 6013: 6011: 6010: 6006: 6004: 6003: 5999: 5997: 5996: 5992: 5991: 5989: 5987: 5983: 5977: 5976: 5975:Wiehenvenator 5972: 5970: 5969: 5965: 5963: 5962: 5958: 5956: 5955: 5951: 5950: 5948: 5946: 5942: 5936: 5935: 5931: 5930: 5927: 5924: 5922: 5918: 5911: 5910: 5906: 5905: 5902: 5898: 5895: 5892: 5887: 5876: 5875: 5871: 5869: 5868: 5864: 5862: 5861: 5857: 5855: 5854: 5850: 5849: 5847: 5845: 5841: 5835: 5834: 5830: 5829: 5826: 5822: 5819: 5816: 5811: 5808: 5805: 5800: 5794: 5793: 5789: 5787: 5786: 5782: 5779: 5778: 5774: 5772: 5771: 5767: 5765: 5764: 5760: 5758: 5757: 5753: 5751: 5750: 5746: 5744: 5743: 5739: 5736: 5735: 5731: 5729: 5728: 5724: 5721: 5720: 5716: 5713: 5712: 5711:Chienkosaurus 5708: 5705: 5704: 5700: 5699: 5696: 5693: 5690: 5685: 5681: 5677: 5672: 5668: 5658: 5651: 5644: 5637: 5611: 5610: 5609: 5608: 5606: 5604: 5600: 5596: 5570: 5569: 5565: 5563: 5562: 5558: 5556: 5555: 5551: 5549: 5548: 5544: 5541: 5540: 5536: 5534: 5533: 5529: 5527: 5526: 5522: 5520: 5519: 5515: 5513: 5512: 5508: 5507: 5505: 5503: 5502:Furileusauria 5499: 5493: 5492: 5488: 5486: 5485: 5481: 5478: 5477: 5476:Rahiolisaurus 5473: 5471: 5470: 5466: 5464: 5463: 5459: 5457: 5456: 5452: 5450: 5449: 5445: 5443: 5442: 5438: 5436: 5435: 5431: 5428: 5427: 5423: 5422: 5419: 5416: 5414: 5410: 5403: 5402: 5398: 5397: 5395: 5393: 5392:Carnotaurinae 5389: 5383: 5382: 5378: 5375: 5374: 5373:Rahiolisaurus 5370: 5368: 5367: 5366:Majungasaurus 5363: 5361: 5360: 5356: 5354: 5353: 5349: 5346: 5345: 5341: 5339: 5338: 5334: 5333: 5331: 5329: 5325: 5319: 5318: 5314: 5312: 5311: 5307: 5305: 5304: 5303:Tarascosaurus 5300: 5298: 5297: 5293: 5291: 5290: 5286: 5284: 5283: 5279: 5277: 5276: 5272: 5270: 5269: 5265: 5263: 5262: 5261:Eoabelisaurus 5258: 5255: 5254: 5250: 5248: 5247: 5246:Chenanisaurus 5243: 5242: 5239: 5236: 5233: 5232:Abelisauridae 5228: 5214: 5213: 5209: 5207: 5206: 5202: 5200: 5199: 5195: 5193: 5192: 5191:Masiakasaurus 5188: 5186: 5185: 5181: 5180: 5178: 5176: 5172: 5166: 5165: 5161: 5159: 5158: 5157:Huinculsaurus 5154: 5152: 5151: 5150:Elaphrosaurus 5147: 5146: 5144: 5142: 5138: 5132: 5131: 5127: 5125: 5124: 5120: 5118: 5117: 5113: 5110: 5109: 5105: 5102: 5101: 5097: 5094: 5093: 5089: 5086: 5085: 5081: 5079: 5078: 5074: 5071: 5070: 5069:Austrocheirus 5066: 5063: 5062: 5058: 5057: 5054: 5051: 5049: 5045: 5038: 5037: 5033: 5030: 5029: 5025: 5022: 5021: 5017: 5014: 5013: 5009: 5008: 5005: 5001: 4998: 4995: 4990: 4980: 4979: 4975: 4973: 4972: 4968: 4967: 4965: 4963: 4959: 4953: 4952: 4948: 4945: 4944: 4940: 4938: 4937: 4933: 4930: 4929: 4925: 4923: 4922: 4921:Berberosaurus 4918: 4917: 4914: 4910: 4907: 4904: 4899: 4892: 4891: 4887: 4885: 4884: 4880: 4879: 4876: 4873: 4870: 4865: 4861: 4857: 4852: 4848: 4839: 4833: 4810: 4809: 4808:Halticosaurus 4805: 4803: 4802: 4801:Teinurosaurus 4798: 4797: 4795: 4791: 4782: 4781: 4780: 4779: 4777: 4775: 4771: 4761: 4760: 4756: 4753: 4752: 4748: 4746: 4745: 4741: 4738: 4737: 4733: 4730: 4729: 4728:Megapnosaurus 4725: 4723: 4722: 4718: 4716: 4715: 4711: 4709: 4708: 4704: 4703: 4701: 4699: 4698:Coelophysidae 4695: 4689: 4688: 4687:Powellvenator 4684: 4682: 4681: 4677: 4675: 4674: 4670: 4668: 4667: 4663: 4660: 4659: 4655: 4654: 4651: 4648: 4646: 4642: 4636: 4635: 4631: 4628: 4627: 4623: 4621: 4620: 4616: 4614: 4613: 4609: 4607: 4606: 4602: 4600: 4599: 4595: 4593: 4592: 4588: 4586: 4585: 4581: 4579: 4578: 4577:Liliensternus 4574: 4572: 4571: 4567: 4565: 4564: 4560: 4558: 4557: 4553: 4550: 4549: 4548:Dolichosuchus 4545: 4543: 4542: 4541:Dilophosaurus 4538: 4537: 4534: 4531: 4528: 4523: 4516: 4515: 4514:Herrerasauria 4511: 4508: 4507: 4503: 4500: 4499: 4495: 4492: 4491: 4487: 4485: 4484: 4480: 4477: 4476: 4472: 4469: 4468: 4467:Daemonosaurus 4464: 4461: 4460: 4456: 4455: 4452: 4449: 4446: 4441: 4437: 4433: 4428: 4424: 4403: 4402: 4401: 4400: 4398: 4396: 4392: 4384: 4380: 4379: 4378: 4377: 4374: 4371: 4369: 4365: 4361: 4353: 4349: 4346: 4344: 4340: 4337: 4335: 4331: 4329: 4325: 4323: 4319: 4318: 4313: 4309: 4305: 4298: 4293: 4291: 4286: 4284: 4279: 4278: 4275: 4271: 4266: 4262: 4261:Majungasaurus 4258: 4254: 4251: 4250:Press release 4248: 4245: 4244:Majungasaurus 4240: 4236: 4235: 4231: 4223: 4219: 4215: 4211: 4207: 4203: 4199: 4195: 4191: 4187: 4183: 4177: 4174: 4169: 4165: 4161: 4157: 4153: 4149: 4145: 4141: 4137: 4133: 4126: 4123: 4118: 4114: 4110: 4106: 4102: 4098: 4094: 4090: 4086: 4082: 4075: 4072: 4067: 4063: 4059: 4055: 4051: 4047: 4043: 4039: 4032: 4029: 4024: 4020: 4016: 4012: 4008: 4004: 4000: 3996: 3989: 3986: 3981: 3977: 3973: 3969: 3965: 3961: 3957: 3953: 3949: 3943: 3941: 3937: 3932: 3928: 3924: 3920: 3916: 3912: 3905: 3902: 3897: 3893: 3889: 3885: 3881: 3877: 3873: 3870: 3866: 3859: 3857: 3855: 3853: 3851: 3849: 3845: 3840: 3836: 3829: 3827: 3821: 3814: 3811: 3805: 3800: 3796: 3792: 3788: 3786: 3777: 3774: 3768: 3763: 3759: 3755: 3751: 3747: 3743: 3736: 3733: 3720: 3716: 3712: 3708: 3704: 3700: 3696: 3692: 3690: 3681: 3678: 3673: 3667: 3663: 3659: 3658:Carpenter, K. 3655: 3648: 3645: 3640: 3636: 3632: 3628: 3624: 3620: 3616: 3613: 3606: 3604: 3602: 3600: 3598: 3594: 3589: 3585: 3581: 3577: 3573: 3569: 3565: 3561: 3557: 3553: 3546: 3544: 3540: 3535: 3531: 3527: 3523: 3519: 3515: 3511: 3507: 3505: 3496: 3494: 3490: 3485: 3481: 3476: 3471: 3467: 3463: 3459: 3455: 3451: 3444: 3441: 3436: 3432: 3428: 3424: 3420: 3416: 3412: 3408: 3404: 3400: 3396: 3392: 3385: 3383: 3381: 3377: 3372: 3368: 3363: 3358: 3353: 3348: 3344: 3340: 3336: 3332: 3328: 3321: 3318: 3313: 3309: 3305: 3301: 3297: 3293: 3289: 3285: 3281: 3277: 3270: 3267: 3262: 3258: 3254: 3250: 3246: 3242: 3238: 3234: 3230: 3226: 3222: 3218: 3211: 3208: 3203: 3199: 3195: 3191: 3187: 3183: 3175: 3172: 3167: 3163: 3159: 3155: 3151: 3147: 3143: 3139: 3133: 3130: 3125: 3119: 3115: 3111: 3107: 3106:Dodson, Peter 3103: 3096: 3094: 3092: 3088: 3083: 3079: 3074: 3069: 3065: 3061: 3057: 3053: 3052: 3047: 3043: 3037: 3035: 3031: 3026: 3022: 3018: 3014: 3010: 3006: 3002: 2999: 2992: 2989: 2984: 2980: 2976: 2972: 2968: 2964: 2960: 2956: 2952: 2945: 2942: 2937: 2933: 2929: 2925: 2921: 2917: 2910: 2907: 2902: 2898: 2894: 2890: 2886: 2879: 2876: 2871: 2867: 2863: 2859: 2855: 2851: 2847: 2844: 2837: 2834: 2829: 2825: 2821: 2817: 2813: 2809: 2805: 2802: 2795: 2792: 2787: 2783: 2779: 2775: 2771: 2767: 2764: 2757: 2755: 2746: 2744: 2742: 2740: 2738: 2736: 2734: 2732: 2730: 2726: 2721: 2717: 2713: 2707: 2703: 2702: 2694: 2691: 2686: 2682: 2678: 2674: 2670: 2666: 2662: 2655: 2652: 2647: 2643: 2639: 2635: 2631: 2627: 2623: 2617: 2614: 2609: 2605: 2601: 2597: 2593: 2589: 2585: 2581: 2574: 2571: 2566: 2562: 2558: 2554: 2550: 2546: 2542: 2538: 2534: 2530: 2523: 2520: 2515: 2511: 2507: 2503: 2499: 2495: 2491: 2487: 2483: 2479: 2472: 2469: 2464: 2460: 2456: 2452: 2448: 2444: 2440: 2436: 2432: 2431:Dodson, Peter 2428: 2421: 2418: 2413: 2409: 2405: 2401: 2397: 2393: 2389: 2385: 2378: 2375: 2370: 2367:(in French). 2366: 2359: 2356: 2351: 2348:(in French). 2347: 2340: 2337: 2332: 2328: 2324: 2320: 2316: 2312: 2308: 2305: 2301: 2295: 2293: 2291: 2289: 2287: 2285: 2283: 2281: 2279: 2275: 2270: 2267:(in French). 2266: 2258: 2255: 2249: 2245: 2242: 2240: 2237: 2236: 2232: 2221: 2218: 2207: 2202: 2200: 2198: 2197: 2192: 2191: 2186: 2185:Majungasaurus 2182: 2178: 2177:titanosaurian 2174: 2173: 2172:Masiakasaurus 2168: 2167: 2163: 2162:dromaeosaurid 2159: 2158: 2153: 2149: 2145: 2141: 2137: 2136:Majungasaurus 2132: 2131:environment. 2130: 2126: 2122: 2118: 2117:Majungasaurus 2114: 2109: 2105: 2102: 2098: 2094: 2093:Majungasaurus 2090: 2086: 2082: 2078: 2074: 2070: 2065: 2063: 2059: 2058:Majungasaurus 2055: 2052: 2051:Maastrichtian 2048: 2044: 2040: 2036: 2035:Majungasaurus 2032: 2028: 2027:Majungasaurus 2020: 2016: 2012: 2008: 2007:Majungasaurus 2003: 1996: 1994: 1992: 1991:Majungasaurus 1988: 1984: 1979: 1977: 1976:quadratojugal 1973: 1969: 1965: 1964:Majungasaurus 1961: 1960:Majungasaurus 1953: 1946: 1944: 1941: 1940:Majungasaurus 1937: 1936:Majungasaurus 1932: 1930: 1926: 1925:osteomyelitis 1922: 1916: 1914: 1910: 1906: 1905:Masiakasaurus 1902: 1901: 1898:specimens of 1895: 1891: 1887: 1883: 1879: 1875: 1871: 1866: 1864: 1860: 1856: 1852: 1848: 1847:Majungasaurus 1844: 1836: 1834: 1832: 1831: 1826: 1822: 1821:Majungasaurus 1817: 1816:Majungasaurus 1813: 1809: 1804: 1802: 1801: 1796: 1795:Majungasaurus 1792: 1788: 1784: 1780: 1779:Majungasaurus 1776: 1772: 1768: 1764: 1760: 1759:Majungasaurus 1756: 1748: 1741: 1739: 1737: 1732: 1728: 1727:Majungasaurus 1724: 1723:Majungasaurus 1719: 1717: 1713: 1709: 1708:Majungasaurus 1705: 1701: 1697: 1693: 1689: 1685: 1681: 1680:Majungasaurus 1677: 1669: 1668:Majungasaurus 1664: 1657: 1655: 1653: 1652:Majungasaurus 1649: 1645: 1641: 1637: 1636:Majungasaurus 1632: 1630: 1629:Majungasaurus 1626: 1625: 1621: 1617: 1616:Majungasaurus 1613: 1612:Majungasaurus 1609: 1608:Majungasaurus 1605: 1604:Majungasaurus 1601: 1600:Majungasaurus 1597: 1596:Majungasaurus 1589: 1587: 1585: 1581: 1580:Majungasaurus 1577: 1576:Majungasaurus 1572: 1571:Majungasaurus 1568: 1567:Majungasaurus 1564: 1563:Majungasaurus 1560: 1556: 1555:Majungasaurus 1550: 1546: 1545:Majungasaurus 1541: 1537: 1534: 1533:Majungasaurus 1529: 1528:Majungasaurus 1525: 1524: 1519: 1518:Majungasaurus 1515: 1511: 1507: 1503: 1502:Majungasaurus 1499: 1498:Majungasaurus 1495: 1491: 1486: 1483: 1482:Majungasaurus 1479: 1475: 1474:Majungasaurus 1470: 1468: 1464: 1460: 1455: 1454:Majungasaurus 1448: 1447: 1442: 1441:Majungasaurus 1437: 1430: 1428: 1426: 1422: 1421:Majungasaurus 1418: 1414: 1413:Majungasaurus 1410: 1406: 1405: 1400: 1396: 1392: 1391:Majungasaurus 1384: 1377: 1372: 1365: 1364: 1356: 1355: 1352: 1350: 1346: 1345: 1338: 1337: 1334: 1333: 1330: 1329: 1328: 1327:Rahiolisaurus 1321: 1320: 1317: 1316: 1313: 1312: 1311: 1304: 1303: 1300: 1299: 1296: 1294: 1290: 1289: 1288:Majungasaurus 1282: 1281: 1278: 1277: 1274: 1272: 1268: 1267: 1260: 1259: 1256: 1255: 1252: 1247: 1246: 1240: 1237: 1236: 1232: 1230: 1226: 1225:Majungasaurus 1222: 1213: 1209: 1207: 1203: 1200:, as well as 1199: 1198: 1193: 1189: 1188: 1183: 1182: 1177: 1173: 1172: 1167: 1166: 1161: 1160:Majungasaurus 1157: 1153: 1149: 1148:Majungasaurus 1145: 1141: 1137: 1132: 1130: 1126: 1122: 1118: 1114: 1111: 1107: 1103: 1099: 1095: 1091: 1087: 1086:Majungasaurus 1080: 1078: 1076: 1072: 1068: 1064: 1063:Majungasaurus 1060: 1055: 1053: 1052:Majungasaurus 1049: 1048:Majungasaurus 1045: 1041: 1037: 1033: 1032:Majungasaurus 1029: 1025: 1021: 1018: 1014: 1010: 1006: 1002: 998: 994: 990: 989:Majungasaurus 986: 985: 980: 976: 975:Majungasaurus 972: 963: 959: 957: 956: 951: 947: 943: 942:Majungasaurus 939: 935: 931: 927: 923: 919: 918:frontal bones 915: 914:Majungasaurus 911: 907: 906:Majungasaurus 903: 902:Majungasaurus 899: 895: 890: 889:Majungasaurus 887:The skull of 885: 883: 882: 876: 875:Majungasaurus 869: 868:Majungasaurus 865: 858: 856: 854: 850: 846: 845:Majungasaurus 842: 841:Majungatholus 838: 834: 830: 828: 823: 819: 815: 811: 810:Majungasaurus 806: 805:Majungasaurus 802: 798: 794: 793:Majungatholus 790: 786: 778: 774: 770: 767: 766:Majungasaurus 763: 758: 754: 752: 751:Majungasaurus 748: 744: 740: 736: 731: 729: 725: 721: 716: 706: 702: 701:Majungasaurus 698: 694: 690: 686: 682: 678: 674: 670: 662: 658: 654: 650: 646: 644: 643: 638: 635: 631: 627: 623: 619: 615: 611: 607: 603: 599: 596: 587: 580: 578: 576: 572: 571: 566: 562: 558: 557:apex predator 554: 553:Majungasaurus 550: 546: 542: 541:Majungasaurus 538: 537: 531: 529: 525: 521: 516: 513: 509: 508:Majungasaurus 504: 502: 501:Majungasaurus 498: 494: 493:Majungatholus 490: 489: 484: 480: 476: 473: 469: 465: 461: 458: 455: 451: 442: 426: 375: 374: 373:Majungasaurus 359: 356: 353:DepĂ©ret, 1896 350: 349:crenatissimus 348: 344: 343: 341: 338: 334: 329: 323: 321: 312: 309: 308:Binomial name 305: 301: 300: 294: 291: 290: 285: 280: 279: 278:Majungasaurus 272: 269: 268: 265: 259: 256: 255: 252: 251:Abelisauridae 246: 243: 242: 239: 233: 231:Superfamily: 230: 229: 226: 223: 220: 217: 216: 213: 210: 207: 204: 203: 200: 197: 194: 191: 190: 187: 184: 181: 180: 177: 174: 171: 170: 167: 164: 161: 160: 155: 150: 146: 143: 138: 134: 129: 115: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 54: 47: 46:Maastrichtian 43: 37: 36:Majungasaurus 33: 30: 19: 8709:Abelisaurids 8602: 8507: 8471: 8400: 8393: 8386: 8379: 8372: 8367:Ornithomimus 8365: 8358: 8351: 8344: 8337: 8330: 8323: 8305:Tyrannomimus 8303: 8295: 8287: 8280: 8275:Deinocheirus 8273: 8266: 8246: 8239: 8232: 8208: 8200: 8195:Nqwebasaurus 8193: 8188:Nedcolbertia 8186: 8179: 8172: 8166:Calamosaurus 8164: 8158:Aviatyrannis 8156: 8149: 8124: 8094: 8087: 8080: 8072: 8065: 8058: 8051: 8043: 8038:Aristosuchus 8036: 8030:Aniksosaurus 8028: 8010:Aniksosaurus 8008: 7978: 7971: 7963: 7909: 7874: 7867: 7860: 7853: 7779: 7772: 7765: 7745: 7738: 7730: 7725:Dynamoterror 7723: 7715: 7695: 7688: 7668: 7661: 7641: 7636:Asiatyrannus 7634: 7610: 7603: 7574:Dryptosaurus 7572: 7564: 7557: 7526: 7519: 7512: 7504: 7497: 7490: 7483: 7476: 7469: 7464:Jinbeisaurus 7462: 7455: 7448: 7440: 7435:Aviatyrannis 7433: 7426: 7392: 7387:Sinotyrannus 7385: 7378: 7371: 7364: 7343: 7336: 7314: 7307: 7266: 7259: 7252: 7244: 7237: 7229: 7221: 7216:Bicentenaria 7214: 7206: 7201:Asiamericana 7199: 7192: 7133: 7074: 7067: 7060: 7053: 7046: 7039: 7032: 7011: 7003: 6996: 6989: 6982: 6976:Deltadromeus 6974: 6966: 6959: 6908:Tyrannotitan 6906: 6899: 6892: 6885: 6865: 6841: 6836:Taurovenator 6834: 6827: 6820: 6815:Lajasvenator 6813: 6806: 6799: 6794:Concavenator 6792: 6784: 6777: 6752: 6744: 6737: 6729: 6721: 6701: 6694: 6686: 6679: 6672: 6665: 6640: 6632: 6624: 6617: 6610:Allosauridae 6578: 6571: 6566:Shidaisaurus 6564: 6557: 6550: 6530: 6522: 6497: 6489: 6482: 6475: 6444: 6436: 6428: 6421: 6394: 6386: 6374:Avetheropoda 6361:Avetheropoda 6332: 6291:Avetheropoda 6258: 6251: 6244:Spinosaurini 6231: 6224: 6217: 6210: 6202: 6174: 6167: 6160: 6139: 6131: 6124: 6119:Protathlitis 6117: 6110: 6085: 6078: 6071: 6063: 6028: 6021: 6014: 6009:Leshansaurus 6007: 6000: 5993: 5973: 5966: 5961:Megalosaurus 5959: 5954:Duriavenator 5952: 5932: 5907: 5891:Megalosauria 5872: 5865: 5860:Marshosaurus 5858: 5853:Condorraptor 5851: 5831: 5790: 5783: 5775: 5768: 5761: 5754: 5747: 5740: 5734:Cruxicheiros 5732: 5725: 5717: 5709: 5701: 5566: 5561:Quilmesaurus 5559: 5552: 5545: 5537: 5530: 5523: 5516: 5509: 5489: 5482: 5474: 5467: 5460: 5453: 5446: 5439: 5432: 5424: 5413:Brachyrostra 5399: 5379: 5371: 5365: 5364: 5357: 5350: 5342: 5335: 5315: 5310:Tralkasaurus 5308: 5301: 5294: 5287: 5280: 5273: 5266: 5259: 5251: 5244: 5212:Vespersaurus 5210: 5205:Velocisaurus 5203: 5196: 5189: 5182: 5162: 5155: 5148: 5128: 5121: 5114: 5106: 5100:Deltadromeus 5098: 5090: 5084:Compsosuchus 5082: 5075: 5067: 5059: 5034: 5026: 5020:Lametasaurus 5018: 5010: 4976: 4971:Ceratosaurus 4969: 4949: 4941: 4934: 4926: 4919: 4903:Ceratosauria 4888: 4881: 4806: 4799: 4757: 4749: 4742: 4734: 4726: 4719: 4712: 4705: 4685: 4680:Podokesaurus 4678: 4671: 4664: 4656: 4632: 4624: 4617: 4610: 4603: 4596: 4589: 4582: 4575: 4568: 4563:Gojirasaurus 4561: 4556:Dracovenator 4554: 4546: 4539: 4527:Neotheropoda 4512: 4504: 4496: 4490:Guaibasaurus 4488: 4481: 4473: 4465: 4459:Chindesaurus 4457: 4347: 4338: 4270: 4260: 4189: 4185: 4176: 4135: 4131: 4125: 4084: 4080: 4074: 4041: 4037: 4031: 3998: 3994: 3988: 3963: 3959: 3914: 3910: 3904: 3879: 3875: 3872: 3868: 3838: 3834: 3825: 3813: 3794: 3790: 3784: 3776: 3749: 3745: 3735: 3723:. Retrieved 3719:the original 3701:(1): 76–90. 3698: 3694: 3688: 3680: 3661: 3647: 3622: 3618: 3615: 3611: 3555: 3551: 3534:the original 3513: 3509: 3503: 3457: 3453: 3443: 3402: 3398: 3394: 3334: 3330: 3320: 3287: 3284:Paleobiology 3283: 3269: 3228: 3224: 3210: 3185: 3181: 3174: 3149: 3145: 3141: 3132: 3113: 3055: 3049: 3008: 3004: 3001: 2997: 2991: 2958: 2954: 2950: 2944: 2919: 2915: 2909: 2892: 2888: 2884: 2878: 2853: 2849: 2846: 2842: 2836: 2811: 2807: 2804: 2800: 2794: 2769: 2766: 2762: 2753: 2700: 2693: 2668: 2664: 2660: 2654: 2629: 2625: 2616: 2591: 2587: 2573: 2540: 2536: 2522: 2481: 2477: 2471: 2463:the original 2442: 2438: 2420: 2387: 2383: 2377: 2368: 2364: 2358: 2349: 2345: 2339: 2314: 2310: 2307: 2303: 2268: 2264: 2257: 2194: 2188: 2184: 2181:Rapetosaurus 2180: 2170: 2164: 2155: 2135: 2133: 2116: 2108:debris flows 2092: 2066: 2057: 2034: 2026: 2024: 2006: 1997:Paleoecology 1990: 1982: 1980: 1963: 1959: 1958: 1939: 1935: 1933: 1917: 1904: 1899: 1894:neural spine 1867: 1846: 1840: 1830:Ceratosaurus 1828: 1820: 1815: 1805: 1798: 1794: 1778: 1775:crocodilians 1758: 1753: 1726: 1722: 1720: 1707: 1679: 1673: 1667: 1651: 1635: 1633: 1628: 1622: 1615: 1611: 1607: 1603: 1599: 1595: 1593: 1584:Rapetosaurus 1583: 1579: 1575: 1570: 1566: 1562: 1559:Rapetosaurus 1558: 1554: 1553: 1549:Rapetosaurus 1548: 1544: 1532: 1527: 1521: 1517: 1501: 1497: 1487: 1481: 1473: 1471: 1453: 1451: 1446:Rapetosaurus 1444: 1440: 1420: 1412: 1404:Ceratosaurus 1402: 1398: 1394: 1390: 1389: 1373:Paleobiology 1342: 1341: 1325: 1324: 1308: 1307: 1287: 1286: 1285: 1264: 1263: 1250: 1224: 1223:2013 places 1220: 1218: 1201: 1195: 1191: 1185: 1179: 1175: 1169: 1163: 1159: 1151: 1147: 1140:evolutionary 1133: 1113:Ceratosauria 1085: 1084: 1066: 1062: 1056: 1051: 1047: 1043: 1039: 1031: 1023: 988: 982: 978: 974: 973:skeleton of 968: 953: 941: 913: 905: 901: 893: 888: 886: 879: 874: 873: 867: 844: 840: 825: 809: 804: 800: 796: 792: 788: 782: 765: 761: 750: 732: 727: 700: 697:Dryptosaurus 696: 693:Megalosaurus 692: 681:RenĂ© Lavocat 666: 656: 655:specimen of 642:Dryptosaurus 640: 636: 629: 621: 614:Megalosaurus 613: 592: 570:Rapetosaurus 568: 552: 540: 536:Megalosaurus 534: 532: 507: 505: 500: 492: 487: 486: 372: 371: 370: 357: 347:Megalosaurus 345: 319: 314: 298: 297: 277: 276: 218: 205: 192: 35: 29: 8627:Wikispecies 8532:Wikispecies 8432:Maniraptora 8421:Maniraptora 8402:Tototlmimus 8332:Anserimimus 8282:Garudimimus 8268:Beishanlong 8210:Valdoraptor 8151:Arkansaurus 8096:Xunmenglong 7973:Juravenator 7767:Tarbosaurus 7643:Nanuqsaurus 7612:Gorgosaurus 7528:Megaraptora 7514:Timurlengia 7246:Sciurumimus 7194:Aratasaurus 7062:Murusraptor 6984:Fukuiraptor 6947:Megaraptora 6829:Shaochilong 6801:Eocarcharia 6754:Megaraptora 6674:Lusovenator 6552:Alpkarakush 6409:Carnosauria 6260:Spinosaurus 6233:Siamosaurus 6133:Suchosaurus 6080:Iberospinus 6016:Magnosaurus 5995:Afrovenator 5968:Torvosaurus 5719:Chilesaurus 5532:Carnotaurus 5525:Caletodraco 5511:Abelisaurus 5337:Arcovenator 5184:Laevisuchus 5175:Noasaurinae 5077:Berthasaura 5048:Noasauridae 4978:Genyodectes 4943:Lukousaurus 4936:Jubbulpuria 4736:Panguraptor 4714:Coelophysis 4707:Camposaurus 4666:Panguraptor 4658:Dracoraptor 4634:Zupaysaurus 4619:Tachiraptor 4598:Sarcosaurus 4498:Nhandumirim 2961:(1): 1–16. 2121:tidal flats 2097:flood plain 2089:seasonality 1972:postorbital 1855:allosaurids 1843:pathologies 1624:Coelophysis 1590:Cannibalism 1492:out of the 1490:mineralized 1395:Carnotaurus 1266:Arcovenator 1197:Abelisaurus 1176:Carnotaurus 1152:Carnotaurus 1136:systematics 1102:Noasauridae 1067:Carnotaurus 1044:Carnotaurus 1024:Carnotaurus 1005:pleurocoels 979:Carnotaurus 971:postcranial 940:, although 910:nasal bones 894:Carnotaurus 881:Carnotaurus 859:Description 835:written in 679:. In 1955, 575:cannibalism 454:abelisaurid 257:Subfamily: 51:70–66  8703:Categories 8360:Gallimimus 8289:Harpymimus 8234:Harpymimus 7855:Iliosuchus 7825:see below↓ 7654:Alioramini 7471:Juratyrant 7457:Eotyrannus 7442:Bagaraatan 7394:Yutyrannus 7328:Coeluridae 7254:Vayuraptor 7100:see below↓ 7076:Tratayenia 7069:Orkoraptor 7055:Megaraptor 6961:Aoniraptor 6894:Mapusaurus 6822:Sauroniops 6786:Altispinax 6739:Neovenator 6681:Siamraptor 6667:Datanglong 6634:Epanterias 6626:Antrodemus 6619:Allosaurus 6600:Allosauria 6423:Altispinax 6388:Gasosaurus 6300:see below↓ 6176:Suchomimus 5777:Sinosaurus 5612:see below↓ 5568:Viavenator 5518:Aucasaurus 5455:Ilokelesia 5426:Dahalokely 5381:Rajasaurus 5359:Indosaurus 5352:Genusaurus 5344:Dahalokely 5268:Indosuchus 5164:Limusaurus 5123:Ligabueino 5116:Kiyacursor 5108:Genusaurus 5092:Dahalokely 5012:Betasuchus 4883:Dornraptor 4783:see below↓ 4759:Segisaurus 4612:Sinosaurus 4475:Eodromaeus 4404:see below↓ 4352:Dinosauria 4334:Sauropsida 4259:about the 3752:: 104553. 2557:1026511658 2371:: 256–259. 2352:: 257–265. 2271:: 176–194. 2250:References 2015:Madagascar 1900:Allosaurus 1886:bone spurs 1800:Indosaurus 1787:cerebellum 1729:, and the 1523:Allosaurus 1443:chasing a 1399:Rajasaurus 1344:Rajasaurus 1310:Indosaurus 1192:Aucasaurus 1181:Ilokelesia 1171:Indosaurus 1165:Rajasaurus 1117:convergent 1110:infraorder 1071:astragalus 1040:Aucasaurus 984:Aucasaurus 922:integument 898:premaxilla 602:Madagascar 472:Cretaceous 464:Madagascar 212:Saurischia 199:Dinosauria 8523:Q18511009 8436:includes 8381:Rativates 8374:Qiupalong 8074:Scipionyx 8067:Mirischia 7740:Lythronax 7732:Labocania 7663:Alioramus 7316:Labocania 7034:Aerosteon 6580:Sinraptor 6484:Erectopus 6219:Irritator 5804:Orionides 5689:Tetanurae 5676:Tetanurae 5603:Tetanurae 5539:Elemgasem 5462:Llukalkan 5441:Elemgasem 5198:Noasaurus 5061:Afromimus 4869:Averostra 4856:Averostra 4774:Averostra 4626:Velocipes 4445:Theropoda 4432:Theropoda 4395:Theropoda 4320:Kingdom: 4304:Theropoda 4168:205013285 3896:130262308 3725:August 3, 3188:: 63–86. 3166:131148538 2870:131213722 2786:129240095 2720:985402380 2671:: 71–89. 2166:Rahonavis 2071:from the 2069:separated 2019:Mahajanga 1874:cartilage 1870:exostosis 1851:tetanuran 1837:Pathology 1827:relative 1783:flocculus 1767:inner ear 1731:tetanuran 1640:scavenged 1156:phylogeny 1144:cladistic 1125:atrophied 1075:calcaneum 1001:vertebrae 818:monograph 565:sauropods 561:ecosystem 441:Mahajanga 292:Species: 225:Theropoda 172:Kingdom: 166:Eukaryota 8612:Wikidata 8517:Wikidata 7965:Coelurus 7373:Kileskus 7366:Guanlong 7338:Coelurus 7231:Gualicho 6731:Gualicho 6112:Baryonyx 5448:Guemesia 5275:Kryptops 5036:Ozraptor 4673:Pendraig 4328:Chordata 4326:Phylum: 4322:Animalia 4253:Archived 4214:11484051 4160:11206544 4117:27078534 4109:16222297 3715:85654858 3639:13873519 3580:16015329 3530:84175628 3484:17148302 3427:12673249 3371:31774829 3331:PLOS ONE 3312:85725299 3278:(2002). 3253:11234010 3112:(eds.). 3082:15306329 3025:54704403 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3073:1691741 2963:Bibcode 2895:: 58A. 2673:Bibcode 2506:9582112 2486:Bibcode 2478:Science 2447:Bibcode 2412:4345348 2392:Bibcode 2085:climate 2077:degrees 1890:osteoma 1863:phalanx 1694:-style 1648:lizards 1459:torsion 1431:Feeding 1417:display 1036:humerus 1020:tendons 950:dentary 946:maxilla 926:keratin 849:neotype 829:Memoirs 779:.MAJ 1) 769:frontal 653:Neotype 593:French 559:in its 512:bipedal 483:species 448:) is a 434:  328:DepĂ©ret 317:† 296:† 284:Lavocat 270:Genus: 120:↓ 8174:Hexing 7862:Kakuru 7450:Dilong 5491:Thanos 5469:Niebla 5289:Rugops 5282:Kurupi 4220:  4212:  4186:Nature 4166:  4158:  4132:Nature 4115:  4107:  4081:Nature 4064:  4021:  3995:Nature 3978:  3894:  3841:: 124. 3713:  3668:  3637:  3586:  3578:  3552:Nature 3528:  3482:  3472:  3433:  3425:  3399:Nature 3369:  3359:  3310:  3259:  3251:  3225:Nature 3164:  3120:  3080:  3070:  3023:  2981:  2934:  2868:  2826:  2784:  2718:  2708:  2644:  2606:  2563:  2555:  2512:  2504:  2410:  2384:Nature 2329:  2157:Vorona 2129:marine 1974:, and 1712:oxygen 1704:sacral 1514:snakes 1478:felids 1467:canids 1401:, and 1221:et al. 1202:Rugops 1094:family 1009:muscle 955:Rugops 938:crowns 814:pelvis 775:dome ( 715:sauros 710:ÏƒÎ±Ï…ÏÎżÏ‚ 634:suffix 510:was a 475:Period 443:lizard 286:, 1955 8686:53957 8591:38582 8438:birds 7478:Moros 7005:Siats 6746:Siats 6688:Siats 4348:Clade 4339:Clade 4218:S2CID 4164:S2CID 4113:S2CID 4019:S2CID 3976:S2CID 3892:S2CID 3831:(PDF) 3711:S2CID 3635:S2CID 3584:S2CID 3526:S2CID 3431:S2CID 3308:S2CID 3257:S2CID 3162:S2CID 3021:S2CID 2979:S2CID 2932:S2CID 2866:S2CID 2824:S2CID 2782:S2CID 2759:(PDF) 2642:S2CID 2604:S2CID 2561:S2CID 2510:S2CID 2408:S2CID 2327:S2CID 2144:frogs 2115:, so 2101:river 2054:stage 1983:et al 1968:jugal 1876:or a 1825:basal 1763:brain 1692:avian 1119:with 1090:clade 1059:tibia 934:sinus 707:word 705:Greek 677:Paris 628:word 626:Latin 567:like 528:teeth 520:skull 450:genus 219:Clade 206:Clade 193:Clade 8655:GBIF 8560:GBIF 8430:see 7048:Maip 4506:Tawa 4381:see 4210:PMID 4156:PMID 4105:PMID 4062:PMID 3727:2007 3666:ISBN 3576:PMID 3480:PMID 3423:PMID 3367:PMID 3249:PMID 3118:ISBN 3078:PMID 2716:OCLC 2706:ISBN 2553:OCLC 2502:PMID 2193:and 2140:fish 2045:and 2005:All 1903:and 1884:and 1857:and 1765:and 1700:lung 1547:and 1494:skin 1194:and 1168:and 1073:and 1042:and 1028:neck 1013:ribs 997:neck 981:and 969:The 853:ICZN 785:FMNH 777:MNHN 764:: a 722:and 695:and 685:MNHN 661:MNHN 431:lit. 59:PreꞒ 8642:EoL 8547:EoL 4202:doi 4190:412 4148:doi 4136:409 4097:doi 4085:437 4054:doi 4042:279 4011:doi 3999:382 3968:doi 3927:doi 3884:doi 3799:doi 3762:doi 3750:115 3703:doi 3627:doi 3568:doi 3556:436 3518:doi 3470:PMC 3462:doi 3415:doi 3403:422 3397:". 3357:PMC 3347:doi 3300:doi 3241:doi 3229:409 3198:doi 3186:100 3154:doi 3068:PMC 3060:doi 3056:271 3013:doi 2971:doi 2924:doi 2897:doi 2858:doi 2816:doi 2774:doi 2681:doi 2634:doi 2596:doi 2545:doi 2494:doi 2482:280 2455:doi 2400:doi 2388:279 2319:doi 2013:of 1915:). 1791:eye 1678:of 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Index

Majungasaurus crenatissimus
Late Cretaceous
Maastrichtian
Ma
PreꞒ
Ꞓ
O
S
D
C
P
T
J
K
Pg
N

Stony Brook University
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Saurischia
Theropoda
Abelisauroidea
Abelisauridae
Majungasaurinae
Majungasaurus

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