337:
solitary bachelors or have ranges overlapping more than one female. The average size of a female territory is 4 hectares (9.9 acres) with a male territory averaging 3.8 hectares (9.4 acres). The lemurs sometimes gather at the places where the territories overlap; there is no aggression on these occasions, but much vocalisation for a period of ten or twenty minutes. The animal moves on all fours, running rapidly along branches, climbing and jumping, mostly at a few metres above the ground, but descending to the forest floor on occasion.
42:
103:
234:
61:
474:
The male and female are in frequent vocal communication throughout the night, but this species in unusual among nocturnal lemurs in not communicating through olfactory signals, and the urine and faeces are allowed to drop to the forest floor. A large cutaneous gland on the throat of the male is used
336:
The social system of the
Masoala fork-marked lemur has been described as pre-gregarious. Some adults live as monogamous couples, sharing the same nest hole by day but moving about independently for at least part of the night (the male often follows a few metres behind the female), but a few live as
307:
The
Masoala fork-marked lemur is so-called because the black stripe along its spine divides on the crown, the two forks continuing on either side of the head to the eye and along either side of the muzzle. The rest of the pelage is some shade of reddish or brownish grey. The head-and-body length is
420:
tree in turn, feeding on the gum and moving on to the next tree. It has a complex itinerary which it regularly follows, and later in the night it visits other less-favoured gum-sources and returns to the previously visited trees and gleans any gum it has missed. The same
362:
species, the toothcomb has evolved to be more robust and is used to scrape the gum that exudes from holes on a tree made by insects. This structure consists of a row of lower incisors that are long and forward pointing. The favoured food sources are trees in the genus
483:
for three to four days in June, giving birth to a single offspring in
November or December. This remains in the nesting hole of the parents at first, but as it gets older it is carried by the mother, clinging to her underside, and when it gets larger, to her back.
444:, which it stalks on the tree bark and among the foliage, catching it with its hands and stuffing it into its mouth. Hunting takes place late in the night when the main gum-gathering activities are over. The animal's
1636:
308:
in the range 227 to 285 mm (9 to 11 in), with a tail of 285 to 370 mm (11 to 15 in). This lemur typically weighs between 300 and 500 g (11 and 18 oz).
1502:
505:
296:
1567:
823:
1621:
1476:
1515:
790:
737:
676:
630:
404:
trees and access to about nine more in overlapping territories. The animal emerges at dusk from its sleeping hole in a large tree such as a
1631:
765:
223:
1541:
1358:
816:
287:
that exudes from insect holes on the surface of certain trees, but it supplements its gum diet with insect prey. This primate is
65:
1349:
1258:
960:
409:
1559:
622:
1520:
1019:
1136:
479:
when males, females and juveniles groom each other, and the male may rub the gland on the female. The female comes into
1154:
933:
1554:
1226:
809:
608:
1572:
1267:
1109:
1055:
1181:
1073:
102:
1100:
1046:
1010:
496:
population is suspected of being in decline as its forest habitat is being destroyed by timber harvesting and
377:
328:
and it is present at altitudes up to about 1,000 m (3,300 ft). It occurs in moist lowland forests.
1299:
1282:
1208:
1118:
942:
924:
385:
and leave characteristic galleries that ooze gum. This lemur has also been observed licking the flowers of
1390:
1340:
1091:
1082:
978:
969:
906:
897:
1199:
1028:
393:
207:
1064:
348:
forests. The
Masoala fork-marked lemur has become specialized for harvesting this substance. Like most
387:
41:
1463:
1437:
1428:
1145:
1127:
553:
951:
915:
292:
288:
50:
1037:
365:
97:
578:
1507:
1580:
1450:
1314:
1241:
1217:
1190:
1172:
786:
733:
672:
636:
626:
321:
184:
780:
729:
662:
1585:
1163:
668:
548:
1533:
616:
233:
1626:
1374:
801:
476:
291:
and a single offspring is born in
November or December. It is declining in numbers due to
17:
1442:
833:
758:
693:
497:
468:
174:
530:
452:. In captivity this lemur was found to be very selective about what it ate, rejecting
1615:
1528:
869:
723:
612:
539:
504:, and receives some protection from being present in several national parks, but the
164:
70:
440:
content, and the
Masoala fork-marked lemur needs to supplement its diet with animal
1455:
604:
426:
349:
1598:
1546:
1489:
1422:
993:
880:
453:
341:
284:
1413:
400:
This species is nocturnal and has a territory containing about three exclusive
277:. It is a small nocturnal animal with large eyes, greyish fur and a long tail.
274:
640:
441:
405:
371:
353:
345:
325:
114:
1407:
851:
437:
391:, and feeding on a "syrup" produced by insect larvae on thin branches of
317:
134:
529:
Louis, E.E.; Randriamampoinana, R.; Bailey, C.A.; Sefczek, T.M. (2020).
1481:
863:
266:
154:
1494:
857:
845:
480:
464:
457:
449:
445:
358:
144:
124:
1384:
1593:
501:
460:
270:
87:
81:
618:
Mammal
Species of the World: A Taxonomic and Geographic Reference
1468:
1388:
805:
782:
Nocturnal
Malagasy primates: Ecology, Physiology, and Behavior
664:
Nocturnal
Malagasy primates: Ecology, Physiology, and Behavior
524:
522:
520:
369:, but certain other tree species also provide gum, including
725:
Lemurs of
Madagascar and the Comoros: The IUCN Red Data Book
508:
has assessed its conservation status as being endangered.
299:
has assessed its conservation status as being endangered.
320:
to northeastern Madagascar. Its range extends from the
381:. Certain beetle larvae burrow just under the bark of
448:
are invariably found to contain fragments of insect
1397:
1312:
1280:
1239:
991:
878:
757:
1637:Taxa named by Henri Marie Ducrotay de Blainville
554:10.2305/IUCN.UK.2020-2.RLTS.T16872A115570701.en
506:International Union for Conservation of Nature
297:International Union for Conservation of Nature
817:
273:found in the coastal forests of northeastern
8:
722:Harcourt, Caroline; Thornback, Jane (1990).
500:agriculture. It is listed on Appendix I of
467:, but feeding avidly on a large moth and a
1385:
824:
810:
802:
425:trees are exploited by day by a bird, the
232:
59:
40:
31:
656:
654:
652:
650:
552:
751:
749:
573:
571:
516:
7:
540:IUCN Red List of Threatened Species
1359:Amber Mountain fork-marked lemur (
785:. Elsevier Science. pp. 80–.
766:The Johns Hopkins University Press
25:
907:Southern fat-tailed dwarf lemur (
1622:IUCN Red List endangered species
340:The diet consists mainly of the
241:Masoala fork-marked lemur range
101:
1350:Pariente's fork-marked lemur (
1259:Coquerel's giant mouse lemur (
961:Montagne d'Ambre dwarf lemur (
934:Lesser iron-gray dwarf lemur (
779:Charles-Dominique, P. (2012).
661:Charles-Dominique, P. (2012).
623:Johns Hopkins University Press
283:is a specialist feeder on the
1:
1227:Northern Rufous mouse lemur (
1020:Madame Berthe's mouse lemur (
760:Walker's Mammals of the World
408:, or the abandoned nest of a
1268:Northern giant mouse lemur (
1137:Margot Marsh's mouse lemur (
615:; Reeder, D. M (eds.).
579:"Checklist of CITES Species"
433:), which also feeds on gum.
410:Coquerel's giant mouse lemur
1332:Masoala fork-marked lemur (
1155:Mittermeier's mouse lemur (
416:), moving directly to each
1653:
1182:Golden-brown mouse lemur (
1074:Reddish-gray mouse lemur (
263:Masoala fork-crowned lemur
35:Masoala fork-marked lemur
18:Masoala fork-crowned lemur
1632:Mammals described in 1839
1371:
1300:Hairy-eared dwarf lemur (
1110:MacArthur's mouse lemur (
925:Furry-eared dwarf lemur (
840:
832:Extant species of family
259:eastern fork-marked lemur
249:Masoala fork-marked lemur
240:
231:
213:
206:
98:Scientific classification
96:
79:
57:
48:
39:
34:
1341:Pale fork-marked lemur (
1056:Ganzhorn's mouse lemur (
898:Fat-tailed dwarf lemur (
692:Briercheck, Ken (1999).
378:Zanthoxylum tsihanimposa
312:Distribution and habitat
1200:Sambirano mouse lemur (
1101:Goodman's mouse lemur (
1029:Bongolava mouse lemur (
1011:Arnhold's mouse lemur (
1209:Simmons' mouse lemur (
1146:Marohita mouse lemur (
1128:Bemanasy mouse lemur (
1119:Claire's mouse lemur (
1047:Danfoss' mouse lemur (
943:Sibree's dwarf lemur (
547:: e.T16872A115570701.
375:spp., and the buds of
344:of trees in temperate
1092:Jonah's mouse lemur (
1083:Jolly's mouse lemur (
979:Thomas' dwarf lemur (
970:Groves' dwarf lemur (
952:Lavasoa dwarf lemur (
916:Greater dwarf lemur (
394:Rhopalocarpus lucidus
257:), also known as the
1322:(Fork-marked lemurs)
1249:(Giant mouse lemurs)
1065:Gerp's mouse lemur (
1038:Boraha mouse lemur (
702:Animal Diversity Web
698:: fork-marked lemur"
1218:Anosy mouse lemur (
1191:Brown mouse lemur (
1173:Pygmy mouse lemur (
352:primates, it has a
293:habitat destruction
51:Conservation status
1164:Gray mouse lemur (
756:Nowak, R. (1999).
671:. pp. 75–95.
613:Wilson, D. E.
605:Groves, C. P.
1609:
1608:
1581:Open Tree of Life
1391:Taxon identifiers
1382:
1381:
1325:
1293:
1252:
1004:
891:
792:978-0-323-15971-5
739:978-2-88032-957-0
728:. IUCN. pp.
678:978-0-323-15971-5
632:978-0-8018-8221-0
322:Masoala Peninsula
245:
244:
91:
74:
16:(Redirected from
1644:
1602:
1601:
1589:
1588:
1576:
1575:
1563:
1562:
1550:
1549:
1537:
1536:
1524:
1523:
1511:
1510:
1498:
1497:
1485:
1484:
1472:
1471:
1459:
1458:
1446:
1445:
1433:
1432:
1431:
1418:
1417:
1416:
1386:
1361:P. electromontis
1324:
1323:
1319:
1292:
1291:
1287:
1251:
1250:
1246:
1202:M. sambiranensis
1139:M. margotmarshae
1103:M. lehilahytsara
1031:M. bongolavensis
1003:
1002:
998:
909:C. adipicaudatus
890:
889:
885:
826:
819:
812:
803:
797:
796:
776:
770:
769:
764:(6th ed.).
763:
753:
744:
743:
719:
713:
712:
710:
708:
689:
683:
682:
669:Elsevier Science
658:
645:
644:
621:(3rd ed.).
609:"Order Primates"
601:
595:
594:
592:
590:
575:
566:
565:
563:
561:
556:
526:
388:Crateva greveana
236:
219:
199:P. furcifer
106:
105:
85:
68:
63:
62:
44:
32:
27:Species of lemur
21:
1652:
1651:
1647:
1646:
1645:
1643:
1642:
1641:
1612:
1611:
1610:
1605:
1597:
1592:
1584:
1579:
1571:
1566:
1558:
1553:
1545:
1540:
1532:
1527:
1519:
1514:
1506:
1501:
1493:
1488:
1480:
1475:
1467:
1462:
1454:
1449:
1443:Phaner_furcifer
1441:
1436:
1429:Phaner furcifer
1427:
1426:
1421:
1412:
1411:
1406:
1399:Phaner furcifer
1393:
1383:
1378:
1367:
1321:
1320:
1318:
1308:
1289:
1288:
1286:
1276:
1248:
1247:
1245:
1235:
1184:M. ravelobensis
1157:M. mittermeieri
1000:
999:
997:
987:
887:
886:
884:
874:
836:
830:
800:
793:
778:
777:
773:
755:
754:
747:
740:
721:
720:
716:
706:
704:
696:Phaner furcifer
691:
690:
686:
679:
660:
659:
648:
633:
625:. p. 114.
603:
602:
598:
588:
586:
577:
576:
569:
559:
557:
533:Phaner furcifer
528:
527:
518:
514:
490:
477:social grooming
414:Mirza coquereli
334:
314:
305:
254:Phaner furcifer
227:
221:
217:Phaner furcifer
215:
202:
100:
92:
75:
64:
60:
53:
28:
23:
22:
15:
12:
11:
5:
1650:
1648:
1640:
1639:
1634:
1629:
1624:
1614:
1613:
1607:
1606:
1604:
1603:
1590:
1577:
1564:
1551:
1538:
1525:
1512:
1499:
1486:
1473:
1460:
1447:
1434:
1419:
1403:
1401:
1395:
1394:
1389:
1380:
1379:
1372:
1369:
1368:
1366:
1365:
1356:
1347:
1338:
1328:
1326:
1310:
1309:
1307:
1306:
1296:
1294:
1290:(mouse lemurs)
1278:
1277:
1275:
1274:
1265:
1255:
1253:
1237:
1236:
1234:
1233:
1224:
1215:
1206:
1197:
1188:
1179:
1170:
1161:
1152:
1143:
1134:
1125:
1116:
1112:M. macarthurii
1107:
1098:
1089:
1080:
1076:M. griseorufus
1071:
1062:
1053:
1044:
1035:
1026:
1017:
1007:
1005:
1001:(mouse lemurs)
989:
988:
986:
985:
976:
967:
958:
954:C. lavasoensis
949:
940:
931:
922:
913:
904:
894:
892:
888:(dwarf lemurs)
876:
875:
873:
872:
866:
860:
854:
848:
841:
838:
837:
834:Cheirogaleidae
831:
829:
828:
821:
814:
806:
799:
798:
791:
771:
745:
738:
714:
684:
677:
646:
631:
596:
567:
515:
513:
510:
498:slash-and-burn
489:
486:
469:praying mantis
436:Gum is low in
333:
330:
324:southwards to
316:This lemur is
313:
310:
304:
301:
243:
242:
238:
237:
229:
228:
222:
211:
210:
204:
203:
196:
194:
190:
189:
182:
178:
177:
175:Cheirogaleidae
172:
168:
167:
162:
158:
157:
152:
148:
147:
142:
138:
137:
132:
128:
127:
122:
118:
117:
112:
108:
107:
94:
93:
80:
77:
76:
58:
55:
54:
49:
46:
45:
37:
36:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
1649:
1638:
1635:
1633:
1630:
1628:
1625:
1623:
1620:
1619:
1617:
1600:
1595:
1591:
1587:
1582:
1578:
1574:
1569:
1565:
1561:
1556:
1552:
1548:
1543:
1539:
1535:
1530:
1526:
1522:
1517:
1513:
1509:
1504:
1500:
1496:
1491:
1487:
1483:
1478:
1474:
1470:
1465:
1461:
1457:
1452:
1448:
1444:
1439:
1435:
1430:
1424:
1420:
1415:
1409:
1405:
1404:
1402:
1400:
1396:
1392:
1387:
1377:
1376:
1370:
1364:
1362:
1357:
1355:
1353:
1348:
1346:
1344:
1343:P. pallescens
1339:
1337:
1335:
1330:
1329:
1327:
1317:
1316:
1311:
1305:
1303:
1298:
1297:
1295:
1285:
1284:
1279:
1273:
1271:
1266:
1264:
1262:
1257:
1256:
1254:
1244:
1243:
1238:
1232:
1230:
1225:
1223:
1221:
1216:
1214:
1212:
1207:
1205:
1203:
1198:
1196:
1194:
1189:
1187:
1185:
1180:
1178:
1176:
1171:
1169:
1167:
1162:
1160:
1158:
1153:
1151:
1149:
1144:
1142:
1140:
1135:
1133:
1131:
1126:
1124:
1122:
1117:
1115:
1113:
1108:
1106:
1104:
1099:
1097:
1095:
1090:
1088:
1086:
1081:
1079:
1077:
1072:
1070:
1068:
1063:
1061:
1059:
1054:
1052:
1050:
1045:
1043:
1041:
1036:
1034:
1032:
1027:
1025:
1023:
1018:
1016:
1014:
1009:
1008:
1006:
996:
995:
990:
984:
982:
977:
975:
973:
968:
966:
964:
963:C. andysabini
959:
957:
955:
950:
948:
946:
941:
939:
937:
936:C. minusculus
932:
930:
928:
923:
921:
919:
914:
912:
910:
905:
903:
901:
896:
895:
893:
883:
882:
877:
871:
870:Strepsirrhini
867:
865:
861:
859:
855:
853:
849:
847:
843:
842:
839:
835:
827:
822:
820:
815:
813:
808:
807:
804:
794:
788:
784:
783:
775:
772:
767:
762:
761:
752:
750:
746:
741:
735:
731:
727:
726:
718:
715:
703:
699:
697:
688:
685:
680:
674:
670:
666:
665:
657:
655:
653:
651:
647:
642:
638:
634:
628:
624:
620:
619:
614:
610:
606:
600:
597:
584:
580:
574:
572:
568:
555:
550:
546:
542:
541:
536:
534:
525:
523:
521:
517:
511:
509:
507:
503:
499:
495:
487:
485:
482:
478:
472:
470:
466:
462:
459:
455:
451:
447:
443:
439:
434:
432:
431:Coua cristata
428:
424:
419:
415:
411:
407:
403:
398:
396:
395:
390:
389:
384:
380:
379:
374:
373:
368:
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361:
360:
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331:
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290:
286:
282:
278:
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272:
268:
264:
260:
256:
255:
250:
239:
235:
230:
225:
220:
218:
212:
209:
208:Binomial name
205:
201:
200:
195:
192:
191:
188:
187:
183:
180:
179:
176:
173:
170:
169:
166:
165:Strepsirrhini
163:
160:
159:
156:
153:
150:
149:
146:
143:
140:
139:
136:
133:
130:
129:
126:
123:
120:
119:
116:
113:
110:
109:
104:
99:
95:
89:
83:
78:
72:
67:
56:
52:
47:
43:
38:
33:
30:
19:
1398:
1373:
1360:
1351:
1342:
1333:
1331:
1313:
1302:A. trichotis
1301:
1281:
1269:
1261:M. coquereli
1260:
1240:
1229:M. tavaratra
1228:
1219:
1210:
1201:
1192:
1183:
1174:
1165:
1156:
1147:
1138:
1130:M. manitatra
1129:
1121:M. mamiratra
1120:
1111:
1102:
1093:
1084:
1075:
1066:
1058:M. ganzhorni
1057:
1048:
1039:
1030:
1021:
1012:
992:
980:
971:
962:
953:
944:
935:
927:C. crossleyi
926:
917:
908:
899:
881:Cheirogaleus
879:
781:
774:
759:
724:
717:
705:. Retrieved
701:
695:
687:
663:
617:
599:
587:. Retrieved
582:
558:. Retrieved
544:
538:
532:
493:
491:
473:
454:grasshoppers
435:
430:
427:crested coua
422:
417:
413:
401:
399:
392:
386:
382:
376:
370:
364:
357:
339:
335:
315:
306:
280:
279:
262:
258:
253:
252:
248:
246:
216:
214:
198:
197:
185:
29:
1490:iNaturalist
1423:Wikispecies
1352:P. parienti
1334:P. furcifer
1211:M. simmonsi
1175:M. myoxinus
1148:M. marohita
1049:M. danfossi
1013:M. arnholdi
585:. UNEP-WCMC
560:19 November
494:P. furcifer
342:natural gum
303:Description
281:P. furcifer
1616:Categories
1166:M. murinus
1085:M. jollyae
1022:M. berthae
994:Microcebus
981:C. thomasi
972:C. grovesi
945:C. sibreei
868:Suborder:
512:References
463:and small
423:Terminalia
418:Terminalia
402:Terminalia
383:Terminalia
366:Terminalia
350:lemuriform
295:, and the
289:monogamous
275:Madagascar
224:Blainville
161:Suborder:
84:Appendix I
66:Endangered
1283:Allocebus
1220:M. tanosi
1094:M. jonahi
1040:M. boraha
900:C. medius
844:Kingdom:
707:2 January
372:Adansonia
354:toothcomb
346:deciduous
326:Toamasina
193:Species:
121:Kingdom:
115:Eukaryota
1594:Species+
1560:12100029
1508:10593686
1414:Q1150927
1408:Wikidata
1375:Category
1193:M. rufus
1067:M. gerpi
918:C. major
864:Primates
858:Mammalia
852:Chordata
850:Phylum:
846:Animalia
641:62265494
607:(2005).
589:18 March
438:nitrogen
171:Family:
155:Primates
145:Mammalia
135:Chordata
131:Phylum:
125:Animalia
111:Domain:
71:IUCN 3.1
1547:1000968
1482:2436443
1270:M. zaza
862:Order:
856:Class:
475:during
465:lizards
397:trees.
332:Ecology
318:endemic
267:species
265:, is a
181:Genus:
151:Order:
141:Class:
86: (
69: (
1627:Lemurs
1586:836057
1573:261734
1521:572863
1469:324406
1315:Phaner
789:
736:
675:
639:
629:
488:Status
481:estrus
461:larvae
458:beetle
450:chitin
446:faeces
406:baobab
359:Phaner
226:, 1839
186:Phaner
1534:16872
1503:IRMNG
1495:43555
1456:4FSQ7
1242:Mirza
732:–69.
611:. In
583:CITES
502:CITES
356:; in
271:lemur
88:CITES
82:CITES
1599:6771
1568:NCBI
1529:IUCN
1516:ITIS
1477:GBIF
787:ISBN
734:ISBN
709:2017
673:ISBN
637:OCLC
627:ISBN
591:2015
562:2021
545:2020
492:The
442:prey
247:The
1555:MSW
1542:MDD
1464:EoL
1451:CoL
1438:ADW
549:doi
285:gum
269:of
261:or
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