1514:
1521:
1506:
130:
106:
1600:
411:"An Empirical Synthetic Pattern Study of Gars (lepisosteiformes) and Closely Related Species, Based Mostly on Skeletal Anatomy. the Resurrection of Holostei"
331:. They are known from only a few specimens from both localities, and may have not been permanent inhabitants of the fossil lakes where they were preserved.
1661:
481:"Exceptional preservation of internal organs in a new fossil species of freshwater shrimp (Caridea: Palaemonoidea) from the Eocene of Messel (Germany)"
1587:
1666:
1651:
532:"Phylogenomics of the Ancient and Species-Depauperate Gars Tracks 150 Million Years of Continental Fragmentation in the Northern Hemisphere"
579:
1513:
661:
1671:
1656:
320:
1520:
1646:
129:
1505:
572:
1641:
1540:
1613:
324:
1070:
799:
347:, which inhabited western North America during the same time period; both genera are classified in the
1284:
1049:
849:
822:
925:
769:
688:
388:, a new gar (Actinopterygii, Lepisosteidae) from the middle Eocene of Grube Messel (Hessen, Germany)"
1152:
1122:
966:
531:
1162:
1142:
1091:
1007:
917:
858:
831:
813:
565:
1186:
987:
865:
1261:
1193:
910:
789:
124:
1618:
980:
903:
341:
and other hard-shelled invertebrate prey. It shares the short, broad snout with the extinct gar
1578:
1251:
896:
351:
1605:
943:
502:
461:
442:"Peculiarities of the Messel fish fauna and their palaeoecological implications: a case study"
422:
337:
was an atypical gar with a short, broad snout and molariform teeth likely adapted to crushing
1207:
1132:
530:
Brownstein, Chase Doran; Yang, Liandong; Friedman, Matt; Near, Thomas J. (20 December 2022).
1488:
1477:
1444:
1368:
1241:
1106:
510:
492:
453:
308:
194:
1455:
883:
779:
348:
1398:
1341:
1225:
959:
952:
646:
618:
515:
296:
171:
1635:
1411:
1331:
1275:
1084:
1014:
998:
973:
838:
480:
383:
284:
63:
1311:
1021:
242:
117:
105:
1563:
1572:
1466:
1433:
1422:
1348:
1177:
1042:
1028:
673:
624:
38:
497:
1378:
1291:
1214:
1035:
932:
752:
730:
700:
588:
457:
359:
338:
304:
273:
184:
83:
48:
506:
479:
de
Mazancourt, Valentin; Wappler, Torsten; Wedmann, Sonja (27 October 2022).
465:
426:
1200:
1077:
1056:
441:
355:
141:
88:
1557:
1321:
1063:
630:
612:
343:
161:
78:
73:
58:
53:
43:
410:
1592:
328:
312:
93:
68:
606:
292:
288:
269:
151:
1534:
276:
1538:
599:
561:
358:, it is assumed that both diverged from one another during the
1302:
295:. It is known from two species, each from a famous freshwater
280:
204:
557:
354:. Although fossils of both genera are known only from the
1484:
1473:
1462:
1451:
1440:
1429:
1418:
1407:
1374:
1364:
1354:
1337:
1327:
1317:
1272:
1257:
1247:
1237:
1174:
1158:
1148:
1138:
1128:
1118:
880:
810:
795:
785:
775:
765:
1547:
1396:
1301:
1271:
1224:
1173:
1105:
997:
942:
879:
848:
821:
809:
750:
699:
672:
645:
573:
8:
1535:
1230:
1111:
888:
818:
758:
747:
678:
651:
642:
596:
580:
566:
558:
104:
22:
514:
496:
446:Palaeobiodiversity and Palaeoenvironments
371:
377:
375:
440:Micklich, Norbert (1 December 2012).
7:
382:Micklich, N.; Klappert, G. (2001).
1662:Prehistoric ray-finned fish genera
14:
1519:
1512:
1504:
128:
1:
1667:Fossil taxa described in 2001
1652:Eocene fish of North America
254:Micklich & Kappert, 2001
224:Micklich & Kappert, 2001
1688:
498:10.1038/s41598-022-23125-9
1501:
1233:
1114:
891:
761:
746:
727:
681:
654:
641:
595:
458:10.1007/s12549-012-0106-4
234:
229:
125:Scientific classification
123:
112:
103:
25:
283:that inhabited western
409:Grande, Lance (2010).
1672:Green River Formation
1657:Eocene fish of Europe
1614:Paleobiology Database
325:Green River Formation
1647:Prehistoric holostei
386:Masillosteus kelleri
251:Masillosteus kelleri
814:Kyphosichthyiformes
239:Masillosteus janeae
536:Systematic Biology
485:Scientific Reports
1629:
1628:
1541:Taxon identifiers
1532:
1531:
1528:
1527:
1499:
1498:
1392:
1391:
1388:
1387:
1101:
1100:
1071:Palaeomacrosemius
944:Callipurbeckiidae
875:
874:
800:Sangiorgioichthys
742:
741:
738:
737:
723:
722:
719:
718:
261:
260:
255:
246:
225:
1679:
1622:
1621:
1609:
1608:
1596:
1595:
1583:
1582:
1581:
1568:
1567:
1566:
1536:
1523:
1517:
1516:
1509:
1508:
1489:Pericentrophorus
1486:
1478:Paracentrophorus
1475:
1464:
1453:
1445:Archaeolepidotus
1442:
1431:
1420:
1409:
1376:
1369:Nhanulepisosteus
1366:
1356:
1339:
1329:
1319:
1285:Dentilepisosteus
1274:
1259:
1249:
1242:Araripelepidotes
1239:
1231:
1176:
1160:
1150:
1140:
1130:
1120:
1112:
1107:Lepisosteiformes
1050:Macrosemiocotzus
889:
882:
850:Lashanichthyidae
823:Kyphosichthyidae
819:
812:
797:
787:
777:
767:
759:
748:
679:
652:
643:
636:
635:
597:
582:
575:
568:
559:
552:
551:
549:
547:
527:
521:
520:
518:
500:
476:
470:
469:
437:
431:
430:
406:
400:
399:
379:
309:Messel Formation
268:("bony one from
253:
241:
223:
216:
195:Lepisosteiformes
133:
132:
116:specimen at the
108:
98:
35:
31:Temporal range:
23:
1687:
1686:
1682:
1681:
1680:
1678:
1677:
1676:
1632:
1631:
1630:
1625:
1617:
1612:
1604:
1599:
1591:
1586:
1577:
1576:
1571:
1562:
1561:
1556:
1543:
1533:
1524:
1511:
1503:
1495:
1456:Austrolepidotes
1384:
1297:
1267:
1220:
1169:
1097:
993:
938:
926:Quasimodichthys
884:Semionotiformes
871:
844:
805:
780:Luoxiongichthys
770:Diandongichthys
755:
734:
715:
695:
668:
637:
602:
601:
591:
586:
556:
555:
545:
543:
529:
528:
524:
478:
477:
473:
439:
438:
434:
421:(2A): iii–871.
408:
407:
403:
381:
380:
373:
368:
222:
214:
127:
99:
97:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
33:
32:
29:
19:
18:Genus of fishes
12:
11:
5:
1685:
1683:
1675:
1674:
1669:
1664:
1659:
1654:
1649:
1644:
1634:
1633:
1627:
1626:
1624:
1623:
1610:
1597:
1584:
1569:
1553:
1551:
1545:
1544:
1539:
1530:
1529:
1526:
1525:
1502:
1500:
1497:
1496:
1494:
1493:
1482:
1471:
1460:
1449:
1438:
1427:
1416:
1404:
1402:
1399:Incertae sedis
1394:
1393:
1390:
1389:
1386:
1385:
1383:
1382:
1372:
1362:
1352:
1345:
1342:Herreraichthys
1335:
1325:
1315:
1307:
1305:
1299:
1298:
1296:
1295:
1288:
1280:
1278:
1269:
1268:
1266:
1265:
1255:
1245:
1234:
1228:
1226:Lepisosteoidei
1222:
1221:
1219:
1218:
1211:
1204:
1197:
1190:
1182:
1180:
1171:
1170:
1168:
1167:
1156:
1153:Lanxangichthys
1146:
1136:
1126:
1123:Adrianaichthys
1115:
1109:
1103:
1102:
1099:
1098:
1096:
1095:
1088:
1081:
1074:
1067:
1060:
1053:
1046:
1039:
1032:
1025:
1018:
1011:
1003:
1001:
995:
994:
992:
991:
984:
977:
970:
967:Occitanichthys
963:
960:Macrosemimimus
956:
953:Callipurbeckia
948:
946:
940:
939:
937:
936:
929:
922:
914:
907:
900:
892:
886:
877:
876:
873:
872:
870:
869:
862:
854:
852:
846:
845:
843:
842:
835:
827:
825:
816:
807:
806:
804:
803:
793:
783:
773:
762:
756:
751:
744:
743:
740:
739:
736:
735:
728:
725:
724:
721:
720:
717:
716:
714:
713:
712:
711:
705:
703:
697:
696:
694:
693:
692:
691:
682:
676:
670:
669:
667:
666:
665:
664:
662:Actinopterygii
655:
649:
647:Actinopterygii
639:
638:
634:
633:
627:
621:
619:Actinopterygii
615:
609:
600:
593:
592:
587:
585:
584:
577:
570:
562:
554:
553:
522:
471:
452:(4): 585–629.
432:
401:
370:
369:
367:
364:
259:
258:
257:
256:
247:
232:
231:
227:
226:
212:
208:
207:
202:
198:
197:
192:
188:
187:
182:
175:
174:
172:Actinopterygii
169:
165:
164:
159:
155:
154:
149:
145:
144:
139:
135:
134:
121:
120:
110:
109:
101:
100:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
36:
30:
17:
13:
10:
9:
6:
4:
3:
2:
1684:
1673:
1670:
1668:
1665:
1663:
1660:
1658:
1655:
1653:
1650:
1648:
1645:
1643:
1642:Lepisosteidae
1640:
1639:
1637:
1620:
1615:
1611:
1607:
1602:
1598:
1594:
1589:
1585:
1580:
1574:
1570:
1565:
1559:
1555:
1554:
1552:
1550:
1546:
1542:
1537:
1522:
1518:
1515:
1507:
1491:
1490:
1483:
1480:
1479:
1472:
1469:
1468:
1461:
1458:
1457:
1450:
1447:
1446:
1439:
1436:
1435:
1428:
1425:
1424:
1417:
1414:
1413:
1412:Acentrophorus
1406:
1405:
1403:
1401:
1400:
1395:
1381:
1380:
1373:
1371:
1370:
1363:
1361:
1360:
1353:
1351:
1350:
1346:
1344:
1343:
1336:
1334:
1333:
1332:Grandemarinus
1326:
1324:
1323:
1316:
1314:
1313:
1309:
1308:
1306:
1304:
1303:Lepisosteidae
1300:
1294:
1293:
1289:
1287:
1286:
1282:
1281:
1279:
1277:
1276:Obaichthyidae
1270:
1264:
1263:
1256:
1254:
1253:
1246:
1244:
1243:
1236:
1235:
1232:
1229:
1227:
1223:
1217:
1216:
1212:
1210:
1209:
1205:
1203:
1202:
1198:
1196:
1195:
1191:
1189:
1188:
1184:
1183:
1181:
1179:
1172:
1165:
1164:
1163:Neosemionotus
1157:
1155:
1154:
1147:
1145:
1144:
1143:Khoratichthys
1137:
1135:
1134:
1127:
1125:
1124:
1117:
1116:
1113:
1110:
1108:
1104:
1094:
1093:
1092:Voelklichthys
1089:
1087:
1086:
1085:Uarbryichthys
1082:
1080:
1079:
1075:
1073:
1072:
1068:
1066:
1065:
1061:
1059:
1058:
1054:
1052:
1051:
1047:
1045:
1044:
1040:
1038:
1037:
1033:
1031:
1030:
1026:
1024:
1023:
1019:
1017:
1016:
1015:Enchelyolepis
1012:
1010:
1009:
1008:Agoultichthys
1005:
1004:
1002:
1000:
999:Macrosemiidae
996:
990:
989:
985:
983:
982:
978:
976:
975:
974:Paralepidotus
971:
969:
968:
964:
962:
961:
957:
955:
954:
950:
949:
947:
945:
941:
935:
934:
930:
928:
927:
923:
920:
919:
918:Neosemionotus
915:
913:
912:
908:
906:
905:
901:
899:
898:
894:
893:
890:
887:
885:
878:
868:
867:
863:
861:
860:
859:Lashanichthys
856:
855:
853:
851:
847:
841:
840:
839:Kyphosichthys
836:
834:
833:
832:Fuyuanichthys
829:
828:
826:
824:
820:
817:
815:
808:
802:
801:
794:
792:
791:
784:
782:
781:
774:
772:
771:
764:
763:
760:
757:
754:
749:
745:
733:
732:
726:
709:
708:
707:
706:
704:
702:
698:
690:
686:
685:
684:
683:
680:
677:
675:
671:
663:
659:
658:
657:
656:
653:
650:
648:
644:
640:
632:
628:
626:
622:
620:
616:
614:
610:
608:
604:
603:
598:
594:
590:
583:
578:
576:
571:
569:
564:
563:
560:
541:
537:
533:
526:
523:
517:
512:
508:
504:
499:
494:
490:
486:
482:
475:
472:
467:
463:
459:
455:
451:
447:
443:
436:
433:
428:
424:
420:
416:
412:
405:
402:
397:
393:
389:
387:
378:
376:
372:
365:
363:
361:
357:
353:
350:
346:
345:
340:
336:
332:
330:
326:
322:
318:
314:
310:
306:
302:
298:
294:
290:
286:
285:North America
282:
278:
275:
271:
267:
266:
252:
248:
244:
240:
236:
235:
233:
228:
221:
220:
213:
210:
209:
206:
205:Lepisosteidae
203:
200:
199:
196:
193:
190:
189:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
153:
150:
147:
146:
143:
140:
137:
136:
131:
126:
122:
119:
115:
111:
107:
102:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
28:
24:
21:
16:
1579:Masillosteus
1549:Masillosteus
1548:
1510:
1487:
1476:
1465:
1454:
1443:
1432:
1421:
1410:
1397:
1377:
1367:
1359:Masillosteus
1358:
1357:
1347:
1340:
1330:
1320:
1312:Atractosteus
1310:
1290:
1283:
1260:
1250:
1240:
1213:
1206:
1199:
1192:
1187:Camerichthys
1185:
1161:
1151:
1141:
1131:
1121:
1090:
1083:
1076:
1069:
1062:
1055:
1048:
1041:
1034:
1027:
1022:Eosemionotus
1020:
1013:
1006:
988:Tlayuamichin
986:
979:
972:
965:
958:
951:
931:
924:
916:
909:
902:
895:
866:Yudaiichthys
864:
857:
837:
830:
798:
788:
778:
768:
729:
629:Infraclass:
544:. Retrieved
542:(1): 213–227
539:
535:
525:
491:(1): 18114.
488:
484:
474:
449:
445:
435:
418:
414:
404:
395:
391:
385:
342:
335:Masillosteus
334:
333:
321:Fossil Butte
316:
300:
265:Masillosteus
264:
263:
262:
250:
238:
219:Masillosteus
218:
217:
178:
118:Field Museum
113:
34:Early Eocene
27:Masillosteus
26:
20:
15:
1573:Wikispecies
1467:Corunegenys
1434:Aphelolepis
1423:Alleiolepis
1349:Lepisosteus
1262:Thaiichthys
1194:Isanichthys
1178:Lepidotidae
1043:Macrosemius
1029:Histionotus
911:Lophionotus
790:Ticinolepis
689:Neopterygii
674:Neopterygii
625:Neopterygii
339:crustaceans
297:lagerstätte
291:during the
1636:Categories
1379:Oniichthys
1292:Obaichthys
1215:Scheenstia
1036:Legnonotus
981:Semiolepis
933:Semionotus
904:Lepidhoyas
753:Ginglymodi
731:Ginglymodi
710:see below↓
701:Ginglymodi
623:Subclass:
589:Ginglymodi
366:References
360:Cretaceous
305:Messel pit
301:M. kelleri
185:Ginglymodi
1564:Q18002378
1252:Pliodetes
1201:Lepidotes
1078:Propterus
1057:Notagogus
897:Hoyasotes
605:Kingdom:
507:2045-2322
466:1867-1608
427:0045-8511
356:Paleogene
352:Cuneatini
317:M. janeae
303:from the
272:") is an
148:Kingdom:
142:Eukaryota
114:M. janeae
1558:Wikidata
1322:Cuneatus
1064:Orthurus
631:Holostei
613:Chordata
611:Phylum:
607:Animalia
398:: 73–81.
344:Cuneatus
230:Species
201:Family:
162:Chordata
158:Phylum:
152:Animalia
138:Domain:
1606:1166796
1593:4839041
1208:Mengius
1133:Beiduyu
617:Class:
516:9613706
329:Wyoming
323:in the
313:Germany
307:in the
274:extinct
211:Genus:
191:Order:
168:Class:
1619:241983
546:5 June
513:
505:
464:
425:
415:Copeia
392:Kaupia
315:, and
293:Eocene
289:Europe
270:Messel
245:, 2010
243:Grande
1601:IRMNG
349:tribe
319:from
277:genus
179:Clade
1588:GBIF
687:see
660:see
548:2023
503:ISSN
462:ISSN
423:ISSN
419:2010
287:and
39:PreꞒ
511:PMC
493:doi
454:doi
327:of
311:of
281:gar
279:of
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450:92
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89:Pg
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581:e
574:t
567:v
550:.
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495::
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456::
429:.
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249:†
237:†
215:†
94:N
84:K
79:J
74:T
69:P
64:C
59:D
54:S
49:O
44:Ꞓ
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