1456:
1441:
1448:
121:
364:
is known only from a single specimen, an upper molar. Although the fossil material is scant, it has enough traits to differ it from all thylacinids. Stylar cusps B and D are well-developed. Furthermore, the aforementioned stylar cusps are compressed to the sides of the molar. The front most
491:
Woodhead, J.; Hand, S.J.; Archer, M.; Graham, I.; Sniderman, K.; Arena, D.A.; Black, K.H.; Godthelp, H.; Creaser, P.; Price, E. (2014). "Developing a radiometrically-dated chronologic sequence for
Neogene biotic change in Australia, from the Riversleigh World Heritage Area of Queensland".
388:
are known only from a single site at
Riversleigh, which dates to the Middle Miocene. The climate during the Middle Miocene would have been warm and permanently wet. Open rainforests would have covered the landscape at the time. Coexisting alongside
461:(2001). "Maximucinus muirheadae, gen. et sp. nov. (Thylacinidae : Marsupialia), from the Miocene of Riversleigh, north-western Queensland, with estimates of body weights for fossil thylacinids".
519:
Travouillon, K.J.; Legendre, S.; Archer, M.; Hand, S.J. (2009). "Palaeoecological analyses of
Riversleigh's Oligo-Miocene sites: implications for Oligo-Miocene climate change in Australia".
432:(i.e. minute protoconule and metaconule cusps), although it was clearly less specialised than that of later thylacinids as it still retains well-developed stylar cusps.
421:. The two families of carnivorous marsupials were likely able to coexist with one another due to differences in both body size and vertical habitat segregation.
353:
word for dog. The species name was chosen to honour fellow researcher
Jeanette Muirhead, for their contributions towards the study of fossil thylacinids.
1590:
1565:
1519:
1447:
1455:
1595:
661:
1575:
1440:
546:
Gillespie, A. K. (2023). "Two new marsupial lion taxa (Marsupialia, Thylacoleonidae) from the early and Middle
Miocene of Australia".
944:
1560:
323:
290:
591:"The pre-Pleistocene fossil thylacinids (Dasyuromorphia: Thylacinidae) and the evolutionary context of the modern thylacine"
376:
Wroe (2001) estimated its weight to be 18 kg (40 lbs), making it the largest thylacinid discovered from the middle
Miocene.
1570:
1580:
930:
916:
120:
1585:
1510:
369:(a small shelf-like structure situated at the base of a tooth's crown) continues into the preparacrista. Both the
909:
654:
1472:
992:
1532:
923:
232:
647:
555:
251:
293:. It was the largest thylacinid of its time, attaining a body size of 18 kg (40 lbs). The genus is
794:
1044:
1003:
825:
571:
115:
1537:
814:
458:
1061:
869:
1357:
1206:
858:
612:
407:
1406:
1385:
1023:
764:
620:
602:
563:
528:
501:
470:
366:
804:
1399:
951:
748:
700:
559:
971:
737:
625:
429:
370:
331:
278:
182:
30:
1554:
1295:
1217:
846:
575:
350:
67:
322:, is an isolated second upper molar. It was collected from the Ringtail site at the
314:
was described as a new genus and species of thylacinid in 2001 by
Stephen Wroe. The
1307:
1241:
1135:
1114:
1106:
1075:
1034:
730:
718:
195:
567:
532:
1504:
1413:
1142:
1099:
897:
670:
395:
42:
1495:
1330:
1316:
1279:
1185:
1158:
1128:
1121:
1013:
401:
327:
282:
274:
87:
52:
616:
505:
1341:
1230:
1196:
1084:
937:
774:
417:
294:
286:
172:
132:
92:
36:
589:
Rovinsky, Douglass S.; Evans, Alistair R.; Adams, Justin W. (2019-09-02).
1489:
1378:
886:
879:
688:
315:
152:
82:
77:
62:
57:
47:
1524:
1392:
1350:
1323:
1264:
1257:
1250:
1175:
607:
267:
97:
72:
1364:
1051:
835:
784:
694:
682:
162:
142:
1466:
474:
1371:
590:
338:
270:
1470:
643:
639:
289:. It is known only a second upper molar found at the
1291:
1202:
1192:
1181:
1171:
1072:
1057:
1040:
1030:
1019:
1009:
999:
988:
905:
875:
865:
854:
831:
821:
810:
800:
790:
780:
770:
760:
726:
548:
1479:
1340:
1306:
1278:
1240:
1216:
1156:
1071:
969:
896:
845:
747:
717:
521:Palaeogeography, Palaeoclimatology, Palaeoecology
330:. The site has been radiometrically dated to the
655:
8:
486:
484:
1467:
1284:
1222:
1164:
981:
977:
753:
714:
710:
662:
648:
640:
453:
451:
449:
447:
445:
373:and metaconule cusps are extremely small.
20:
624:
606:
441:
318:and only known specimen, catalogued as
7:
428:shows some specializations towards
14:
1591:Rupelian genus first appearances
1566:Prehistoric mammals of Australia
1454:
1446:
1439:
393:would have been the thylacinids
119:
324:Riversleigh World Heritage Area
297:, containing only one species,
291:Riversleigh World Heritage Area
1:
568:10.1080/03115518.2022.2152096
463:Australian Journal of Zoology
1596:Prehistoric marsupial genera
533:10.1016/j.palaeo.2009.02.025
337:The genus name combines the
16:Extinct species of marsupial
1576:Tortonian genus extinctions
1612:
1437:
1287:
1225:
1167:
984:
980:
756:
713:
709:
677:
405:, and the thylacoleonids
238:
231:
116:Scientific classification
114:
23:
506:10.1016/j.gr.2014.10.004
1561:Prehistoric thylacines
1511:Maximucinus muirheadae
1481:Maximucinus muirheadae
300:Maximucinus muirheadae
277:that lived during the
245:Maximucinus muirheadae
1533:Paleobiology Database
1571:Oligocene marsupials
560:2023Alch...47..506G
1581:Miocene marsupials
608:10.7717/peerj.7457
307:History and naming
224:M. muirheadae
1586:Riversleigh fauna
1548:
1547:
1473:Taxon identifiers
1464:
1463:
1435:
1434:
1431:
1430:
1427:
1426:
1423:
1422:
1274:
1273:
1152:
1151:
965:
964:
961:
960:
494:Gondwana Research
408:Wakaleo oldfieldi
334:, ~14.2-12.9 Ma.
259:
258:
1603:
1541:
1540:
1528:
1527:
1515:
1514:
1513:
1500:
1499:
1498:
1468:
1459:
1458:
1451:
1450:
1443:
1293:
1285:
1223:
1204:
1194:
1183:
1173:
1165:
1074:
1059:
1042:
1032:
1021:
1011:
1001:
990:
982:
978:
907:
877:
867:
856:
833:
823:
812:
802:
792:
782:
772:
762:
754:
728:
715:
711:
664:
657:
650:
641:
631:
630:
628:
610:
586:
580:
579:
543:
537:
536:
516:
510:
509:
488:
479:
478:
455:
418:Lekaneleo myersi
326:, north-western
247:
243:
207:
194:
124:
123:
109:
39:
29:Temporal range:
21:
1611:
1610:
1606:
1605:
1604:
1602:
1601:
1600:
1551:
1550:
1549:
1544:
1536:
1531:
1523:
1518:
1509:
1508:
1503:
1494:
1493:
1488:
1475:
1465:
1460:
1453:
1445:
1419:
1407:Pseudantechinus
1336:
1302:
1270:
1236:
1212:
1161:
1148:
1067:
974:
957:
931:Microperoryctes
892:
841:
749:Peramelemorphia
743:
705:
701:Australidelphia
673:
668:
637:
635:
634:
588:
587:
583:
545:
544:
540:
518:
517:
513:
490:
489:
482:
475:10.1071/ZO01044
457:
456:
443:
438:
413:W. vanderleueri
382:
359:
309:
281:in what is now
255:
249:
241:
240:
227:
205:
192:
118:
110:
108:
107:
106:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
35:14.2β12.9
34:
33:
27:
17:
12:
11:
5:
1609:
1607:
1599:
1598:
1593:
1588:
1583:
1578:
1573:
1568:
1563:
1553:
1552:
1546:
1545:
1543:
1542:
1529:
1516:
1501:
1485:
1483:
1477:
1476:
1471:
1462:
1461:
1438:
1436:
1433:
1432:
1429:
1428:
1425:
1424:
1421:
1420:
1418:
1417:
1410:
1403:
1396:
1389:
1382:
1375:
1368:
1361:
1354:
1346:
1344:
1338:
1337:
1335:
1334:
1327:
1320:
1312:
1310:
1304:
1303:
1301:
1300:
1288:
1282:
1276:
1275:
1272:
1271:
1269:
1268:
1261:
1254:
1246:
1244:
1238:
1237:
1235:
1234:
1226:
1220:
1214:
1213:
1211:
1210:
1200:
1190:
1179:
1168:
1162:
1157:
1154:
1153:
1150:
1149:
1147:
1146:
1139:
1132:
1125:
1118:
1111:
1103:
1096:
1089:
1080:
1078:
1069:
1068:
1066:
1065:
1055:
1048:
1038:
1028:
1017:
1007:
997:
985:
975:
972:Dasyuromorphia
970:
967:
966:
963:
962:
959:
958:
956:
955:
948:
941:
934:
927:
920:
913:
902:
900:
894:
893:
891:
890:
883:
873:
863:
851:
849:
843:
842:
840:
839:
829:
819:
808:
798:
788:
778:
768:
757:
751:
745:
744:
742:
741:
734:
723:
721:
707:
706:
704:
703:
697:
691:
685:
678:
675:
674:
669:
667:
666:
659:
652:
644:
633:
632:
581:
554:(4): 506β521.
538:
527:(1β4): 24β37.
511:
500:(1): 153β167.
480:
469:(6): 603β314.
440:
439:
437:
434:
430:hypercarnivory
381:
378:
358:
355:
345:, large, with
332:Middle Miocene
308:
305:
279:Middle Miocene
257:
256:
250:
236:
235:
229:
228:
220:
218:
214:
213:
203:
199:
198:
190:
186:
185:
183:Dasyuromorphia
180:
176:
175:
170:
166:
165:
160:
156:
155:
150:
146:
145:
140:
136:
135:
130:
126:
125:
112:
111:
103:
102:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
40:
31:Middle Miocene
28:
15:
13:
10:
9:
6:
4:
3:
2:
1608:
1597:
1594:
1592:
1589:
1587:
1584:
1582:
1579:
1577:
1574:
1572:
1569:
1567:
1564:
1562:
1559:
1558:
1556:
1539:
1534:
1530:
1526:
1521:
1517:
1512:
1506:
1502:
1497:
1491:
1487:
1486:
1484:
1482:
1478:
1474:
1469:
1457:
1452:
1449:
1442:
1416:
1415:
1411:
1409:
1408:
1404:
1402:
1401:
1400:Phascolosorex
1397:
1395:
1394:
1393:Parantechinus
1390:
1388:
1387:
1386:Neophascogale
1383:
1381:
1380:
1376:
1374:
1373:
1369:
1367:
1366:
1362:
1360:
1359:
1355:
1353:
1352:
1348:
1347:
1345:
1343:
1339:
1333:
1332:
1328:
1326:
1325:
1321:
1319:
1318:
1314:
1313:
1311:
1309:
1305:
1298:
1297:
1296:Whollydooleya
1290:
1289:
1286:
1283:
1281:
1277:
1267:
1266:
1262:
1260:
1259:
1255:
1253:
1252:
1248:
1247:
1245:
1243:
1239:
1233:
1232:
1228:
1227:
1224:
1221:
1219:
1218:Sminthopsinae
1215:
1209:
1208:
1201:
1199:
1198:
1191:
1188:
1187:
1180:
1178:
1177:
1170:
1169:
1166:
1163:
1160:
1155:
1145:
1144:
1140:
1138:
1137:
1133:
1131:
1130:
1126:
1124:
1123:
1119:
1117:
1116:
1112:
1109:
1108:
1104:
1102:
1101:
1097:
1095:
1094:
1090:
1087:
1086:
1082:
1081:
1079:
1077:
1070:
1064:
1063:
1056:
1054:
1053:
1049:
1047:
1046:
1039:
1037:
1036:
1029:
1026:
1025:
1018:
1016:
1015:
1008:
1006:
1005:
998:
995:
994:
987:
986:
983:
979:
976:
973:
968:
954:
953:
949:
947:
946:
942:
940:
939:
935:
933:
932:
928:
926:
925:
921:
919:
918:
914:
912:
911:
904:
903:
901:
899:
895:
889:
888:
884:
882:
881:
874:
872:
871:
864:
861:
860:
853:
852:
850:
848:
847:Thylacomyidae
844:
838:
837:
830:
828:
827:
820:
817:
816:
809:
807:
806:
799:
797:
796:
789:
787:
786:
779:
777:
776:
769:
767:
766:
759:
758:
755:
752:
750:
746:
740:
739:
735:
733:
732:
725:
724:
722:
720:
716:
712:
708:
702:
698:
696:
692:
690:
686:
684:
680:
679:
676:
672:
665:
660:
658:
653:
651:
646:
645:
642:
638:
627:
622:
618:
614:
609:
604:
600:
596:
592:
585:
582:
577:
573:
569:
565:
561:
557:
553:
549:
542:
539:
534:
530:
526:
522:
515:
512:
507:
503:
499:
495:
487:
485:
481:
476:
472:
468:
464:
460:
454:
452:
450:
448:
446:
442:
435:
433:
431:
427:
424:The molar of
422:
420:
419:
414:
410:
409:
404:
403:
398:
397:
392:
387:
379:
377:
374:
372:
368:
363:
356:
354:
352:
351:Ancient Greek
348:
344:
340:
335:
333:
329:
325:
321:
317:
313:
306:
304:
302:
301:
296:
292:
288:
284:
280:
276:
272:
269:
265:
264:
253:
248:
246:
237:
234:
233:Binomial name
230:
226:
225:
219:
216:
215:
212:
211:
204:
201:
200:
197:
191:
188:
187:
184:
181:
178:
177:
174:
171:
168:
167:
164:
161:
158:
157:
154:
151:
148:
147:
144:
141:
138:
137:
134:
131:
128:
127:
122:
117:
113:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
38:
32:
26:
22:
19:
1480:
1444:
1412:
1405:
1398:
1391:
1384:
1377:
1370:
1363:
1356:
1349:
1329:
1322:
1315:
1308:Phascogalini
1294:
1263:
1256:
1251:Antechinomys
1249:
1242:Sminthopsini
1229:
1205:
1195:
1184:
1174:
1141:
1136:Tyarrpecinus
1134:
1127:
1120:
1115:Ngamalacinus
1113:
1107:Mutpuracinus
1105:
1098:
1092:
1091:
1083:
1076:Thylacinidae
1060:
1050:
1043:
1035:Malleodectes
1033:
1022:
1012:
1002:
991:
952:Rhynchomeles
950:
943:
936:
929:
922:
915:
908:
885:
878:
868:
857:
834:
824:
813:
803:
795:Kutjamarcoot
793:
783:
773:
763:
736:
731:Naraboryctes
729:
719:Notoryctidae
699:Superorder:
636:
598:
594:
584:
551:
547:
541:
524:
520:
514:
497:
493:
466:
462:
425:
423:
416:
412:
406:
400:
394:
390:
385:
383:
380:Paleobiology
375:
361:
360:
346:
342:
336:
319:
311:
310:
299:
298:
262:
261:
260:
244:
239:
223:
222:
209:
208:
196:Thylacinidae
169:Infraclass:
24:
18:
1505:Wikispecies
1414:Sarcophilus
1365:Dasyuroides
1265:Sminthopsis
1143:Wabulacinus
1100:Muribacinus
1093:Maximucinus
1052:Myrmecobius
1045:Mayigriphus
1004:Dasylurinja
993:Ankotarinja
898:Peramelidae
826:Silvicultor
671:Agreodontia
426:Maximucinus
396:Muribacinus
391:Maximucinus
386:Maximucinus
384:Fossils of
371:protoconule
362:Maximucinus
357:Description
312:Maximucinus
263:Maximucinus
210:Maximucinus
173:Marsupialia
25:Maximucinus
1555:Categories
1358:Dasykaluta
1351:Dasycercus
1331:Phascogale
1317:Antechinus
1280:Dasyurinae
1186:Ganbulanyi
1159:Dasyuridae
1129:Thylacinus
1122:Nimbacinus
1014:Joculusium
945:Peroryctes
917:Echymipera
815:Numbigilga
738:Notoryctes
436:References
402:Nimbacinus
328:Queensland
283:Queensland
275:thylacinid
1342:Dasyurini
1231:Planigale
1197:Glaucodon
1085:Badjcinus
1062:Wakamatha
938:Perameles
870:Ischnodon
775:Chaeropus
681:Kingdom:
617:2167-8359
601:: e7457.
576:256157821
320:QM F30331
295:monotypic
287:Australia
217:Species:
139:Kingdom:
133:Eukaryota
1525:11186801
1496:Q6795826
1490:Wikidata
1379:Myoictis
1372:Dasyurus
1207:Urrayira
887:Macrotis
880:Liyamayi
859:Bulbadon
695:Mammalia
689:Chordata
687:Phylum:
683:Animalia
459:Wroe, S.
367:cingulum
316:holotype
189:Family:
163:Mammalia
153:Chordata
149:Phylum:
143:Animalia
129:Domain:
1324:Murexia
1258:Ningaui
1176:Barinya
924:Isoodon
765:Bulungu
693:Class:
626:6727838
556:Bibcode
343:maximus
268:extinct
242:†
221:†
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