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The palate has a pronounced, sagittally elongated vault that is formed primarily by the vomers. The vomers contact the premaxillae at a level with the maxillary tooth margin towards the tip of the rostrum. However, it does curve strongly dorsally towards the palatines, and so we see the choanae very
539:
were ankylosed to the jaws. Although there are indications of deep implantation, it is still uncertain whether the mode of attachment could be described as thecodont. Looking at the maxillary teeth, a circular cross-section is maintained throughout the teeth and the majority are worn out heavily to
517:
The postorbital is composed of the anteroventral, posterodorsal, and dorsomedial processes in equal length. The anteroventral process does not come into contact with the ectopterygoid but it does indeed reach farther down the medial side of the jugal. The posterodorsal process forms a great portion
495:
has a broadly triangular shape with a wide temporal region that tapers sharply along the orbits, expands abruptly at the prefrontals, and then tapers to the blunt rostrum. There is a large, median external naris located at the front of the rostrum that faces dorsally and cranially. The orbits face
513:
On the maxilla, there is a distinct notch that contributes most of the border of a dorsal fenestra. It appeared to be the cranialmost of a series of maxillary foramina that extended across the lateral face of the maxilla. This most likely hinted to cutaneous blood vessels and nerves in that area.
551:
In 1963, M.E. Malan observed an interesting pattern in the positioning of the maxillary and dentary teeth. The middle section, where the medial expansion of the maxilla is wideset, had a zig-zag arrangement of teeth while the first and last four teeth are aligned in a row that is parallel to the
478:
with multiple rows of maxillary and dentary teeth, with each row consisting of only a small number of teeth. The two premaxillary teeth are approximately twice the size of the maxillary teeth. The maxillary teeth are inset medially and project below the internal naris. There is also an occlusion
444:
SAM 7416, another paratype, consists of an articulated vertebral column composed of the last dozen presacrals, both sacrals and at least the first 15 caudal vertebrae, fragments of right forelimb, pelvic girdle, complete right femur, right crus and partial left crus, and right and left tarsi and
364:
near the town of Aliwal North in the Cape
Province of South Africa. The block of sandstone contained many intermingled partial skeletons of several small reptiles, and after careful sorting, Watson considered the unidentified skeletons to belong to a single species, which he named
440:
SAM 6536, another paratype, consists of a virtually complete skull with lower jaws, articulated cervical vertebrae and ribs, dorsal vertebrae and ribs, complete left scapulocoracoid and partial right scapula, interclavicle, clavicles, distal end of left humerus, and gastralia.
399:
looked more closely at the partial skeletons and immediately determined it to be in fact skeletons of two distinct, though related, species. Broom designated an articulated skeleton with a single external naris and a pair of supposed acrodont premaxillary teeth as the type of
521:
Information of the lower jaw had to be put together from the pieces of the different SAM fossils as there is not one that had a complete lower jaw. When looking at the different parts however, there are indications of at least six elements present in the lower jaw of the
483:. Looking at the spine, the length of the axis neural spine is greater than the length of axis centrum. It also has a craniocaudally narrow neural spine of the third cervical and a prominent midventral groove on the first two caudal centra.
437:, consists of an incomplete right maxilla, an articulated series of the last ten presacrals, both sacrals, and first six caudals, partial forelimbs, left and right pelvic girdles, right hind limb, as well as element of left tarsus.
426:, consists of a partial rostrum, palate, braincase, lower jaws, sections of articulated presacral vertebral column, nine articulated caudal vertebrae, portions of scapula and pelvic girdle, and partial forelimb and hindlimbs.
534:
Evident on SAM 5882, 6046, and 6536 was a clear line of separation between the tooth and surrounding bone indicating that the teeth are implanted in the jaw, which went against the previous idea that the marginal teeth of
2499:
1826:
2209:
1795:
3988:
3688:
2466:
2504:
4057:
997:"A New Species of Basal Rhynchosaur (Diapsida: Archosauromorpha) from the Early Middle Triassic of South Africa, and the Early Evolution of Rhynchosauria"
2471:
510:
far above the tooth margin. To withdraw the choana further from the oral cavity, the vomerine and palatine borders of the choana are recessed dorsally.
3975:
1333:
4077:
4072:
1788:
1298:
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2310:
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could have had multiple teeth on at least the maxilla. This hypothesis was supported when examining the erosional features on SAM 6536.
506:, is the presence of a beak-like rostrum, formed primarily by huge premaxillary teeth rather than by tapering, edentulous premaxillae.
3693:
2245:
4067:
3911:
3790:
1781:
1130:"The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms"
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1741:
4062:
3745:
3683:
3080:
350:
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between the vomerine teeth and dentary teeth with saddle-shaped vomers that overhang the dorsal side of the premaxillary
3646:
3341:
3085:
2509:
226:
3336:
3070:
1326:
358:
1257:
Sues, Hans-Dieter (2005). "Evolution of
Herbivory in Terrestrial Vertebrates: Perspectives from the Fossil Record".
138:
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2354:
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314:
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maxilla. In the dentary, a simpler arrangement with only a slight zig-zag arrangement. Comparing this feature to
3386:
2255:
1906:
1842:
1088:"The Early Triassic Rhynchosaur Mesosuchus Browni and the Interrelationships of Basal Archosauromorph Reptiles"
474:
pertain to the skull due to the relatively large amount of skull material known. It was a small rhynchosaurian
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1992:
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of the smooth cranial rim of the upper temporal fenestra and contacts the parietal below the postfrontal.
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was an ancestral crocodile with close affinities to other presumed primitive crocodilians such as
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2016:
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laterally and slightly cranially. One feature on the skull that can distinguish the
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This article contains quotations from this source, which is available under the
653:
602:
283:
206:
43:
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1184:
404:, and the remainder of the specimens were assigned to a new genus and species,
3631:
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3466:
3451:
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3426:
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3280:
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3165:
3125:
3115:
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2698:
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2412:
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1951:
1722:
1598:
1406:
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969:
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88:
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1155:
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in 1912 after examining a block of sandstone kept in a private collection of
3579:
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3220:
3039:
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2944:
2899:
2879:
2839:
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2638:
2617:
2601:
2540:
2535:
2224:
1390:
1293:
574:
545:
480:
370:
313:, and from the paratypes SAM 6046, SAM 6536, SAM 7416 and SAM 7701 from the
150:
93:
37:
1243:
1173:
1099:
117:
1773:
558:, which also possessed a zig-zag pattern of teeth, they hypothesized that
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83:
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526:: dentary, splenial, surangular, angular, prearticular, and articular.
502:
475:
291:
273:
180:
98:
73:
1013:
996:
3461:
1360:
514:
This notch is hidden by an overlap of the premaxilla by the maxilla.
160:
3937:
1272:
Martin D. Ezcurra; Felipe C. Montefeltro; Richard J. Butler (2016).
1199:"The Relationships of the Euparkeriidae and the Rise of Archosauria"
16:
Extinct genus of reptiles from early Middle
Triassic of South Africa
1041:
Watson, David M.S. (1913). "Mesosuchus Browni, Gen. Et Spec. Nov".
276:
1311:
3941:
3839:
3811:
3667:
2488:
2401:
2163:
1815:
1777:
1353:
1315:
1060:
Ezcurra, Martin (2016). "The Early
Evolution of Rhynchosaurs".
1185:
Creative
Commons Attribution 4.0 International (CC BY 4.0)
2500:
List of fossiliferous stratigraphic units in South Africa
1827:
List of fossiliferous stratigraphic units in South Africa
500:
from all other rhynchosaurs, with possible exception of
353:. This block was found in the middle deposits of the
325:
is quite small, spanning around 30 cm in length.
3950:
3718:
3702:
3619:
3588:
3567:
3551:
3500:
3354:
3329:
3098:
3053:
2822:
2631:
2610:
2549:
2518:
2373:
2342:
2309:
2238:
2210:
Tectonic evolution of the
Barberton greenstone belt
2135:
1962:
1876:
1835:
1663:
1615:
1563:
1510:
1459:
1432:
1405:
1092:Philosophical Transactions of the Royal Society
466:, as it is the only known species. Most of the
3689:Geological Commission of the Cape of Good Hope
2467:Department of Mineral Resources (South Africa)
2505:Archean life in the Barberton Greenstone Belt
1789:
1327:
1123:
1121:
1119:
8:
373:of the name suggests, Watson believed that
3938:
3918:
3836:
3808:
3699:
3664:
3326:
3050:
2515:
2485:
2472:Department of Mineral Resources and Energy
2398:
2235:
2160:
1832:
1812:
1796:
1782:
1774:
1569:
1516:
1507:
1438:
1411:
1402:
1350:
1334:
1320:
1312:
116:
20:
1292:
1233:
1163:
1145:
1107:
1022:
1012:
1001:Zoological Journal of the Linnean Society
987:
540:short, blunt pegs. This suggests that
1274:"The Early Evolution of Rhynchosaurs"
7:
1081:
1079:
1077:
1075:
1073:
1071:
1036:
1034:
853:
829:
802:
778:
754:
706:
696:
672:
594:
584:
4058:Middle Triassic reptiles of Africa
3694:Geological Society of South Africa
1281:Frontiers in Ecology and Evolution
1062:Frontiers in Ecology and Evolution
14:
2246:List of volcanoes in South Africa
3917:
3906:
3905:
3893:
3879:
3791:Eduard Meine van Zinderen-Bakker
1758:
1757:
1740:
1304:from the original on 2024-02-20.
1178:
972:
137:
3076:Pristerognathus Assemblage Zone
3684:Evolutionary Studies Institute
3081:Tapinocephalus Assemblage Zone
1:
4078:Taxa named by D. M. S. Watson
4073:Fossil taxa described in 1912
3066:Daptocephalus Assemblage Zone
3061:Cistecephalus Assemblage Zone
2462:Minerals Council South Africa
2319:1620 Robben Island earthquake
3669:Research and administration
3342:Lystrosaurus Assemblage Zone
3086:Tropidostoma Assemblage Zone
1043:Records of the Albany Museum
347:David Meredith Seares Watson
329:was discovered and named by
294:stage) deposits of Eastern
3337:Cynognathus Assemblage Zone
3071:Eodicynodon Assemblage Zone
1128:Ezcurra, Martín D. (2016).
1094:. 353.1368 (1368): 501–41.
462:come from the diagnosis of
272:("middle crocodile") is an
4094:
4048:Prehistoric reptile genera
2365:Vredefort impact structure
2355:Morokweng impact structure
2329:2006 Mozambique earthquake
1206:Royal Society Open Science
3873:
3846:
3835:
3818:
3807:
3766:Daniel Rossouw Kannemeyer
3674:
3663:
3647:Waterloo Farm lagerstätte
2510:Waterloo Farm lagerstätte
2495:
2484:
2408:
2397:
2291:Barberton Greenstone Belt
2170:
2159:
1822:
1811:
1753:
1738:
1572:
1519:
1506:
1487:
1441:
1414:
1401:
1349:
875:
858:
851:
834:
827:
807:
800:
783:
776:
759:
752:
726:
711:
704:
694:
677:
670:
650:
633:
616:
599:
592:
458:All diagnostic traits of
242:
237:
134:Scientific classification
132:
124:
115:
23:
3387:Coelophysis rhodesiensis
2256:Pilanesberg Game Reserve
1907:Table Mountain Sandstone
1843:Bushveld Igneous Complex
1197:Sookias, Roland (2016).
995:Butler, Richard (2015).
4068:Fossils of South Africa
3900:South Africa portal
3736:Charles Kimberlin Brain
2324:1969 Tulbagh earthquake
2039:Abrahamskraal Formation
1993:Sundays River Formation
1917:Abrahamskraal Formation
1805:Geology of South Africa
1294:10.3389/fevo.2015.00142
1259:Cambridge: Cambridge UP
302:. It is known from the
3751:Anusuya Chinsamy-Turan
3679:Council for Geoscience
3642:West Coast Fossil Park
2334:2014 Orkney earthquake
2215:Terra Australis Orogen
2165:Tectonics and orogeny
1978:Buffelskloof Formation
1100:10.1098/rstb.1998.0225
4063:Triassic South Africa
4015:Paleobiology Database
2261:Prince Edward Islands
1817:Geological formations
1086:Dilkes, D.W. (1998).
429:SAM 6046, one of the
355:Burgersdorp Formation
3823:Geology of Cape Town
3781:John Talbot Robinson
3731:Lieuwe Dirk Boonstra
2437:Coal in South Africa
2428:Banket (mining term)
2125:Normandien Formation
1868:Transvaal Supergroup
714:Mariante rhynchosaur
395:. However, in 1913,
345:were first found by
337:Discovery and naming
125:Life restoration of
3786:George William Stow
3627:Cradle of Humankind
2381:Bellsbank meteorite
2274:Finsch diamond mine
2230:Witwatersrand Basin
2205:Makhonjwa Mountains
2076:Whitehill Formation
2054:Middleton Formation
1942:Whitehill Formation
1932:Ganigobis Formation
1892:Ganigobis Formation
1226:10.1098/rsos.150674
1218:2016RSOS....350674S
980:Paleontology portal
407:Euparkeria capensis
3886:Geology portal
3761:Alfred W. Crompton
3726:Andrew Geddes Bain
3010:Pristerognathoides
2450:Titanium in Africa
2445:Platinum in Africa
2441:Iron ore in Africa
2284:Koffiefontein mine
2279:Jagersfontein Mine
2190:Gondwanide orogeny
2088:Bushveld Sandstone
2059:Teekloof Formation
1988:Kirkwood Formation
1937:Teekloof Formation
1848:Fig Tree Formation
1695:Macrocephalosaurus
1665:Hyperodapedontinae
1617:Stenaulorhynchinae
1147:10.7717/peerj.1778
824:Hyperodapedontinae
701:Stenaulorhynchinae
4030:
4029:
4002:Open Tree of Life
3944:Taxon identifiers
3935:
3934:
3869:
3868:
3851:Pothole (geology)
3831:
3830:
3813:Regional articles
3803:
3802:
3799:
3798:
3756:H. Basil S. Cooke
3659:
3658:
3655:
3654:
3432:Heterodontosaurus
3350:
3349:
3094:
3093:
3025:Promoschorhynchus
2480:
2479:
2455:Uranium in Africa
2403:Mineral deposits
2393:
2392:
2389:
2388:
2155:
2154:
2151:
2150:
2120:Molteno Formation
2115:Katberg Formation
2093:Clarens Formation
2049:Katberg Formation
2044:Balfour Formation
2012:Drakensberg Group
2000:Kalahari Deposits
1922:Balfour Formation
1853:Komatii Formation
1771:
1770:
1749:
1748:
1736:
1735:
1732:
1731:
1502:
1501:
1498:
1497:
1483:
1482:
1479:
1478:
1014:10.1111/zoj.12246
962:
961:
953:
952:
944:
943:
935:
934:
926:
925:
917:
916:
908:
907:
899:
898:
890:
889:
741:
740:
577:based on Ezcurra
555:Captorhinus aguti
472:Mesosuchus browni
470:used to identify
464:Mesosuchus browni
367:Mesosuchus browni
265:
264:
259:
233:
4085:
4023:
4022:
4010:
4009:
3997:
3996:
3984:
3983:
3971:
3970:
3969:
3939:
3921:
3920:
3909:
3908:
3898:
3897:
3896:
3884:
3883:
3837:
3809:
3700:
3665:
3327:
3051:
2980:Macroscelesaurus
2860:Cistecephaloides
2845:Blattoidealestes
2669:Cistecephaloides
2572:Deltacephalaspis
2516:
2486:
2399:
2236:
2161:
2110:Elliot Formation
2098:Elliot Formation
2029:Karoo Supergroup
1863:Onverwacht Group
1833:
1813:
1798:
1791:
1784:
1775:
1761:
1760:
1744:
1654:Stenaulorhynchus
1570:
1517:
1508:
1449:Archosauromorpha
1439:
1434:Archosauromorpha
1412:
1403:
1396:
1395:
1382:Archosauromorpha
1351:
1336:
1329:
1322:
1313:
1306:
1305:
1303:
1296:
1278:
1269:
1263:
1262:
1254:
1248:
1247:
1237:
1203:
1194:
1188:
1182:
1177:
1167:
1149:
1125:
1114:
1113:
1111:
1083:
1066:
1065:
1057:
1051:
1050:
1038:
1029:
1028:
1026:
1016:
992:
982:
977:
976:
975:
854:
830:
803:
779:
755:
730:Stenaulorhynchus
707:
697:
673:
595:
585:
414:Fossil specimens
309:5882, a partial
253:
225:
218:
205:
194:Archosauromorpha
142:
141:
120:
110:
40:
29:Temporal range:
21:
4093:
4092:
4088:
4087:
4086:
4084:
4083:
4082:
4033:
4032:
4031:
4026:
4018:
4013:
4005:
4000:
3992:
3987:
3979:
3974:
3965:
3964:
3959:
3946:
3936:
3931:
3894:
3892:
3878:
3865:
3856:Steep structure
3842:
3827:
3814:
3795:
3719:Paleontologists
3714:
3698:
3670:
3651:
3615:
3611:Vulpes skinneri
3584:
3563:
3547:
3543:Ptychoceratodus
3508:Acanthohoplites
3496:
3472:Ptychoceratodus
3346:
3325:
3276:Ptychoceratodus
3251:Platycraniellus
3176:Ictidosuchoides
3090:
3049:
3030:Scalopocephalus
3015:Pristerognathus
2930:Homodontosaurus
2818:
2744:Ictidosuchoides
2684:Dicynodontoides
2664:Charassognathus
2627:
2606:
2582:Gondwanascorpio
2545:
2514:
2491:
2476:
2404:
2385:
2369:
2338:
2305:
2234:
2220:Transvaal Basin
2200:Kalahari Craton
2195:Kaapvaal Craton
2166:
2147:
2131:
2083:Stormberg Group
2022:Uitenhage Group
1973:Uitenhage Group
1958:
1902:Bokkeveld Group
1872:
1831:
1818:
1807:
1802:
1772:
1767:
1745:
1728:
1659:
1633:Brasinorhynchus
1611:
1565:Rhynchosauridae
1559:
1494:
1475:
1455:
1428:
1397:
1356:
1355:
1345:
1340:
1310:
1309:
1301:
1276:
1271:
1270:
1266:
1256:
1255:
1251:
1201:
1196:
1195:
1191:
1127:
1126:
1117:
1085:
1084:
1069:
1059:
1058:
1054:
1040:
1039:
1032:
994:
993:
989:
978:
973:
971:
968:
963:
954:
945:
936:
927:
918:
909:
900:
891:
742:
667:Rhynchosauridae
568:
532:
489:
456:
451:
416:
362:Assemblage Zone
339:
331:D. M. S. Watson
288:Middle Triassic
224:
216:
203:
136:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
31:Middle Triassic
27:
17:
12:
11:
5:
4091:
4089:
4081:
4080:
4075:
4070:
4065:
4060:
4055:
4053:Anisian genera
4050:
4045:
4035:
4034:
4028:
4027:
4025:
4024:
4011:
3998:
3985:
3972:
3956:
3954:
3948:
3947:
3942:
3933:
3932:
3930:
3929:
3922:
3915:
3903:
3889:
3874:
3871:
3870:
3867:
3866:
3864:
3863:
3858:
3853:
3847:
3844:
3843:
3840:
3833:
3832:
3829:
3828:
3826:
3825:
3819:
3816:
3815:
3812:
3805:
3804:
3801:
3800:
3797:
3796:
3794:
3793:
3788:
3783:
3778:
3776:James Kitching
3773:
3768:
3763:
3758:
3753:
3748:
3743:
3738:
3733:
3728:
3722:
3720:
3716:
3715:
3713:
3712:
3706:
3704:
3697:
3696:
3691:
3686:
3681:
3675:
3672:
3671:
3668:
3661:
3660:
3657:
3656:
3653:
3652:
3650:
3649:
3644:
3639:
3634:
3629:
3623:
3621:
3617:
3616:
3614:
3613:
3608:
3606:Panthera shawi
3603:
3598:
3592:
3590:
3586:
3585:
3583:
3582:
3577:
3571:
3569:
3565:
3564:
3562:
3561:
3555:
3553:
3549:
3548:
3546:
3545:
3540:
3535:
3530:
3528:Macroscaphites
3525:
3520:
3515:
3510:
3504:
3502:
3498:
3497:
3495:
3494:
3489:
3484:
3482:Sefapanosaurus
3479:
3474:
3469:
3464:
3459:
3457:Massospondylus
3454:
3449:
3444:
3439:
3434:
3429:
3424:
3419:
3414:
3409:
3404:
3402:Erythrotherium
3399:
3394:
3389:
3384:
3379:
3374:
3369:
3364:
3358:
3356:
3352:
3351:
3348:
3347:
3345:
3344:
3339:
3333:
3331:
3324:
3323:
3321:Zorillodontops
3318:
3313:
3308:
3303:
3298:
3293:
3288:
3283:
3278:
3273:
3268:
3263:
3258:
3253:
3248:
3246:Plateosauravus
3243:
3238:
3233:
3231:Olivierosuchus
3228:
3223:
3218:
3213:
3211:Microgomphodon
3208:
3203:
3201:Melanorosaurus
3198:
3193:
3188:
3183:
3178:
3173:
3168:
3163:
3158:
3153:
3151:Erythrotherium
3148:
3143:
3138:
3133:
3128:
3123:
3118:
3113:
3108:
3106:Batrachosuchus
3102:
3100:
3096:
3095:
3092:
3091:
3089:
3088:
3083:
3078:
3073:
3068:
3063:
3057:
3055:
3048:
3047:
3045:Uranocentrodon
3042:
3037:
3032:
3027:
3022:
3017:
3012:
3007:
3002:
2997:
2992:
2987:
2982:
2977:
2972:
2967:
2962:
2960:Lanthanostegus
2957:
2952:
2947:
2942:
2937:
2932:
2927:
2922:
2917:
2912:
2907:
2902:
2897:
2892:
2890:Cynariognathus
2887:
2882:
2877:
2872:
2867:
2862:
2857:
2852:
2850:Brachyprosopus
2847:
2842:
2837:
2832:
2826:
2824:
2823:Beaufort Group
2820:
2819:
2817:
2816:
2811:
2806:
2801:
2796:
2791:
2786:
2781:
2776:
2771:
2766:
2761:
2756:
2751:
2746:
2741:
2736:
2731:
2726:
2721:
2716:
2711:
2706:
2701:
2696:
2691:
2686:
2681:
2676:
2671:
2666:
2661:
2656:
2651:
2646:
2644:Alopecognathus
2641:
2635:
2633:
2629:
2628:
2626:
2625:
2620:
2614:
2612:
2608:
2607:
2605:
2604:
2599:
2594:
2589:
2584:
2579:
2574:
2569:
2564:
2559:
2553:
2551:
2547:
2546:
2544:
2543:
2538:
2533:
2531:Onychopterella
2528:
2522:
2520:
2513:
2512:
2507:
2502:
2496:
2493:
2492:
2489:
2482:
2481:
2478:
2477:
2475:
2474:
2469:
2464:
2458:
2457:
2452:
2447:
2442:
2439:
2433:
2432:
2431:
2430:
2420:
2415:
2409:
2406:
2405:
2402:
2395:
2394:
2391:
2390:
2387:
2386:
2384:
2383:
2377:
2375:
2371:
2370:
2368:
2367:
2362:
2360:Tswaing crater
2357:
2352:
2350:Kalkkop crater
2346:
2344:
2343:Impact craters
2340:
2339:
2337:
2336:
2331:
2326:
2321:
2315:
2313:
2307:
2306:
2304:
2303:
2298:
2293:
2288:
2287:
2286:
2281:
2276:
2271:
2263:
2258:
2253:
2248:
2242:
2240:
2233:
2232:
2227:
2222:
2217:
2212:
2207:
2202:
2197:
2192:
2187:
2185:Ellisras Basin
2182:
2180:Cape Fold Belt
2177:
2171:
2168:
2167:
2164:
2157:
2156:
2153:
2152:
2149:
2148:
2146:
2145:
2139:
2137:
2133:
2132:
2130:
2129:
2128:
2127:
2122:
2117:
2112:
2104:
2103:
2102:
2101:
2100:
2095:
2090:
2080:
2079:
2078:
2068:
2063:
2062:
2061:
2056:
2051:
2046:
2041:
2034:Beaufort Group
2026:
2025:
2024:
2019:
2017:Enon Formation
2014:
2005:
2004:
2003:
2002:
1997:
1996:
1995:
1990:
1985:
1983:Enon Formation
1980:
1966:
1964:
1960:
1959:
1957:
1956:
1955:
1954:
1946:
1945:
1944:
1939:
1934:
1929:
1924:
1919:
1911:
1910:
1909:
1904:
1896:
1895:
1894:
1889:
1883:Carboniferous
1880:
1878:
1874:
1873:
1871:
1870:
1865:
1860:
1855:
1850:
1845:
1839:
1837:
1836:Palaeoarchaean
1830:
1829:
1823:
1820:
1819:
1816:
1809:
1808:
1803:
1801:
1800:
1793:
1786:
1778:
1769:
1768:
1766:
1765:
1754:
1751:
1750:
1747:
1746:
1739:
1737:
1734:
1733:
1730:
1729:
1727:
1726:
1719:
1712:
1705:
1702:Oryctorhynchus
1698:
1691:
1684:
1677:
1669:
1667:
1661:
1660:
1658:
1657:
1650:
1643:
1636:
1629:
1621:
1619:
1613:
1612:
1610:
1609:
1602:
1595:
1588:
1581:
1573:
1567:
1561:
1560:
1558:
1557:
1550:
1543:
1536:
1529:
1520:
1514:
1504:
1503:
1500:
1499:
1496:
1495:
1488:
1485:
1484:
1481:
1480:
1477:
1476:
1474:
1473:
1472:
1471:
1465:
1463:
1457:
1456:
1454:
1453:
1452:
1451:
1442:
1436:
1430:
1429:
1427:
1426:
1425:
1424:
1415:
1409:
1399:
1398:
1394:
1393:
1384:
1375:
1369:
1363:
1354:
1347:
1346:
1341:
1339:
1338:
1331:
1324:
1316:
1308:
1307:
1264:
1249:
1189:
1115:
1067:
1052:
1030:
986:
985:
984:
983:
967:
964:
960:
959:
956:
955:
951:
950:
947:
946:
942:
941:
938:
937:
933:
932:
929:
928:
924:
923:
920:
919:
915:
914:
911:
910:
906:
905:
902:
901:
897:
896:
893:
892:
888:
887:
884:
883:
874:
871:
870:
867:
866:
857:
852:
850:
847:
846:
843:
842:
833:
828:
826:
820:
819:
816:
815:
806:
801:
799:
796:
795:
792:
791:
782:
777:
775:
772:
771:
768:
767:
758:
753:
751:
748:
747:
744:
743:
739:
738:
735:
734:
725:
722:
721:
718:
717:
710:
705:
703:
695:
693:
690:
689:
686:
685:
676:
671:
669:
663:
662:
659:
658:
649:
646:
645:
642:
641:
632:
629:
628:
625:
624:
615:
612:
611:
608:
607:
598:
593:
591:
583:
567:
564:
544:is in fact an
531:
528:
488:
485:
468:autapomorphies
455:
452:
450:
447:
418:SAM 5882, the
415:
412:
338:
335:
263:
262:
261:
260:
254:Watson, 1912 (
240:
239:
235:
234:
214:
210:
209:
201:
197:
196:
191:
184:
183:
178:
174:
173:
168:
164:
163:
158:
154:
153:
148:
144:
143:
130:
129:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
4090:
4079:
4076:
4074:
4071:
4069:
4066:
4064:
4061:
4059:
4056:
4054:
4051:
4049:
4046:
4044:
4041:
4040:
4038:
4021:
4016:
4012:
4008:
4003:
3999:
3995:
3990:
3986:
3982:
3977:
3973:
3968:
3962:
3958:
3957:
3955:
3953:
3949:
3945:
3940:
3928:
3927:
3923:
3916:
3914:
3913:
3904:
3902:
3901:
3890:
3888:
3887:
3882:
3876:
3875:
3872:
3862:
3859:
3857:
3854:
3852:
3849:
3848:
3845:
3838:
3834:
3824:
3821:
3820:
3817:
3810:
3806:
3792:
3789:
3787:
3784:
3782:
3779:
3777:
3774:
3772:
3769:
3767:
3764:
3762:
3759:
3757:
3754:
3752:
3749:
3747:
3744:
3742:
3739:
3737:
3734:
3732:
3729:
3727:
3724:
3723:
3721:
3717:
3711:
3710:Hans Merensky
3708:
3707:
3705:
3701:
3695:
3692:
3690:
3687:
3685:
3682:
3680:
3677:
3676:
3673:
3666:
3662:
3648:
3645:
3643:
3640:
3638:
3637:Scylacorhinus
3635:
3633:
3630:
3628:
3625:
3624:
3622:
3618:
3612:
3609:
3607:
3604:
3602:
3599:
3597:
3594:
3593:
3591:
3587:
3581:
3578:
3576:
3575:Africanacetus
3573:
3572:
3570:
3566:
3560:
3557:
3556:
3554:
3550:
3544:
3541:
3539:
3536:
3534:
3531:
3529:
3526:
3524:
3521:
3519:
3516:
3514:
3511:
3509:
3506:
3505:
3503:
3499:
3493:
3490:
3488:
3485:
3483:
3480:
3478:
3475:
3473:
3470:
3468:
3465:
3463:
3460:
3458:
3455:
3453:
3450:
3448:
3447:Litargosuchus
3445:
3443:
3442:Lesothosaurus
3440:
3438:
3435:
3433:
3430:
3428:
3425:
3423:
3420:
3418:
3417:Gigantoscelus
3415:
3413:
3410:
3408:
3405:
3403:
3400:
3398:
3395:
3393:
3390:
3388:
3385:
3383:
3380:
3378:
3375:
3373:
3370:
3368:
3367:Abrictosaurus
3365:
3363:
3360:
3359:
3357:
3353:
3343:
3340:
3338:
3335:
3334:
3332:
3328:
3322:
3319:
3317:
3314:
3312:
3309:
3307:
3304:
3302:
3299:
3297:
3294:
3292:
3289:
3287:
3286:Scaloposaurus
3284:
3282:
3279:
3277:
3274:
3272:
3271:Proterosuchus
3269:
3267:
3264:
3262:
3261:Progalesaurus
3259:
3257:
3254:
3252:
3249:
3247:
3244:
3242:
3239:
3237:
3234:
3232:
3229:
3227:
3224:
3222:
3219:
3217:
3214:
3212:
3209:
3207:
3204:
3202:
3199:
3197:
3194:
3192:
3189:
3187:
3186:Lepidodendron
3184:
3182:
3179:
3177:
3174:
3172:
3169:
3167:
3164:
3162:
3161:Euskelosaurus
3159:
3157:
3156:Eucnemesaurus
3154:
3152:
3149:
3147:
3146:Erythrosuchus
3144:
3142:
3141:Ericiolacerta
3139:
3137:
3134:
3132:
3129:
3127:
3124:
3122:
3119:
3117:
3114:
3112:
3111:Blikanasaurus
3109:
3107:
3104:
3103:
3101:
3097:
3087:
3084:
3082:
3079:
3077:
3074:
3072:
3069:
3067:
3064:
3062:
3059:
3058:
3056:
3052:
3046:
3043:
3041:
3038:
3036:
3033:
3031:
3028:
3026:
3023:
3021:
3018:
3016:
3013:
3011:
3008:
3006:
3003:
3001:
2998:
2996:
2995:Pelictosuchus
2993:
2991:
2988:
2986:
2983:
2981:
2978:
2976:
2973:
2971:
2970:Lepidodendron
2968:
2966:
2963:
2961:
2958:
2956:
2953:
2951:
2948:
2946:
2943:
2941:
2940:Ictidochampsa
2938:
2936:
2933:
2931:
2928:
2926:
2923:
2921:
2918:
2916:
2913:
2911:
2908:
2906:
2903:
2901:
2898:
2896:
2893:
2891:
2888:
2886:
2885:Cryptocynodon
2883:
2881:
2878:
2876:
2873:
2871:
2868:
2866:
2863:
2861:
2858:
2856:
2853:
2851:
2848:
2846:
2843:
2841:
2838:
2836:
2835:Anomocephalus
2833:
2831:
2828:
2827:
2825:
2821:
2815:
2812:
2810:
2807:
2805:
2804:Thliptosaurus
2802:
2800:
2799:Theriognathus
2797:
2795:
2794:Stereosternum
2792:
2790:
2787:
2785:
2782:
2780:
2779:Scylacosaurus
2777:
2775:
2774:Procynosuchus
2772:
2770:
2769:Odontocyclops
2767:
2765:
2762:
2760:
2757:
2755:
2752:
2750:
2749:Lepidodendron
2747:
2745:
2742:
2740:
2739:Ictidognathus
2737:
2735:
2732:
2730:
2727:
2725:
2722:
2720:
2717:
2715:
2712:
2710:
2707:
2705:
2704:Dwykaselachus
2702:
2700:
2697:
2695:
2692:
2690:
2687:
2685:
2682:
2680:
2677:
2675:
2674:Crapartinella
2672:
2670:
2667:
2665:
2662:
2660:
2657:
2655:
2652:
2650:
2649:Anomocephalus
2647:
2645:
2642:
2640:
2637:
2636:
2634:
2630:
2624:
2623:Lepidodendron
2621:
2619:
2616:
2615:
2613:
2611:Carboniferous
2609:
2603:
2600:
2598:
2595:
2593:
2590:
2588:
2585:
2583:
2580:
2578:
2575:
2573:
2570:
2568:
2565:
2563:
2562:Antarctilamna
2560:
2558:
2555:
2554:
2552:
2548:
2542:
2539:
2537:
2534:
2532:
2529:
2527:
2524:
2523:
2521:
2517:
2511:
2508:
2506:
2503:
2501:
2498:
2497:
2494:
2490:Paleontology
2487:
2483:
2473:
2470:
2468:
2465:
2463:
2460:
2459:
2456:
2453:
2451:
2448:
2446:
2443:
2440:
2438:
2435:
2434:
2429:
2426:
2425:
2424:
2423:Witwatersrand
2421:
2419:
2418:Merensky Reef
2416:
2414:
2411:
2410:
2407:
2400:
2396:
2382:
2379:
2378:
2376:
2372:
2366:
2363:
2361:
2358:
2356:
2353:
2351:
2348:
2347:
2345:
2341:
2335:
2332:
2330:
2327:
2325:
2322:
2320:
2317:
2316:
2314:
2312:
2308:
2302:
2299:
2297:
2294:
2292:
2289:
2285:
2282:
2280:
2277:
2275:
2272:
2270:
2267:
2266:
2264:
2262:
2259:
2257:
2254:
2252:
2249:
2247:
2244:
2243:
2241:
2237:
2231:
2228:
2226:
2223:
2221:
2218:
2216:
2213:
2211:
2208:
2206:
2203:
2201:
2198:
2196:
2193:
2191:
2188:
2186:
2183:
2181:
2178:
2176:
2173:
2172:
2169:
2162:
2158:
2144:
2141:
2140:
2138:
2134:
2126:
2123:
2121:
2118:
2116:
2113:
2111:
2108:
2107:
2105:
2099:
2096:
2094:
2091:
2089:
2086:
2085:
2084:
2081:
2077:
2074:
2073:
2072:
2069:
2067:
2064:
2060:
2057:
2055:
2052:
2050:
2047:
2045:
2042:
2040:
2037:
2036:
2035:
2032:
2031:
2030:
2027:
2023:
2020:
2018:
2015:
2013:
2010:
2009:
2007:
2006:
2001:
1998:
1994:
1991:
1989:
1986:
1984:
1981:
1979:
1976:
1975:
1974:
1971:
1970:
1968:
1967:
1965:
1961:
1953:
1950:
1949:
1947:
1943:
1940:
1938:
1935:
1933:
1930:
1928:
1925:
1923:
1920:
1918:
1915:
1914:
1912:
1908:
1905:
1903:
1900:
1899:
1897:
1893:
1890:
1888:
1885:
1884:
1882:
1881:
1879:
1875:
1869:
1866:
1864:
1861:
1859:
1858:Moodies Group
1856:
1854:
1851:
1849:
1846:
1844:
1841:
1840:
1838:
1834:
1828:
1825:
1824:
1821:
1814:
1810:
1806:
1799:
1794:
1792:
1787:
1785:
1780:
1779:
1776:
1764:
1756:
1755:
1752:
1743:
1725:
1724:
1720:
1718:
1717:
1713:
1711:
1710:
1706:
1704:
1703:
1699:
1697:
1696:
1692:
1690:
1689:
1688:Isalorhynchus
1685:
1683:
1682:
1681:Hyperodapedon
1678:
1676:
1675:
1671:
1670:
1668:
1666:
1662:
1656:
1655:
1651:
1649:
1648:
1644:
1642:
1641:
1637:
1635:
1634:
1630:
1628:
1627:
1623:
1622:
1620:
1618:
1614:
1608:
1607:
1606:Rhynchosaurus
1603:
1601:
1600:
1596:
1594:
1593:
1589:
1587:
1586:
1582:
1580:
1579:
1575:
1574:
1571:
1568:
1566:
1562:
1556:
1555:
1551:
1549:
1548:
1544:
1542:
1541:
1537:
1535:
1534:
1530:
1527:
1526:
1522:
1521:
1518:
1515:
1513:
1512:Rhynchosauria
1509:
1505:
1493:
1492:
1491:Rhynchosauria
1486:
1469:
1468:
1467:
1466:
1464:
1462:
1461:Rhynchosauria
1458:
1450:
1446:
1445:
1444:
1443:
1440:
1437:
1435:
1431:
1423:
1419:
1418:
1417:
1416:
1413:
1410:
1408:
1404:
1400:
1392:
1388:
1385:
1383:
1379:
1376:
1374:
1370:
1368:
1364:
1362:
1358:
1357:
1352:
1348:
1344:
1343:Rhynchosauria
1337:
1332:
1330:
1325:
1323:
1318:
1317:
1314:
1300:
1295:
1290:
1286:
1282:
1275:
1268:
1265:
1260:
1253:
1250:
1245:
1241:
1236:
1231:
1227:
1223:
1219:
1215:
1212:(3): 150674.
1211:
1207:
1200:
1193:
1190:
1186:
1181:
1175:
1171:
1166:
1161:
1157:
1153:
1148:
1143:
1139:
1135:
1131:
1124:
1122:
1120:
1116:
1110:
1105:
1101:
1097:
1093:
1089:
1082:
1080:
1078:
1076:
1074:
1072:
1068:
1063:
1056:
1053:
1048:
1044:
1037:
1035:
1031:
1025:
1020:
1015:
1010:
1007:(3): 571–88.
1006:
1002:
998:
991:
988:
981:
970:
965:
958:
957:
949:
948:
940:
939:
931:
930:
922:
921:
913:
912:
904:
903:
895:
894:
886:
885:
882:
881:
880:
879:Hyperodapedon
873:
872:
869:
868:
865:
864:
863:
856:
855:
849:
848:
845:
844:
841:
840:
839:
838:Isalorhynchus
832:
831:
825:
822:
821:
818:
817:
814:
813:
812:
805:
804:
798:
797:
794:
793:
790:
789:
788:
781:
780:
774:
773:
770:
769:
766:
765:
764:
757:
756:
750:
749:
746:
745:
737:
736:
733:
732:
731:
724:
723:
720:
719:
716:
715:
709:
708:
702:
699:
698:
692:
691:
688:
687:
684:
683:
682:
681:Rhynchosaurus
675:
674:
668:
665:
664:
661:
660:
657:
656:
655:
648:
647:
644:
643:
640:
639:
638:
631:
630:
627:
626:
623:
622:
621:
614:
613:
610:
609:
606:
605:
604:
597:
596:
590:
589:Rhynchosauria
587:
586:
582:
580:
576:
572:
565:
563:
561:
557:
556:
549:
547:
543:
538:
529:
527:
525:
519:
515:
511:
507:
505:
504:
499:
494:
491:The skull of
486:
484:
482:
477:
473:
469:
465:
461:
453:
448:
446:
442:
438:
436:
432:
427:
425:
421:
413:
411:
409:
408:
403:
398:
394:
393:
392:Ornithosuchus
388:
387:
386:Erythrosuchus
382:
381:
380:Proterosuchus
376:
372:
368:
363:
361:
356:
352:
348:
344:
336:
334:
332:
328:
324:
320:
318:
315:Aliwal North
312:
308:
305:
301:
297:
293:
289:
285:
282:
278:
275:
271:
270:
257:
252:
251:
247:
244:
243:
241:
236:
232:
228:
223:
222:
215:
212:
211:
208:
207:Rhynchosauria
202:
199:
198:
195:
192:
189:
186:
185:
182:
179:
176:
175:
172:
169:
166:
165:
162:
159:
156:
155:
152:
149:
146:
145:
140:
135:
131:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
4043:Rhynchosaurs
3951:
3924:
3910:
3891:
3877:
3771:Andre Keyser
3746:Alfred Brown
3741:Robert Broom
3533:Nqwebasaurus
3523:Kangnasaurus
3487:Sphenosuchus
3412:Geranosaurus
3392:Dracovenator
3372:Antetonitrus
3316:Ufudocyclops
3301:Tetracynodon
3291:Scymnosaurus
3205:
3196:Lystrosaurus
3181:Kannemeyeria
3136:Elliotherium
3035:Silpholestes
2985:Moschorhinus
2965:Leontosaurus
2935:Icticephalus
2925:Euchambersia
2875:Colobodectes
2865:Choerosaurus
2830:Aelurosaurus
2809:Trochosaurus
2789:Simorhinella
2784:Scymnosaurus
2729:Glossopteris
2714:Eunotosaurus
2592:Serenichthys
2577:Diplacanthus
2557:Africanaspis
2301:Karoo-Ferrar
1721:
1716:Supradapedon
1714:
1707:
1700:
1693:
1686:
1679:
1672:
1652:
1645:
1638:
1631:
1626:Ammorhynchus
1624:
1604:
1597:
1590:
1585:Eifelosaurus
1583:
1576:
1552:
1546:
1545:
1538:
1531:
1523:
1489:
1386:
1377:
1284:
1280:
1267:
1258:
1252:
1209:
1205:
1192:
1137:
1133:
1091:
1061:
1055:
1046:
1042:
1024:11449/167867
1004:
1000:
990:
877:
876:
860:
859:
836:
835:
809:
808:
785:
784:
761:
760:
728:
727:
712:
679:
678:
652:
651:
635:
634:
619:
618:
617:
601:
600:
578:
570:
569:
559:
553:
550:
541:
536:
533:
523:
520:
516:
512:
508:
501:
497:
492:
490:
471:
463:
459:
457:
443:
439:
434:
428:
423:
417:
405:
401:
397:Robert Broom
390:
384:
378:
374:
366:
359:
351:Alfred Brown
342:
340:
326:
322:
316:
300:South Africa
268:
267:
266:
249:
248:
220:
219:
187:
126:
24:
18:
3601:Homo naledi
3589:Pleistocene
3538:Paranthodon
3513:Algoasaurus
3492:Tritheledon
3477:Pulanesaura
3407:Fabrosaurus
3382:Coelophysis
3330:Assemblages
3311:Trirachodon
3306:Thrinaxodon
3256:Procolophon
3241:Pentasaurus
3226:Noteosuchus
3216:Micropholis
3191:Lydekkerina
3131:Dinanomodon
3121:Cynognathus
3054:Assemblages
3020:Procolophon
3005:Pristerodon
3000:Polycynodon
2955:Ictidostoma
2950:Ictidodraco
2915:Eohyosaurus
2910:Eodicynodon
2905:Dinanomodon
2855:Broomistega
2724:Glanosuchus
2709:Endothiodon
2694:Dinanomodon
2659:Bulbasaurus
2587:Priscomyzon
2311:Earthquakes
2296:Drakensberg
2251:Pilanesberg
2175:Algoa Basin
2143:Algoa Group
2066:Dwyka Group
1969:Cretaceous
1948:Ordovician
1927:Dwyka Group
1887:Dwyka Group
1709:Paradapedon
1647:Mesodapedon
1640:Elorhynchus
1554:Noteosuchus
1533:Eohyosaurus
654:Eohyosaurus
603:Noteosuchus
449:Description
360:Cynognathus
286:from early
284:rhynchosaur
4037:Categories
3952:Mesosuchus
3703:Geologists
3632:Planolites
3596:Boskop Man
3501:Cretaceous
3467:Pegomastax
3452:Lycorhinus
3437:Ledumahadi
3427:Gyposaurus
3377:Arcusaurus
3296:Shansiodon
3281:Regisaurus
3266:Prolacerta
3206:Mesosuchus
3166:Galesaurus
3126:Diademodon
3116:Bolotridon
2920:Emyduranus
2895:Cynosaurus
2870:Clelandina
2759:Mesosaurus
2754:Lycosuchus
2719:Galechirus
2699:Dinogorgon
2679:Cynosaurus
2526:Notiodella
2519:Ordovician
2413:Kimberlite
2374:Meteorites
2265:Diatremes
2071:Ecca Group
1952:Soom Shale
1723:Teyumbaita
1599:Langeronyx
1547:Mesosuchus
1470:see below↓
1422:Sauropsida
1407:Sauropsida
1373:Sauropsida
966:References
862:Teyumbaita
763:Langeronyx
620:Mesosuchus
571:Mesosuchus
560:Mesosuchus
542:Mesosuchus
537:Mesosuchus
524:Mesosuchus
498:Mesosuchus
493:Mesosuchus
460:Mesosuchus
435:Mesosuchus
424:Mesosuchus
402:Mesosuchus
375:Mesosuchus
343:Mesosuchus
327:Mesosuchus
323:Mesosuchus
317:Euparkeria
269:Mesosuchus
221:Mesosuchus
127:Mesosuchus
25:Mesosuchus
3580:Nummulite
3559:Nummulite
3552:Paleogene
3518:Baculites
3236:Orosaurus
3221:Myosaurus
3040:Tamboeria
2990:Paliguana
2975:Lycideops
2945:Ictidodon
2900:Diictodon
2880:Compsodon
2840:Basilodon
2734:Hofmeyria
2689:Diictodon
2654:Basilodon
2639:Abdalodon
2618:Dadoxylon
2602:Umzantsia
2541:Soomaspis
2536:Promissum
2239:Volcanism
2225:Witteberg
2106:Triassic
2008:Jurassic
1898:Devonian
1877:Paleozoic
1391:Crocopoda
1359:Kingdom:
1156:2167-8359
1140:: e1778.
1049:: 296–97.
575:cladogram
566:Phylogeny
546:herbivore
530:Dentition
481:symphysis
454:Diagnosis
431:paratypes
371:etymology
369:. As the
357:, in the
341:Bones of
333:in 1912.
250:M. browni
157:Kingdom:
151:Eukaryota
36:246
3967:Q1186485
3961:Wikidata
3926:Glossary
3912:Category
3422:Gryponyx
3397:Eocursor
3362:Aardonyx
3355:Jurassic
3099:Triassic
2814:Youngina
2764:Moschops
2597:Tutusius
2567:Bainella
2550:Devonian
2269:Big Hole
2136:Cenozoic
1963:Mesozoic
1913:Permian
1763:Category
1674:Beesiiwo
1578:Bentonyx
1525:Colobops
1367:Chordata
1365:Phylum:
1361:Animalia
1299:Archived
1287:: 1–24.
1244:27069658
1187:license.
1174:27162705
787:Bentonyx
581:(2016):
420:holotype
311:skeleton
304:holotype
238:Species
181:Reptilia
171:Chordata
167:Phylum:
161:Animalia
147:Domain:
4007:4126184
3994:1449596
3981:4821494
3861:Swazian
3568:Neogene
3171:Howesia
2632:Permian
1592:Fodonyx
1540:Howesia
1371:Class:
1235:4821269
1214:Bibcode
1165:4860341
1109:1692244
1064:: 1–23.
811:Fodonyx
637:Howesia
503:Howesia
476:diapsid
445:pedes.
292:Anisian
290:(early
274:extinct
213:Genus:
200:Order:
177:Class:
105:↓
3462:Ngwevu
1242:
1232:
1172:
1162:
1154:
1106:
579:et al.
389:, and
227:Watson
4020:38209
3989:IRMNG
3841:Other
3620:Other
1387:Clade
1378:Clade
1302:(PDF)
1277:(PDF)
1202:(PDF)
1134:PeerJ
573:in a
487:Skull
281:basal
277:genus
188:Clade
3976:GBIF
1447:see
1420:see
1240:PMID
1170:PMID
1152:ISSN
422:for
319:site
296:Cape
256:type
231:1912
44:PreꞒ
1289:doi
1230:PMC
1222:doi
1160:PMC
1142:doi
1104:PMC
1096:doi
1019:hdl
1009:doi
1005:174
433:of
307:SAM
279:of
4039::
4017::
4004::
3991::
3978::
3963::
1389::
1380::
1297:.
1283:.
1279:.
1238:.
1228:.
1220:.
1208:.
1204:.
1168:.
1158:.
1150:.
1136:.
1132:.
1118:^
1102:.
1090:.
1070:^
1045:.
1033:^
1017:.
1003:.
999:.
548:.
410:.
383:,
321:.
298:,
229:,
190::
94:Pg
38:Ma
33:,
1797:e
1790:t
1783:v
1528:?
1335:e
1328:t
1321:v
1291::
1285:3
1261:.
1246:.
1224::
1216::
1210:3
1176:.
1144::
1138:4
1112:.
1098::
1047:2
1027:.
1021::
1011::
258:)
246:†
217:†
204:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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