1855:
1844:
368:
1850:
136:
1861:
1884:
527:
642:
118:
618:
to the teeth of velociraptorine dromaeosaurids. The authors recommended caution when referring to isolated teeth from the Late
Jurassic or Early Cretaceous to the Dromaeosauridae (explicitly citing Milner's 2002 study and Sweetman's 2004 study as examples of studies that identified isolated theropod
608:
dromaeosaurid. If this placement is correct, it would have been one of the oldest dromaeosaurids known, the first to be described, and the first known from
Britain. However, Rauhut, Milner and Moore-Fay (2010) pointed out the great similarity of the teeth of basal
410:. Later several other teeth and specimen BMNH 48207, another dentary fragment from a somewhat smaller individual, were referred to the species. Owen in 1878 also assumed some fossilised scutes, of a type for which he coined the name "granicones", belonged to
406:, dating from the middle Berriasian. It consists of an about three inch long left dentary fragment with nine teeth. The holotype was once thought to be lost but was rediscovered during the 1970s in the
779:
Vertebrate paleobiodiversity of the Early
Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J/K boundary
514:
for "destroyer", a reference to "the adaptations of the teeth for piercing, cutting, and lacerating the prey" of a form he estimated to be equal in size to the present
2003:
704:"Morphology, histology and identification of the 'granicones' from the Purbeck Limestone Formation (Lower Cretaceous: Berriasian) of Dorset, southern England"
1037:
1949:
952:"Cranial osteology and phylogenetic position of the theropod dinosaur Proceratosaurus bradleyi (Woodward, 1910) from the Middle Jurassic of England"
2013:
839:
918:"The first record of velociraptorine dinosaurs (Saurischia, Theropoda) from the Wealden (Early Cretaceous, Barremian) of southern England"
2018:
1877:
655:
675:
Owen, R. (1878). "On the
Fossils called "Granicones"; being a Contribution to the Histology of the Exo‐skeleton in "Reptilia"".
1854:
1843:
1030:
1860:
1849:
135:
1134:
1023:
399:
2008:
507:
457:
348:
1904:
1975:
1887:
563:
250:
738:"Microvertebrate biodiversity from Cherves-De-Cognac (Lower Cretaceous, Berriasian: Charente, France)"
1936:
1188:
407:
597:
1605:
567:
575:
1342:
1161:
812:
403:
376:
340:
275:
130:
438:
found at the
Berriasian aged Cherves-de-Cognac locality, specimen CHEm03.537, was referred to a
367:
1980:
764:
1962:
1802:
1693:
1629:
1525:
1318:
835:
630:
can be diagnosed for having a tooth denticle size difference index ranging from 1-14 to 1-55.
620:
1967:
1726:
1672:
1532:
1510:
963:
932:
804:
756:
718:
684:
647:
610:
605:
559:
448:
336:
324:
30:
1786:
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1234:
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1119:
1046:
614:
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320:
778:
1444:
688:
586:
571:
536:
515:
503:
917:
808:
781:. Geodiversitas, Museum National d’Histoire Naturelle Paris, In press. ffhal-03264773f
703:
1997:
1765:
1679:
1571:
1556:
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1488:
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1471:
1406:
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1328:
984:
968:
951:
894:
816:
757:
702:
Barrett, P. M.; Clarke, J. B.; Brinkman, D. B.; Chapman, S. D.; Ensom, P. C. (2002).
589:
310:
68:
737:
1824:
1735:
1656:
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1614:
1593:
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1517:
1384:
1377:
1370:
1360:
1307:
1292:
777:
Ronan Allain, Romain Vullo, Lee Rozada, Jérémy
Anquetin, Renaud Bourgeais, et al..
580:
480:
468:
425:
395:
285:
226:
936:
875:
856:
498:) meaning "one who admonishes" or "a monitor," in reference to the similarity of
1779:
1757:
1708:
1664:
1621:
1586:
1564:
1503:
1414:
1349:
1173:
555:
488:
43:
600:
examined five good specimens of fossil teeth and confirmed that they belong to
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637:
415:
332:
193:
88:
53:
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1335:
1146:
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1009:
551:
526:
206:
147:
93:
37:
722:
1921:
1715:
1686:
1460:
1270:
1070:
383:
316:
313:
180:
167:
83:
78:
63:
58:
48:
876:"Monograph on the fossil Reptilia of the Wealden and Purbeck formations"
857:"Monograph on the fossil Reptilia of the Wealden and Purbeck formations"
755:
Bonde, N. (2012). "Danish
Dinosaurs: A Review". In Godefroit, P. (ed.).
390:, was collected by Charles Willcox, an amateur paleontologist living at
1954:
1817:
1103:
391:
344:
98:
73:
619:
teeth as belonging to dromaeosaurids), as these teeth might belong to
1392:
1284:
1064:
985:"Theropod dinosaurs of the Purbeck Limestone Group, southern England"
895:"Theropod dinosaurs of the Purbeck Limestone Group, southern England"
547:
435:
419:
352:
328:
157:
1898:
792:
117:
1015:
793:"On some Fossil Reptilian and Mammalian Remains from the Purbecks"
525:
511:
366:
306:
1941:
1902:
1057:
1019:
125:
Portions of the holotype mandible and close up of a tooth
950:
Rauhut, O. W. M.; Milner, A. C.; Moore-Fay, S. (2010).
736:
Pouech, J.; Mazin, J. M.; Billon-Bruyat, J. P. (2006).
585:. In 2002 however, a re-examination of the fossils by
331:
teeth and jaw fragments found in rocks of the middle
1012:
from the Milner paper, on the
Dinosaur Mailing List.
592:
showed that they most likely belonged to a subadult
1911:
1725:
1639:
1604:
1542:
1470:
1359:
1317:
1254:
1199:
1172:
1145:
1118:
554:; later he changed his mind concluding it was a
414:but these were in 2002 shown to be limb or tail
534:(centre) depicted as a dromaeosaur capturing a
1031:
339:) age in the Cherty Freshwater Member of the
8:
797:Quarterly Journal of the Geological Society
452:. Additional teeth have been attributed to
1899:
1548:
1262:
1251:
1178:
1151:
1124:
1115:
1054:
1038:
1024:
1016:
677:Journal of the Royal Microscopical Society
562:did understand it was a dinosaur. In 1934
116:
20:
967:
956:Zoological Journal of the Linnean Society
832:Dinosaurs: The Encyclopedia, Supplement 2
834:. McFarland & Company. p. 686.
667:
604:, and concluded that this species is a
546:was originally classified by Owen as a
763:. Indiana University Press. pp.
442:sp. Some large specimens referred to
7:
2004:Early Cretaceous dinosaurs of Europe
689:10.1111/j.1365-2818.1878.tb01721.x
656:Timeline of dromaeosaurid research
359:would have been a small predator.
14:
809:10.1144/GSL.JGS.1854.010.01-02.48
1883:
1882:
1859:
1853:
1848:
1842:
969:10.1111/j.1096-3642.2009.00591.x
640:
134:
989:Special Papers in Palaeontology
899:Special Papers in Palaeontology
742:Mesozoic Terrestrial Ecosystems
494:, a contraction of νουθέτητης (
394:, from the Feather Quarry near
880:The Palaeontographical Society
861:The Palaeontographical Society
1:
2014:Fossil taxa described in 1854
937:10.1016/j.cretres.2004.01.004
355:. If it was a dromaeosaurid,
502:teeth to those of a modern
479:was named and described by
2035:
2019:Taxa named by Richard Owen
483:in 1854. The generic name
467:contains one species (the
398:in a marine deposition of
16:Extinct genus of dinosaurs
1872:
1840:
1551:
1265:
1250:
1231:
1181:
1154:
1127:
1114:
1053:
623:tyrannosauroids instead.
578:even renamed the species
281:
274:
256:
249:
131:Scientific classification
129:
124:
115:
23:
916:Sweetman, S. C. (2004).
400:Cherty Freshwater Member
458:Angeac-Charente bonebed
434:. In 2006 a tooth from
349:Angeac-Charente bonebed
305:is the name given to a
983:Milner, A. C. (2002).
893:Milner, A. C. (2002).
723:10.1006/cres.2002.1002
540:
446:may instead belong to
379:
1976:Paleobiology Database
1010:Extensive description
626:According to Milner,
564:William Elgin Swinton
529:
370:
830:Glut, D. F. (2002).
759:Bernissart Dinosaurs
487:is derived from the
408:Dorset County Museum
363:Discovery and naming
1606:Saurornitholestinae
925:Cretaceous Research
711:Cretaceous Research
602:Nuthetes destructor
473:Nuthetes destructor
460:in western France.
263:Nuthetes destructor
541:
404:Lulworth Formation
380:
377:Lulworth Formation
341:Lulworth Formation
327:, known only from
242:N. destructor
1991:
1990:
1963:Open Tree of Life
1905:Taxon identifiers
1896:
1895:
1868:
1867:
1838:
1837:
1834:
1833:
1803:Saurornitholestes
1694:Saurornitholestes
1630:Saurornitholestes
1319:Halszkaraptorinae
1246:
1245:
1242:
1241:
1227:
1226:
1223:
1222:
1219:
1218:
874:Owen, R. (1878).
863:. supp. 5: 31–39.
855:Owen, R. (1873).
841:978-0-7864-1166-5
791:Owen, R. (1854).
566:thought it was a
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1727:Velociraptorinae
1673:Dromaeosauroides
1549:
1533:Zhongjianosaurus
1511:Sinornithosaurus
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884:
883:
882:. supp. 8: 1–19.
871:
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820:
803:(1–2): 420–433.
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648:Dinosaurs portal
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560:Richard Lydekker
456:from the nearby
449:Dromaeosauroides
337:Early Cretaceous
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31:Early Cretaceous
29:Temporal range:
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1787:Luanchuanraptor
1721:
1641:Dromaeosaurinae
1635:
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1544:Eudromaeosauria
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1257:Dromaeosauridae
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1235:Dromaeosauridae
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1201:Dromaeosauridae
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1135:Avemetatarsalia
1120:Avemetatarsalia
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621:proceratosaurid
615:Proceratosaurus
606:velociraptorine
558:. Only in 1888
530:Restoration of
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1004:External links
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962:(1): 155–195.
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683:(5): 233–236.
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611:tyrannosauroid
598:Steve Sweetman
587:paleontologist
572:Megalosauridae
570:member of the
537:Durlstotherium
523:
522:Classification
520:
516:Bengal monitor
504:monitor lizard
432:"H." bakewelli
364:
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325:tyrannosauroid
311:coelurosaurian
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1572:Dineobellator
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1496:Hesperonychus
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1489:Graciliraptor
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1482:Changyuraptor
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1407:Ornithodesmus
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1400:Neuquenraptor
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1329:Halszkaraptor
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986:
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976:
970:
965:
961:
957:
953:
946:
943:
938:
934:
930:
926:
919:
912:
909:
904:
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896:
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867:
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848:
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837:
833:
826:
823:
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787:
784:
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774:
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766:
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724:
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705:
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682:
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661:
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649:
638:
633:
631:
629:
624:
622:
617:
616:
612:
607:
603:
599:
595:
594:dromaeosaurid
591:
590:Angela Milner
588:
584:
582:
577:
573:
569:
565:
561:
557:
553:
549:
545:
539:
538:
533:
528:
521:
519:
517:
513:
509:
508:specific name
505:
501:
497:
493:
490:
486:
482:
478:
477:N. destructor
474:
470:
466:
461:
459:
455:
451:
450:
445:
441:
437:
433:
429:
427:
421:
417:
413:
409:
405:
401:
397:
393:
389:
385:
378:
374:
373:N. destructor
371:Tooth tip of
369:
362:
360:
358:
354:
350:
347:and also the
346:
342:
338:
334:
330:
326:
322:
321:dromaeosaurid
318:
315:
312:
308:
304:
303:
291:(Steel, 1970)
289:
287:
283:
282:
280:
277:
273:
266:
264:
255:
252:
251:Binomial name
248:
244:
243:
237:
234:
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228:
223:
222:
215:
212:
211:
208:
205:
202:
199:
198:
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176:
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162:
159:
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153:
152:
149:
146:
143:
142:
137:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
1912:
1847:
1825:Velociraptor
1823:
1816:
1809:
1801:
1794:
1793:
1785:
1778:
1771:
1764:
1756:
1748:
1741:
1736:Acheroraptor
1734:
1714:
1707:
1700:
1692:
1685:
1678:
1671:
1663:
1657:Dakotaraptor
1655:
1650:Achillobator
1648:
1628:
1620:
1615:Atrociraptor
1613:
1594:Zhenyuanlong
1592:
1585:
1579:Tianyuraptor
1577:
1570:
1563:
1555:
1531:
1524:
1518:Tianyuraptor
1516:
1509:
1502:
1494:
1487:
1480:
1459:
1451:
1443:
1436:
1428:
1420:
1413:
1405:
1398:
1391:
1385:Dakotaraptor
1383:
1378:Buitreraptor
1376:
1371:Austroraptor
1369:
1361:Unenlagiinae
1348:
1341:
1334:
1327:
1308:Zhenyuanlong
1306:
1298:
1293:Tianyuraptor
1291:
1283:
1276:
1269:
1233:
1099:
1090:
1081:
992:
988:
978:
959:
955:
945:
928:
924:
911:
902:
898:
888:
879:
869:
860:
850:
831:
825:
800:
796:
786:
773:
758:
750:
741:
731:
714:
710:
697:
680:
676:
670:
627:
625:
613:
601:
581:Megalosaurus
579:
576:Rodney Steel
543:
542:
535:
531:
499:
495:
491:
484:
481:Richard Owen
476:
472:
469:type species
464:
462:
453:
447:
443:
439:
431:
426:Helochelydra
423:
411:
396:Durlston Bay
387:
381:
372:
356:
301:
300:
299:
286:Megalosaurus
284:
262:
257:
241:
240:
220:
219:
200:
187:
174:
24:
18:
1780:Linheraptor
1758:Deinonychus
1709:Yurgovuchia
1665:Deinonychus
1622:Bambiraptor
1587:Vectiraptor
1565:Bambiraptor
1504:Microraptor
1415:Pamparaptor
1350:Natovenator
1189:Maniraptora
1174:Maniraptora
556:crocodilian
496:nouthetetes
489:Koine Greek
422:, possibly
388:DORCM G 913
319:, either a
1998:Categories
1876:See also:
1750:Boreonykus
1702:Utahraptor
1453:Variraptor
1422:Pyroraptor
1300:Variraptor
1278:Pyroraptor
1210:see below↓
1086:Dinosauria
1077:Sauropsida
995:: 191–201.
905:: 191–201.
662:References
583:destructor
574:. In 1970
463:The genus
416:osteoderms
333:Berriasian
288:destructor
269:Owen, 1854
194:Saurischia
181:Dinosauria
1743:Adasaurus
1438:Unenlagia
1430:Rahonavis
1336:Hulsanpes
1162:Theropoda
1147:Theropoda
1095:Theropoda
1063:Kingdom:
817:130752835
744:: 96–100.
492:nouthetes
428:" anglica
375:from the
235:Species:
207:Theropoda
154:Kingdom:
148:Eukaryota
36:143
1942:10578592
1922:Wikidata
1913:Nuthetes
1888:Category
1878:Timeline
1795:Nuthetes
1716:Zapsalis
1687:Itemirus
1461:Ypupiara
1343:Mahakala
1271:Daurlong
1071:Chordata
1069:Phylum:
1065:Animalia
634:See also
628:Nuthetes
568:juvenile
544:Nuthetes
532:Nuthetes
500:Nuthetes
485:Nuthetes
465:Nuthetes
454:Nuthetes
444:Nuthetes
440:Nuthetes
412:Nuthetes
384:holotype
357:Nuthetes
317:dinosaur
314:theropod
302:Nuthetes
276:Synonyms
221:Nuthetes
168:Chordata
164:Phylum:
158:Animalia
144:Domain:
25:Nuthetes
1968:4946089
1955:4822941
1928:Q957171
1818:Tsaagan
1104:Paraves
1075:Class:
552:varanid
402:of the
392:Swanage
345:England
260:†
239:†
213:Genus:
105:↓
1557:Balaur
1526:Wulong
1393:Diuqin
1285:Shanag
838:
815:
550:and a
548:lizard
506:. The
436:France
420:turtle
353:France
329:fossil
229:, 1854
1981:66532
1100:Clade
1091:Clade
1082:Clade
921:(PDF)
813:S2CID
767:–449.
707:(PDF)
512:Latin
418:of a
323:or a
307:genus
201:Clade
188:Clade
175:Clade
1950:GBIF
1811:Shri
1773:Kuru
1187:see
1160:see
1133:see
836:ISBN
382:The
227:Owen
44:PreꞒ
1937:EoL
964:doi
960:158
933:doi
805:doi
765:435
719:doi
685:doi
510:is
471:),
430:or
351:in
343:in
309:of
2000::
1978::
1965::
1952::
1939::
1924::
1102::
1093::
1084::
993:68
991:.
987:.
958:.
954:.
929:25
927:.
923:.
903:68
901:.
897:.
878:.
859:.
811:.
801:10
799:.
795:.
740:.
715:23
713:.
709:.
679:.
596:.
518:.
475:.
386:,
203::
190::
177::
94:Pg
38:Ma
33:,
1806:?
1798:?
1790:?
1761:?
1753:?
1697:?
1668:?
1660:?
1625:?
1597:?
1582:?
1560:?
1521:?
1499:?
1474:?
1456:?
1448:?
1433:?
1425:?
1410:?
1388:?
1363:?
1321:?
1303:?
1288:?
1039:e
1032:t
1025:v
972:.
966::
939:.
935::
844:.
819:.
807::
725:.
721::
691:.
687::
681:1
424:"
335:(
217:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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