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Nymphalidae

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have the submedial vein (vein 1) unbranched and in one subfamily forked near the base; the medial vein has three branches, veins 2, 3, and 4; veins 5 and 6 arise from the points of junction of the discocellulars; the subcostal vein and its continuation beyond the apex of cell, vein 7, has never more
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The trait for which these butterflies are most known is the use of only four legs; the reason their forelegs have become vestigial is not yet completely clear. Some suggest the forelegs are used to amplify the sense of smell, because some species possess a brush-like set of soft hair called
1482:"Papers presented at the Mammalian Phylogeny symposium during the 2002 Annual Meeting of the Society for Molecular Biology and Evolution, Sorrento, Italy, June 13–16, 2002 (Chapter: Towards a better understanding of the higher systematics of Nymphalidae (Lepidoptera: Papilionoidea))" 1313:, which has led researchers to believe the forelegs are used to improve signaling and communication between the species, while standing in the other four. This ability proves useful in terms of reproduction and the overall health of the species, and it is the leading theory so far. 354:, because they are known to stand on only four legs while the other two are curled up; in some species, these forelegs have a brush-like set of hairs, which gives this family its other common name. Many species are brightly coloured and include popular species such as the 303: 434:
have internal (1a) and precostal veins. The cell in both wings is closed or open, often closed in the fore, open in the hindwing. The dorsal margin of the hindwing is channelled to receive the abdomen in many of the forms.
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than four branches, veins 8–11; 8 and 9 always arise from vein 7, 10, and 11 sometimes from vein 7 but more often free, i.e., given off by the subcostal vein before apex of the cell.
1461: 3892: 346:, they are usually medium-sized to large butterflies. Most species have a reduced pair of forelegs and many hold their colourful wings flat when resting. They are also called 308: 3766: 3805: 274: 442:
always have two grooves on the underside; the club is variable in shape. Throughout the family, the front pair of legs in the male, and with three exceptions (
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was unequivocally implied in the formation of the name (Code Article 11.7.1.1). The attribution of the Nymphalidae to Rafinesque has now been widely adopted.
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In the adult butterflies, the first pair of legs is small or reduced, giving the family the other names of four-footed or brush-footed butterflies. The
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Rafinesque introduced the name Nymphalia as a subfamily name in diurnal Lepidoptera. Rafinesque did not include
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Niklas Wahlberg, Elisabet Weingartner & Sören Nylin (2003). Gisella Caccone & Giacomo Bernardi (ed.).
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butterflies, the caterpillars often have head spines or projections. Mostly edible species, have some
342:, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily 3882: 3859: 3820: 3688: 3603: 1981: 1973: 1964: 1708: 1540: 1130: 1112: 1041: 3127: 2902: 2894: 1467: 1238: 923: 232: 1121: 49: 3846: 1560: 843: 584:
Most species have long wings, and some have transparent wings. Host plants are in the families
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were formerly well-recognized as distinct families due to insufficient study.
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Wolfe, Joanna M.; Oliver, Jeffrey C.; Monteiro, Antónia (2011-01-01).
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Analyse de la Nature, ou Tableau de l'Univers et des Corps Organisés.
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Caterpillars resemble those of the Danainae and feed on Apocynaceae.
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Vane-Wright & de Jong, 2003: 167; Pelham, 2008; Wahlberg, 2010
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effect that helps the butterflies blend into their surroundings.
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Catalogue of the Butterflies of the United States and Canada
3154:(gossamer-winged butterflies: blues, coppers and relatives) 3375:(daggers, sallows, owlet moths, quakers, cutworms, darts) 415:
are hairy or spiky with projections on the head, and the
3362:(underwing, tiger, tussock, litter, snout, owlet moths) 492:
are of unclear position, reflecting the fact that some
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The Fauna of British India, Including Ceylon and Burma
525:, earlier treated as the distinct family Libytheidae) 3563: 3483: 3439: 3405: 3350: 3263: 3242: 3221: 3192: 3171: 3092: 3071: 3050: 3021: 3003: 2985: 2964: 2935: 2914: 2893: 2872: 2851: 2830: 2793: 2764: 2676: 2658: 2543: 2481: 2442: 2392: 2374: 2347: 2322: 2301: 2272: 2251: 2207: 2180: 2151: 2122: 2072: 2063: 2033: 2024: 2006: 1997: 1972: 1963: 1941: 1932: 1888: 1845: 1802: 1663:Canadian Biodiversity Information Facility (2003): 858:Colourful tropical butterflies, they are noted for 646:, they are restricted to host plants in the family 2813:(castniid moths: giant butterfly-moths, sun moths) 853:(earlier treated as distinct family Heliconiidae) 476:, and are in the male often very inconspicuous. 918:of the heliconiine clade, also includes tribes 771:(owls, neotropical with 70–80 species, mostly 209: 1716: 1375:Jean Barravecceia: Palermo. 224 pages, p 127. 8: 862:. All species use host plants in the family 3893:Taxa named by Constantine Samuel Rafinesque 811:Host plants are in the families Arecaceae, 464:of the forelegs is considerable, e.g., the 404:For an explanation of technical terms, see 3551: 3162:(brush-footed, or four-footed butterflies) 2371: 2367: 2204: 2177: 2069: 2030: 2003: 1969: 1938: 1929: 1925: 1911: 1885: 1871: 1842: 1828: 1799: 1785: 1723: 1709: 1701: 575:species, sometimes considered a subfamily 40: 31: 1594:"Butterfly watch: four legs vs. six legs" 1427: 835:of the nymphaline clade, excludes former 3341:(hawk moths, sphinx moths and hornworms) 1623:Butterflies through Binoculars, The West 3138:(whites, yellows, orangetips, sulphurs) 1683:, Brisbane butterflies web site (2005). 1681:Danaids and Browns - Family Nymphalidae 1364: 846:, and tribes Pseudergolini and Coeini) 784:Host plants in the families Arecaceae, 488:of the Nymphalidae is complex. Several 3254:(eggars, snout moths, or lappet moths) 1637:Life Histories of Cascadia Butterflies 1628:Guppy, Crispin S. and Shepard, Jon H. 633:(about six species, restricted to the 1635:James, David G. and Nunnallee, David 1535: 1533: 1531: 1529: 1527: 1490:Molecular Phylogenetics and Evolution 1451: 1449: 1447: 1290:(includes several subspecies such as 7: 3821:e1ca03a2-3aec-414c-8039-e544ff1bb0d8 3733:24c56ea2-9787-4cf5-995b-fcdcfeb0b576 747:Include the spectacular neotropical 1696:Nymphalidae from all over the world 775:, sometimes considered a subfamily 738:, sometimes considered a subfamily 608:, sometimes considered a subfamily 2776:(carpenter millers, or goat moths) 2693:(burnet, forester, or smoky moths) 1353:List of fritillaries (butterflies) 670:. Host plants are in the families 406:External morphology of Lepidoptera 25: 2097:(Amazonian primitive ghost moths) 292:Small pearl-bordered fritillary ( 1337: 1323: 1024:Example species from this family 1003:, Poaceae, Rubiaceae, Rutaceae, 881:Host plants are in the families 53: 2113:(African primitive ghost moths) 1630:Butterflies of British Columbia 1257:Small pearl-bordered fritillary 2417:(Old World spiny-winged moths) 1569:10.1016/B0-12-226865-2/00039-0 938:Host plants are in the family 819:, Heliconiaceae, Poaceae, and 753:, its food plants include the 1: 3315:(Kentish glory and relatives) 2134:(New Zealand primitive moths) 1503:10.1016/S1055-7903(03)00052-6 392:among the listed genera, but 3461:(oriental swallowtail moths) 2739:(planthopper parasite moths) 1551:Encyclopedia of Biodiversity 1275:Texan crescentspot butterfly 765:, Poaceae, and Sapindaceae. 549:Host plant families include 3535:Taxonomy of the Lepidoptera 3122:(American moth-butterflies) 3083:(Old World butterfly-moths) 3062:(picture-winged leaf moths) 1812:(mandibulate archaic moths) 1651:The Butterflies of Cascadia 1086:Cracker butterflies, genus 960:(formerly in Limenitidinae) 3914: 3033:(tropical fruitworm moths) 2570:(case-bearers, case moths) 2334:(trumpet leaf miner moths) 2089:(swift moths, ghost moths) 1543:(2001). "Nymphalidae". In 403: 250:Crimson-spotted forester ( 3530: 3511: 3283:(American silkworm moths) 3114:(swallowtail butterflies) 2784:(dudgeon carpenter moths) 2493:(false diamondback moths) 2404:(burrowing webworm moths) 2370: 1928: 1924: 1910: 1884: 1870: 1841: 1827: 1798: 1784: 1740: 1400:Journal of Insect Science 1371:Rafinesque, C.S. (1815). 1296:Speyeria zerene hippolyta 1216:Polygonia interrogationis 480:Systematics and phylogeny 223: 216: 144: 139: 50:Scientific classification 48: 39: 34: 3275:(Australian lappet moth) 2284:(pigmy, or midget moths) 1687:Tree of Life Web Project 1468:Taylor and Francis, Ltd. 1279:Anthanassa texana texana 1051:California tortoiseshell 348:brush-footed butterflies 27:Largest butterfly family 2105:(miniature ghost moths) 604:(about 6–10 species in 503:within the family are: 352:four-footed butterflies 3484:Superfamily unassigned 2905:(bristle-legged moths) 2454:(ribbed cocoon makers) 2263:(Andean endemic moths) 2219:(fairy longhorn moths) 1672:"Family Nymphalidae". 1464:: Butterflies Volume I 1078:Common snout butterfly 327: 299: 285: 281:Tirumala septentrionis 271: 257: 243: 3842:Paleobiology Database 3391:(prominents, kittens) 2926:(fringe-tufted moths) 1982:Acanthopteroctetidae 1974:Acanthopteroctetoidea 1649:Pyle, Robert Michael 1234:Coenonympha pamphilus 1055:Nymphalis californica 314: 291: 277: 264:Monarch butterflies ( 263: 249: 235: 3728:Fauna Europaea (new) 2903:Schreckensteiniidae 2895:Schreckensteinioidea 2884:(false burnet moths) 2755:(slug, or cup moths) 2313:(Gondwanaland moths) 2292:(white eyecap moths) 2050:(archaic bell moths) 1412:10.1673/031.011.6601 1270:Cithaerias andromeda 1113:Grayling (butterfly) 1082:Libytheana carinenta 226:About 5,700 species 3233:(sack bearer moths) 2955:(false plume moths) 2947:(many-plumed moths) 2805:(little bear moths) 2747:(slug caterpillars) 2602:(long-horned moths) 2586:(grass-miner moths) 1984:(archaic sun moths) 1621:Glassberg, Jeffrey 1598:Scientific American 1239:Small tortoiseshell 1104:Edith's checkerspot 3888:Butterfly families 1665:Family Nymphalidae 1144:Euphydryas aurinia 1135:Limenitis lorquini 1126:Asterocampa celtis 1015:, Urticaceae, and 885:, Passifloraceae, 419:have shiny spots. 328: 300: 286: 272: 258: 244: 3870: 3869: 3829:Open Tree of Life 3557:Taxon identifiers 3548: 3547: 3526: 3525: 3522: 3521: 3507: 3506: 3503: 3502: 3499: 3498: 3475: 3462: 3426: 3418: 3392: 3384: 3376: 3363: 3342: 3334: 3316: 3308: 3300: 3292: 3284: 3276: 3255: 3234: 3213: 3205: 3184: 3163: 3155: 3147: 3139: 3131: 3123: 3115: 3105: 3084: 3063: 3042: 3041:(fruitworm moths) 3034: 2977: 2956: 2948: 2927: 2906: 2885: 2864: 2843: 2842:(metalmark moths) 2822: 2821:(clearwing moths) 2814: 2806: 2785: 2777: 2756: 2748: 2740: 2717: 2694: 2650: 2642: 2629: 2628:(concealer moths) 2621: 2603: 2595: 2587: 2579: 2571: 2530: 2507: 2504:Glyphipterigidae 2494: 2473:Roeslerstammiidae 2463: 2455: 2434: 2426: 2418: 2405: 2343: 2342: 2335: 2314: 2293: 2285: 2264: 2243: 2220: 2147: 2146: 2143: 2142: 2135: 2114: 2106: 2098: 2090: 2059: 2058: 2051: 2020: 2019: 1993: 1992: 1985: 1959: 1958: 1906: 1905: 1902: 1901: 1890:Heterobathmioidea 1866: 1865: 1862: 1861: 1856: 1823: 1822: 1819: 1818: 1813: 1642:Pelham, Jonathan 1578:978-0-12-226865-6 1541:Philip J. DeVries 1345:Arthropods portal 1292:Oregon silverspot 1284:Zerene fritillary 1243:Nymphalis urticae 1207:Danaus chrysippus 1167:Monarch butterfly 1162:Nymphalis antiopa 1131:Lorquin's admiral 1122:Hackberry emperor 1117:Hipparchia semele 1108:Euphydryas editha 1064:Polygonia c-album 933:(mostly tropical) 860:Müllerian mimicry 829:heliconiine clade 523:snout butterflies 360:monarch butterfly 312: 278:Dark blue tiger ( 230: 229: 135: 16:(Redirected from 3905: 3863: 3862: 3850: 3849: 3837: 3836: 3824: 3823: 3814: 3813: 3801: 3800: 3798:NBNSYS0000160544 3788: 3787: 3775: 3774: 3762: 3761: 3749: 3748: 3736: 3735: 3723: 3722: 3710: 3709: 3697: 3696: 3684: 3683: 3671: 3670: 3658: 3657: 3645: 3644: 3635: 3634: 3625: 3624: 3612: 3611: 3599: 3598: 3597: 3584: 3583: 3582: 3552: 3474:(geometer moths) 3473: 3460: 3454:Pseudobistonidae 3424: 3416: 3390: 3382: 3374: 3361: 3340: 3332: 3314: 3306: 3298: 3290: 3282: 3274: 3253: 3232: 3211: 3203: 3182: 3161: 3153: 3145: 3137: 3129: 3121: 3113: 3104: 3097: 3082: 3061: 3040: 3032: 2975: 2954: 2946: 2925: 2904: 2883: 2862: 2841: 2820: 2812: 2804: 2783: 2775: 2754: 2746: 2738: 2715: 2692: 2648: 2640: 2627: 2619: 2601: 2593: 2585: 2577: 2576:Cosmopterigidae 2569: 2528: 2505: 2492: 2461: 2453: 2452:Bucculatricidae 2432: 2424: 2416: 2403: 2372: 2368: 2333: 2312: 2291: 2283: 2262: 2241: 2218: 2205: 2178: 2133: 2112: 2104: 2096: 2088: 2070: 2049: 2031: 2004: 1983: 1970: 1939: 1930: 1926: 1912: 1896:Heterobathmiidae 1886: 1872: 1854: 1843: 1829: 1811: 1810:Micropterigidae 1804:Micropterigoidea 1800: 1786: 1725: 1718: 1711: 1702: 1609: 1608: 1606: 1604: 1589: 1583: 1582: 1554: 1537: 1522: 1521: 1519: 1513:. Archived from 1486: 1477: 1471: 1470: 1453: 1442: 1441: 1431: 1391: 1385: 1382: 1376: 1369: 1347: 1342: 1341: 1333: 1328: 1327: 1252:Pyronia tithonus 1225:Vanessa atalanta 1171:Danaus plexippus 1140:Marsh fritillary 1013:Scrophulariaceae 912:nymphaline clade 508:libytheine clade 334:are the largest 313: 267:Danaus plexippus 253:Euphaedra themis 130: 58: 57: 44: 32: 21: 3913: 3912: 3908: 3907: 3906: 3904: 3903: 3902: 3873: 3872: 3871: 3866: 3858: 3853: 3845: 3840: 3832: 3827: 3819: 3817: 3809: 3804: 3796: 3791: 3783: 3778: 3770: 3765: 3757: 3752: 3744: 3739: 3731: 3726: 3718: 3713: 3705: 3700: 3692: 3687: 3679: 3674: 3666: 3661: 3653: 3648: 3640: 3638: 3630: 3628: 3620: 3615: 3607: 3602: 3593: 3592: 3587: 3578: 3577: 3572: 3559: 3549: 3544: 3540:Lists by region 3518: 3517:is not a clade. 3513:Note: division 3495: 3479: 3435: 3401: 3346: 3307:(Dryandra moth) 3299:(Brahmin moths) 3259: 3238: 3217: 3188: 3167: 3098: 3096: 3088: 3067: 3046: 3031:Copromorphidae 3023:Copromorphoidea 3017: 2999: 2981: 2960: 2931: 2910: 2889: 2868: 2863:(tortrix moths) 2847: 2826: 2789: 2760: 2716:(flannel moths) 2699:Himantopteridae 2672: 2654: 2600:Lecithoceridae 2594:(twirler moths) 2539: 2477: 2462:(Douglas moths) 2444:Gracillarioidea 2438: 2425:(bagworm moths) 2410:Arrhenophanidae 2388: 2376:Simaethistoidea 2362: 2360: 2358: 2356: 2354: 2352: 2350: 2339: 2318: 2297: 2268: 2247: 2199: 2197: 2195: 2193: 2191: 2189: 2187: 2185: 2183: 2172: 2170: 2168: 2166: 2164: 2162: 2160: 2158: 2156: 2154: 2139: 2132:Mnesarchaeidae 2124:Mnesarchaeoidea 2118: 2111:Prototheoridae 2055: 2043:Aenigmatineidae 2016: 2008:Lophocoronoidea 1989: 1955: 1920: 1898: 1880: 1878:Heterobathmiina 1858: 1853:Agathiphagidae 1847:Agathiphagoidea 1837: 1815: 1794: 1780: 1736: 1729: 1660: 1618: 1616:Further reading 1613: 1612: 1602: 1600: 1591: 1590: 1586: 1579: 1539: 1538: 1525: 1517: 1484: 1479: 1478: 1474: 1455: 1454: 1445: 1393: 1392: 1388: 1383: 1379: 1370: 1366: 1361: 1343: 1336: 1329: 1322: 1319: 1306: 1301: 1288:Speyeria zerene 1266:Andromeda satyr 1180:Morpho menelaus 1153:Maniola jurtina 1099:Chlosyne janais 1032:Actinote zikani 1026: 821:Selaginellaceae 668:Batesian mimics 555:Asclepiadoideae 482: 409: 402: 384: 324:yellow ironweed 302: 225: 224:Over 600 genera 129: 52: 28: 23: 22: 15: 12: 11: 5: 3911: 3909: 3901: 3900: 3895: 3890: 3885: 3875: 3874: 3868: 3867: 3865: 3864: 3851: 3838: 3825: 3815: 3802: 3789: 3776: 3763: 3750: 3737: 3724: 3715:Fauna Europaea 3711: 3698: 3685: 3672: 3659: 3646: 3636: 3626: 3613: 3600: 3585: 3569: 3567: 3561: 3560: 3555: 3546: 3545: 3543: 3542: 3537: 3531: 3528: 3527: 3524: 3523: 3520: 3519: 3512: 3509: 3508: 3505: 3504: 3501: 3500: 3497: 3496: 3494: 3493: 3487: 3485: 3481: 3480: 3478: 3477: 3469: 3464: 3456: 3451: 3445: 3443: 3437: 3436: 3434: 3433: 3428: 3420: 3411: 3409: 3403: 3402: 3400: 3399: 3394: 3386: 3378: 3370: 3365: 3356: 3354: 3348: 3347: 3345: 3344: 3336: 3328: 3323: 3318: 3310: 3302: 3294: 3286: 3278: 3269: 3267: 3261: 3260: 3258: 3257: 3252:Lasiocampidae 3248: 3246: 3244:Lasiocampoidea 3240: 3239: 3237: 3236: 3227: 3225: 3219: 3218: 3216: 3215: 3207: 3198: 3196: 3190: 3189: 3187: 3186: 3177: 3175: 3169: 3168: 3166: 3165: 3157: 3149: 3141: 3133: 3125: 3117: 3108: 3106: 3090: 3089: 3087: 3086: 3077: 3075: 3069: 3068: 3066: 3065: 3056: 3054: 3048: 3047: 3045: 3044: 3036: 3027: 3025: 3019: 3018: 3016: 3015: 3009: 3007: 3001: 3000: 2998: 2997: 2991: 2989: 2987:Whalleyanoidea 2983: 2982: 2980: 2979: 2974:Pterophoridae 2970: 2968: 2966:Pterophoroidea 2962: 2961: 2959: 2958: 2950: 2941: 2939: 2933: 2932: 2930: 2929: 2920: 2918: 2912: 2911: 2909: 2908: 2899: 2897: 2891: 2890: 2888: 2887: 2878: 2876: 2870: 2869: 2867: 2866: 2857: 2855: 2849: 2848: 2846: 2845: 2836: 2834: 2828: 2827: 2825: 2824: 2816: 2808: 2799: 2797: 2791: 2790: 2788: 2787: 2779: 2770: 2768: 2762: 2761: 2759: 2758: 2750: 2742: 2734: 2729: 2724: 2719: 2714:Megalopygidae 2711: 2709:Somabrachyidae 2706: 2701: 2696: 2688: 2682: 2680: 2674: 2673: 2671: 2670: 2664: 2662: 2656: 2655: 2653: 2652: 2649:(timber moths) 2644: 2641:(flower moths) 2636: 2634:Pterolonchidae 2631: 2623: 2620:(mompha moths) 2615: 2610: 2605: 2597: 2589: 2581: 2578:(cosmet moths) 2573: 2568:Coleophoridae 2565: 2560: 2558:Batrachedridae 2555: 2549: 2547: 2541: 2540: 2538: 2537: 2532: 2529:(ermine moths) 2527:Yponomeutidae 2524: 2519: 2514: 2509: 2501: 2496: 2487: 2485: 2483:Yponomeutoidea 2479: 2478: 2476: 2475: 2470: 2468:Gracillariidae 2465: 2457: 2448: 2446: 2440: 2439: 2437: 2436: 2433:(fungus moths) 2428: 2420: 2412: 2407: 2398: 2396: 2390: 2389: 2387: 2386: 2384:Simaethistidae 2380: 2378: 2365: 2345: 2344: 2341: 2340: 2338: 2337: 2328: 2326: 2320: 2319: 2317: 2316: 2311:Palaephatidae 2307: 2305: 2303:Palaephatoidea 2299: 2298: 2296: 2295: 2287: 2278: 2276: 2270: 2269: 2267: 2266: 2257: 2255: 2249: 2248: 2246: 2245: 2237: 2232: 2227: 2222: 2213: 2211: 2202: 2175: 2149: 2148: 2145: 2144: 2141: 2140: 2138: 2137: 2128: 2126: 2120: 2119: 2117: 2116: 2108: 2103:Palaeosetidae 2100: 2092: 2084: 2078: 2076: 2067: 2061: 2060: 2057: 2056: 2054: 2053: 2048:Neopseustidae 2045: 2039: 2037: 2035:Neopseustoidea 2028: 2022: 2021: 2018: 2017: 2014:Lophocoronidae 2012: 2010: 2001: 1995: 1994: 1991: 1990: 1988: 1987: 1978: 1976: 1967: 1961: 1960: 1957: 1956: 1954: 1953: 1947: 1945: 1943:Eriocranioidea 1936: 1922: 1921: 1915: 1908: 1907: 1904: 1903: 1900: 1899: 1894: 1892: 1882: 1881: 1875: 1868: 1867: 1864: 1863: 1860: 1859: 1851: 1849: 1839: 1838: 1832: 1825: 1824: 1821: 1820: 1817: 1816: 1808: 1806: 1796: 1795: 1789: 1782: 1781: 1779: 1778: 1772: 1766: 1760: 1754: 1748: 1741: 1738: 1737: 1730: 1728: 1727: 1720: 1713: 1705: 1699: 1698: 1693: 1684: 1677: 1667: 1659: 1658:External links 1656: 1655: 1654: 1647: 1640: 1633: 1626: 1617: 1614: 1611: 1610: 1584: 1577: 1557:Academic Press 1545:Simon A. Levin 1523: 1520:on 2007-10-25. 1497:(3): 473–484. 1472: 1457:Bingham, C. T. 1443: 1386: 1377: 1363: 1362: 1360: 1357: 1356: 1355: 1349: 1348: 1334: 1331:Insects portal 1318: 1315: 1305: 1302: 1300: 1299: 1281: 1272: 1263: 1261:Boloria selene 1254: 1245: 1236: 1227: 1218: 1209: 1200: 1191: 1189:Vanessa cardui 1182: 1173: 1164: 1158:Mourning cloak 1155: 1146: 1137: 1128: 1119: 1110: 1101: 1092: 1084: 1075: 1073:Junonia coenia 1069:Common buckeye 1066: 1057: 1048: 1039: 1027: 1025: 1022: 1021: 1020: 1001:Plantaginaceae 993:Flacourtiaceae 985:Convolvulaceae 981:Caprifoliaceae 972: 971: 961: 955: 944: 943: 935: 934: 908: 907: 901: 900: 899: 898: 878: 877: 864:Passifloraceae 855: 854: 825: 824: 808: 807: 800: 799: 798: 797: 796:, and Poaceae. 781: 780: 763:Menispermaceae 744: 743: 728: 727: 692:Flacourtiaceae 680:Convolvulaceae 659: 658: 652: 651: 642:Mimics of the 639: 638: 625:satyrine clade 621: 620: 619: 618: 614: 613: 598: 597: 581: 580: 557:(subfamily of 546: 545: 527: 526: 499:The five main 481: 478: 401: 400:Classification 398: 383: 380: 368:tortoiseshells 295:Boloria selene 228: 227: 221: 220: 214: 213: 206: 205: 200: 195: 190: 185: 180: 175: 170: 165: 160: 155: 150: 142: 141: 137: 136: 124: 120: 119: 114: 110: 109: 104: 100: 99: 94: 90: 89: 84: 80: 79: 74: 70: 69: 64: 60: 59: 46: 45: 37: 36: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 3910: 3899: 3898:Papilionoidea 3896: 3894: 3891: 3889: 3886: 3884: 3881: 3880: 3878: 3861: 3856: 3852: 3848: 3843: 3839: 3835: 3830: 3826: 3822: 3816: 3812: 3807: 3803: 3799: 3794: 3790: 3786: 3781: 3777: 3773: 3768: 3764: 3760: 3755: 3751: 3747: 3742: 3738: 3734: 3729: 3725: 3721: 3716: 3712: 3708: 3703: 3699: 3695: 3690: 3686: 3682: 3677: 3673: 3669: 3664: 3660: 3656: 3651: 3647: 3643: 3637: 3633: 3627: 3623: 3618: 3614: 3610: 3605: 3601: 3596: 3590: 3586: 3581: 3575: 3571: 3570: 3568: 3566: 3562: 3558: 3553: 3541: 3538: 3536: 3533: 3532: 3529: 3516: 3510: 3492: 3489: 3488: 3486: 3482: 3476: 3470: 3468: 3465: 3463: 3459:Epicopeiidae 3457: 3455: 3452: 3450: 3447: 3446: 3444: 3442: 3438: 3432: 3429: 3427: 3421: 3419: 3413: 3412: 3410: 3408: 3404: 3398: 3397:Oenosandridae 3395: 3393: 3389:Notodontidae 3387: 3385: 3379: 3377: 3371: 3369: 3366: 3364: 3358: 3357: 3355: 3353: 3349: 3343: 3337: 3335: 3329: 3327: 3324: 3322: 3319: 3317: 3311: 3309: 3303: 3301: 3295: 3293: 3287: 3285: 3281:Apatelodidae 3279: 3277: 3271: 3270: 3268: 3266: 3262: 3256: 3250: 3249: 3247: 3245: 3241: 3235: 3231:Mimallonidae 3229: 3228: 3226: 3224: 3223:Mimallonoidea 3220: 3214: 3208: 3206: 3204:(snout moths) 3200: 3199: 3197: 3195: 3191: 3185: 3179: 3178: 3176: 3174: 3170: 3164: 3158: 3156: 3150: 3148: 3142: 3140: 3134: 3132: 3126: 3124: 3118: 3116: 3112:Papilionidae 3110: 3109: 3107: 3102: 3095: 3094:Papilionoidea 3091: 3085: 3081:Callidulidae 3079: 3078: 3076: 3074: 3073:Calliduloidea 3070: 3064: 3058: 3057: 3055: 3053: 3049: 3043: 3039:Carposinidae 3037: 3035: 3029: 3028: 3026: 3024: 3020: 3014: 3011: 3010: 3008: 3006: 3002: 2996: 2995:Whalleyanidae 2993: 2992: 2990: 2988: 2984: 2978: 2976:(plume moths) 2972: 2971: 2969: 2967: 2963: 2957: 2951: 2949: 2943: 2942: 2940: 2938: 2934: 2928: 2924:Epermeniidae 2922: 2921: 2919: 2917: 2916:Epermenioidea 2913: 2907: 2901: 2900: 2898: 2896: 2892: 2886: 2880: 2879: 2877: 2875: 2871: 2865: 2859: 2858: 2856: 2854: 2850: 2844: 2838: 2837: 2835: 2833: 2829: 2823: 2817: 2815: 2809: 2807: 2801: 2800: 2798: 2796: 2792: 2786: 2782:Dudgeoneidae 2780: 2778: 2772: 2771: 2769: 2767: 2763: 2757: 2751: 2749: 2743: 2741: 2737:Epipyropidae 2735: 2733: 2732:Cyclotornidae 2730: 2728: 2725: 2723: 2720: 2718: 2712: 2710: 2707: 2705: 2702: 2700: 2697: 2695: 2689: 2687: 2686:Heterogynidae 2684: 2683: 2681: 2679: 2675: 2669: 2666: 2665: 2663: 2661: 2660:Galacticoidea 2657: 2651: 2647:Xyloryctidae 2645: 2643: 2639:Scythrididae 2637: 2635: 2632: 2630: 2626:Oecophoridae 2624: 2622: 2616: 2614: 2613:Metachandidae 2611: 2609: 2606: 2604: 2598: 2596: 2590: 2588: 2584:Elachistidae 2582: 2580: 2574: 2572: 2566: 2564: 2563:Blastobasidae 2561: 2559: 2556: 2554: 2553:Autostichidae 2551: 2550: 2548: 2546: 2542: 2536: 2533: 2531: 2525: 2523: 2520: 2518: 2515: 2513: 2510: 2508: 2506:(sedge moths) 2502: 2500: 2497: 2495: 2491:Acrolepiidae 2489: 2488: 2486: 2484: 2480: 2474: 2471: 2469: 2466: 2464: 2460:Douglasiidae 2458: 2456: 2450: 2449: 2447: 2445: 2441: 2435: 2429: 2427: 2421: 2419: 2415:Eriocottidae 2413: 2411: 2408: 2406: 2402:Acrolophidae 2400: 2399: 2397: 2395: 2391: 2385: 2382: 2381: 2379: 2377: 2373: 2369: 2366: 2364: 2346: 2336: 2332:Tischeriidae 2330: 2329: 2327: 2325: 2324:Tischerioidea 2321: 2315: 2309: 2308: 2306: 2304: 2300: 2294: 2288: 2286: 2282:Nepticulidae 2280: 2279: 2277: 2275: 2274:Nepticuloidea 2271: 2265: 2261:Andesianidae 2259: 2258: 2256: 2254: 2253:Andesianoidea 2250: 2244: 2242:(yucca moths) 2238: 2236: 2235:Incurvariidae 2233: 2231: 2228: 2226: 2223: 2221: 2215: 2214: 2212: 2210: 2206: 2203: 2201: 2179: 2176: 2174: 2150: 2136: 2130: 2129: 2127: 2125: 2121: 2115: 2109: 2107: 2101: 2099: 2095:Neotheoridae 2093: 2091: 2085: 2083: 2080: 2079: 2077: 2075: 2071: 2068: 2066: 2062: 2052: 2046: 2044: 2041: 2040: 2038: 2036: 2032: 2029: 2027: 2023: 2015: 2011: 2009: 2005: 2002: 2000: 1999:Lophocoronina 1996: 1986: 1980: 1979: 1977: 1975: 1971: 1968: 1966: 1965:Acanthoctesia 1962: 1952: 1951:Eriocraniidae 1949: 1948: 1946: 1944: 1940: 1937: 1935: 1931: 1927: 1923: 1919: 1913: 1909: 1897: 1893: 1891: 1887: 1883: 1879: 1873: 1869: 1857: 1855:(kauri moths) 1850: 1848: 1844: 1840: 1836: 1830: 1826: 1814: 1807: 1805: 1801: 1797: 1793: 1787: 1783: 1777: 1773: 1771: 1767: 1765: 1761: 1759: 1755: 1753: 1749: 1747: 1743: 1742: 1739: 1734: 1726: 1721: 1719: 1714: 1712: 1707: 1706: 1703: 1697: 1694: 1692: 1688: 1685: 1682: 1678: 1675: 1671: 1668: 1666: 1662: 1661: 1657: 1652: 1648: 1645: 1641: 1638: 1634: 1631: 1627: 1624: 1620: 1619: 1615: 1599: 1595: 1592:Gould, S. E. 1588: 1585: 1580: 1574: 1570: 1566: 1562: 1558: 1553: 1552: 1546: 1542: 1536: 1534: 1532: 1530: 1528: 1524: 1516: 1512: 1508: 1504: 1500: 1496: 1492: 1491: 1483: 1476: 1473: 1469: 1465: 1463: 1458: 1452: 1450: 1448: 1444: 1439: 1435: 1430: 1425: 1421: 1417: 1413: 1409: 1405: 1401: 1397: 1390: 1387: 1381: 1378: 1374: 1368: 1365: 1358: 1354: 1351: 1350: 1346: 1340: 1335: 1332: 1326: 1321: 1316: 1314: 1312: 1303: 1297: 1293: 1289: 1285: 1282: 1280: 1276: 1273: 1271: 1267: 1264: 1262: 1258: 1255: 1253: 1249: 1246: 1244: 1240: 1237: 1235: 1231: 1228: 1226: 1222: 1219: 1217: 1213: 1212:Question mark 1210: 1208: 1204: 1201: 1199: 1195: 1192: 1190: 1186: 1183: 1181: 1177: 1174: 1172: 1168: 1165: 1163: 1159: 1156: 1154: 1150: 1147: 1145: 1141: 1138: 1136: 1132: 1129: 1127: 1123: 1120: 1118: 1114: 1111: 1109: 1105: 1102: 1100: 1096: 1095:Crimson patch 1093: 1091: 1090: 1085: 1083: 1079: 1076: 1074: 1070: 1067: 1065: 1061: 1058: 1056: 1052: 1049: 1047: 1043: 1040: 1038: 1034: 1033: 1029: 1028: 1023: 1018: 1014: 1010: 1006: 1002: 998: 995:, Lamiaceae, 994: 990: 989:Euphorbiaceae 986: 982: 978: 975:Some species 974: 973: 969: 965: 962: 959: 956: 953: 952:Limenitidinae 950:(formerly in 949: 946: 945: 941: 937: 936: 932: 929: 928: 927: 925: 924:Pseudergolini 921: 917: 913: 906: 905:Limenitidinae 903: 902: 896: 892: 888: 887:Sterculiaceae 884: 880: 879: 875: 871: 868: 867: 865: 861: 857: 856: 852: 849: 848: 847: 845: 841: 838: 834: 830: 822: 818: 814: 810: 809: 805: 802: 801: 795: 791: 790:Heliconiaceae 787: 783: 782: 778: 774: 770: 767: 766: 764: 760: 756: 752: 751: 746: 745: 741: 737: 733: 730: 729: 725: 721: 717: 713: 709: 705: 701: 697: 693: 689: 685: 684:Euphorbiaceae 681: 677: 673: 669: 665: 661: 660: 657: 654: 653: 649: 645: 641: 640: 636: 632: 629: 628: 627: 626: 616: 615: 611: 607: 603: 600: 599: 595: 591: 587: 583: 582: 578: 574: 570: 567: 566: 564: 560: 556: 552: 548: 547: 543: 540: 539: 538: 536: 532: 531:danaine clade 524: 520: 517: 516: 515: 513: 509: 504: 502: 497: 495: 491: 487: 479: 477: 475: 471: 467: 463: 459: 458: 453: 452: 447: 446: 441: 436: 433: 428: 425: 420: 418: 414: 407: 399: 397: 395: 391: 390: 381: 379: 377: 373: 369: 365: 361: 357: 353: 349: 345: 344:Papilionoidea 341: 337: 333: 325: 321: 317: 297: 296: 290: 283: 282: 276: 269: 268: 262: 255: 254: 248: 242: 240: 234: 222: 219: 215: 212: 211: 204: 201: 199: 196: 194: 191: 189: 188:Limenitidinae 186: 184: 181: 179: 176: 174: 171: 169: 166: 164: 161: 159: 156: 154: 151: 149: 146: 145: 143: 138: 133: 128: 125: 122: 121: 118: 117:Papilionoidea 115: 113:Superfamily: 112: 111: 108: 105: 102: 101: 98: 95: 92: 91: 88: 85: 82: 81: 78: 75: 72: 71: 68: 65: 62: 61: 56: 51: 47: 43: 38: 33: 30: 19: 3564: 3491:Millieriidae 3472:Geometridae 3441:Geometroidea 3425:(gold moths) 3383:(tuft moths) 3333:(saturniids) 3331:Saturniidae 3321:Eupterotidae 3305:Carthaeidae 3297:Brahmaeidae 3291:(silk moths) 3212:(grass moth) 3183:(teak moths) 3160:Nymphalidae 3159: 3146:(metalmarks) 3128:Hesperiidae 2861:Tortricidae 2853:Tortricoidea 2840:Choreutidae 2832:Choreutoidea 2803:Brachodidae 2753:Limacodidae 2727:Anomoeotidae 2668:Galacticidae 2592:Gelechiidae 2545:Gelechioidea 2535:Ypsolophidae 2512:Heliodinidae 2290:Opostegidae 2230:Heliozelidae 2082:Anomosetidae 2026:Neopseustina 1776:Holometabola 1774:Superorder: 1768:Infraclass: 1679:Peter Chew: 1673: 1650: 1643: 1636: 1629: 1622: 1601:. Retrieved 1597: 1587: 1550: 1515:the original 1494: 1488: 1475: 1460: 1403: 1399: 1389: 1380: 1372: 1367: 1307: 1295: 1287: 1278: 1269: 1260: 1251: 1242: 1233: 1224: 1215: 1206: 1197: 1188: 1185:Painted lady 1179: 1170: 1161: 1152: 1149:Meadow brown 1143: 1134: 1125: 1116: 1107: 1098: 1087: 1081: 1072: 1063: 1054: 1045: 1036: 1030: 999:, Moraceae, 997:Loranthaceae 991:, Fagaceae, 916:sister group 911: 909: 851:Heliconiinae 833:sister group 828: 826: 786:Bromeliaceae 761:, Fabaceae, 759:Bignoniaceae 748: 740:Amathusiinae 676:Celastraceae 624: 622: 590:Gesneriaceae 530: 528: 507: 505: 498: 483: 455: 451:Pseudergolis 449: 443: 437: 429: 421: 413:caterpillars 410: 393: 387: 385: 382:Nomenclature 372:fritillaries 351: 347: 331: 329: 293: 279: 265: 251: 237: 210:article text 207: 178:Heliconiinae 140:Subfamilies 126: 35:Nymphalidae 29: 3883:Nymphalidae 3754:iNaturalist 3632:Nymphalidae 3622:Nymphalidae 3609:Nymphalidae 3595:Nymphalidae 3589:Wikispecies 3565:Nymphalidae 3449:Sematuridae 3423:Cimeliidae 3417:(hook-tips) 3415:Drepanidae 3407:Drepanoidea 3339:Sphingidae 3326:Phiditiidae 3313:Endromidae 3289:Bombycidae 3273:Anthelidae 3265:Bombycoidea 3181:Hyblaeidae 3173:Hyblaeoidea 3152:Lycaenidae 3144:Riodinidae 3101:butterflies 3060:Thyrididae 3052:Thyridoidea 2953:Tineodidae 2945:Alucitidae 2937:Alucitoidea 2811:Castniidae 2745:Dalceridae 2691:Zygaenidae 2678:Zygaenoidea 2522:Plutellidae 2517:Lyonetiidae 2499:Bedelliidae 2240:Prodoxidae 2225:Cecidosidae 2087:Hepialidae 2074:Hepialoidea 1792:Zeugloptera 1733:Lepidoptera 1691:Nymphalidae 1674:Insecta.pro 1670:Nymphalidae 1559:. pp.  1230:Small heath 1221:Red admiral 1203:Plain tiger 1176:Blue morpho 1017:Verbenaceae 1009:Sapindaceae 964:Nymphalinae 777:Brassolinae 773:crepuscular 736:Amathusiini 734:(including 724:Sapindaceae 720:Santalaceae 631:Calinaginae 610:Tellervinae 606:Australasia 586:Apocynaceae 573:Neotropical 571:(about 300 559:Apocynaceae 551:Apocynaceae 519:Libytheinae 494:subfamilies 340:butterflies 332:Nymphalidae 198:Nymphalinae 183:Libytheinae 158:Calinaginae 127:Nymphalidae 107:Lepidoptera 3877:Categories 3515:Monotrysia 3373:Noctuidae 3368:Euteliidae 3352:Noctuoidea 3210:Crambidae 3202:Pyralidae 3194:Pyraloidea 3130:(skippers) 3120:Hedylidae 2704:Lacturidae 2618:Momphidae 2423:Psychidae 1934:Dacnonypha 1762:Subclass: 1752:Arthropoda 1466:. London: 1359:References 1304:Morphology 1248:Gatekeeper 1005:Salicaceae 968:Biblidinae 958:Cyrestinae 948:Biblidinae 931:Apaturinae 895:Urticaceae 883:Asteraceae 817:Cyperaceae 769:Brassolini 712:Rhamnaceae 704:Piperaceae 672:Annonaceae 656:Charaxinae 602:Tellervini 594:Solanaceae 577:Ithomiinae 417:chrysalids 318:butterfly 168:Cyrestinae 163:Charaxinae 153:Biblidinae 148:Apaturinae 132:Rafinesque 87:Arthropoda 3467:Uraniidae 3360:Erebidae 3136:Pieridae 2882:Urodidae 2874:Urodoidea 2819:Sesiidae 2795:Sesioidea 2774:Cossidae 2766:Cossoidea 2608:Lypusidae 2431:Tineidae 2394:Tineoidea 2217:Adelidae 2209:Adeloidea 1916:Suborder 1876:Suborder 1835:Aglossata 1833:Suborder 1790:Suborder 1764:Pterygota 1744:Kingdom: 1420:1536-2442 1406:(1): 66. 1198:Aglais io 1089:Hamadryas 1042:Archdukes 891:Tiliaceae 874:Acraeinae 844:Cyrestini 840:Biblidini 804:Satyrinae 755:Arecaceae 732:Morphinae 700:Myrtaceae 696:Lauraceae 662:Tropical 635:Himalayas 569:Ithomiini 486:phylogeny 470:Satyrinae 432:hindwings 424:forewings 394:Nymphalis 389:Nymphalis 320:nectaring 239:Aglais io 236:Peacock ( 218:Diversity 203:Satyrinae 193:Morphinae 73:Kingdom: 67:Eukaryota 18:Nymphalid 3663:BugGuide 3639:BioLib: 3629:BAMONA: 3574:Wikidata 3381:Nolidae 3005:Immoidea 2065:Exoporia 1918:Glossata 1770:Neoptera 1750:Phylum: 1746:Animalia 1735:families 1511:12927132 1459:(1905). 1438:21867433 1317:See also 1044:, genus 1037:Actinote 1035:, genus 940:Ulmaceae 870:Acraeini 794:Musaceae 716:Rutaceae 688:Fabaceae 648:Moraceae 644:Danainae 563:Moraceae 542:Danainae 466:Danainae 457:Calinaga 445:Libythea 440:antennae 364:admirals 356:emperors 316:Crescent 208:and see 173:Danainae 123:Family: 83:Phylum: 77:Animalia 63:Domain: 3580:Q156449 3013:Immidae 2722:Aididae 1758:Insecta 1756:Class: 1731:Extant 1547:(ed.). 1429:3281478 1194:Peacock 977:migrate 813:Araceae 708:Poaceae 561:), and 462:atrophy 376:cryptic 103:Order: 97:Insecta 93:Class: 3860:989033 3847:135150 3834:358161 3818:NZOR: 3785:117276 3772:114362 3707:1NYMPF 3431:Doidae 1653:(2002) 1646:(2008) 1639:(2011) 1632:(2001) 1625:(2001) 1575:  1563:–573. 1509:  1436:  1426:  1418:  1046:Lexias 920:Coeini 893:, and 837:tribes 750:Morpho 722:, and 664:canopy 592:, and 501:clades 474:thorax 454:, and 370:, and 336:family 134:, 1815 3855:WoRMS 3811:33415 3767:IRMNG 3759:47922 3642:17299 1603:7 Sep 1518:(PDF) 1485:(PDF) 1311:setae 1060:Comma 535:basal 512:basal 3806:NCBI 3780:ITIS 3746:7017 3741:GBIF 3720:7202 3702:EPPO 3650:BOLD 1605:2013 1573:ISBN 1507:PMID 1434:PMID 1416:ISSN 922:and 910:The 842:and 827:The 623:The 529:The 506:The 490:taxa 484:The 468:and 438:The 430:The 422:The 330:The 3793:NBN 3694:876 3689:EoL 3681:DGC 3676:CoL 3668:192 3655:723 3617:AFD 3604:ADW 1565:doi 1561:559 1499:doi 1424:PMC 1408:doi 350:or 338:of 322:on 3879:: 3857:: 3844:: 3831:: 3808:: 3795:: 3782:: 3769:: 3756:: 3743:: 3730:: 3717:: 3704:: 3691:: 3678:: 3665:: 3652:: 3619:: 3606:: 3591:: 3576:: 1689:: 1596:. 1571:. 1555:. 1526:^ 1505:. 1495:28 1493:. 1487:. 1446:^ 1432:. 1422:. 1414:. 1404:11 1402:. 1398:. 1294:, 1286:, 1277:, 1268:, 1259:, 1250:, 1241:, 1232:, 1223:, 1214:, 1205:, 1196:, 1187:, 1178:, 1169:, 1160:, 1151:, 1142:, 1133:, 1124:, 1115:, 1106:, 1097:, 1080:, 1071:, 1062:, 1053:, 1011:, 1007:, 987:, 983:, 926:) 889:, 866:. 815:, 792:, 788:, 757:, 718:, 714:, 710:, 706:, 702:, 698:, 694:, 690:, 686:, 682:, 678:, 674:, 588:, 565:. 553:, 537:) 514:) 448:, 366:, 362:, 358:, 3103:) 3099:( 2363:a 2361:i 2359:s 2357:y 2355:r 2353:t 2351:i 2349:D 2200:a 2198:i 2196:s 2194:y 2192:r 2190:t 2188:o 2186:n 2184:o 2182:M 2173:a 2171:r 2169:u 2167:e 2165:n 2163:o 2161:r 2159:e 2157:t 2155:e 2153:H 1724:e 1717:t 1710:v 1676:. 1607:. 1581:. 1567:: 1501:: 1440:. 1410:: 1298:) 1019:. 970:) 954:) 914:( 897:. 876:) 831:( 823:. 779:) 742:) 726:. 650:. 637:) 612:) 596:. 579:) 533:( 521:( 510:( 408:. 326:. 298:) 284:) 270:) 256:) 241:) 20:)

Index

Nymphalid

Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Lepidoptera
Papilionoidea
Nymphalidae
Rafinesque
Apaturinae
Biblidinae
Calinaginae
Charaxinae
Cyrestinae
Danainae
Heliconiinae
Libytheinae
Limenitidinae
Morphinae
Nymphalinae
Satyrinae
article text
Diversity

Aglais io

Euphaedra themis

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