238:(or Archaediptera) which includes several Triassic fossil members. The family has characteristics associated with the Nematocera as well as the Brachycera. The antennae are shortened as in the Brachycera and these flies are long, having a snout with vestigeal mouthparts, non-differentiated abdominal segments with large cerci. The wings are narrow and hair-fringed and have very weak venation. They are known to form cloud-like swarms in summer and the short-lived non-feeding adults have wings that fracture at the base shortly after mating.
634:
117:
2967:
233:
in northern regions of the world, including northeastern North
America, Japan, the Himalayas, and eastern Russia. Around a dozen extant species are known, with two fossil species found in amber, extending back to the Mid Cretaceous. Under an alternative classification, they are considered the only
293:
have atrophied mouthparts. Nymphomyiidae are unusual in that the adults are ventrally holoptic, meaning they possess two eyes that meet on the underside of the head. Adults form large swarms above water. One or two generations may breed in a single year depending on the region and climate.
289:, retaining various larval features. They have strap-like wings with a very reduced venation, and the wing margins have long fringes like those of the Thysanoptera. The wings break at the base after mating. The antennae are very reduced. Species in the genus
269:
so distinct that he put it in a separate superfamily
Nymphomyioidea related to Triassic Dictyodipteridae which are in a suborder Archidiptera. Modern classifications put all the species in a single genus
502:
Schneeberg, Katharina; Friedrich, Frank; Courtney, Gregory W.; Wipfler, Benjamin; Beutel, Rolf G. (2012). "The larvae of
Nymphomyiidae (Diptera, Insecta) – Ancestral and highly derived?".
549:
Harper, P.P.; Lauzon, Michel (2012). "Life Cycle of the Nymph Fly
Palaeodipteron Walkeri Ide 1965 (Diptera: Nymphomyiidae) in the White Mountains of Southern Québec".
2887:
2913:
755:
3008:
2861:
2900:
646:
Courtney, G. 1998. First records of the
Nymphomyiidae (Diptera) in Nepal. Proceedings of the Entomological Society of Washington 100: 595–597.
3037:
613:
Courtney, G.W. (1994). "Biosystematics of the
Nymphomyiidae (Insecta: Diptera): life history, morphology, and phylogenetic relationships".
683:
Rüdiger Wagner; Patrick Müller (2020). "The oldest of the rare: A new species of the family
Nymphomyiidae (Diptera) from Burmese amber".
748:
2905:
458:
Ide, F.P. 1965. A fly of the archaic family
Nymphomyiidae (Diptera) from North America. Canadian Entomologist, 97: 496-507.
3001:
710:
Wagner, R.; Hoffeins, C.; Hoffeins, H. W. (2000). "A fossil nymphomyiid (Diptera) from the Baltic and
Bitterfeld amber".
530:
3027:
576:
D. Huang; A. Nel; C. Cai; Q. Lin & M. S. Engel (2013). "Amphibious flies and paedomorphism in the
Jurassic period".
529:
Pape, Thomas; Blagoderov, Vladimir; Mostovski, Michail B. (2011). "Order Diptera Linnaeus, 1758". In Z. Q. Zhang (ed.).
116:
3032:
2809:
2918:
3042:
741:
2994:
2737:
370:
354:
330:
633:
241:
The family Nymphomyiidae has several species which were originally placed in separate genera of their own.
2752:
1916:
346:
2814:
378:
338:
2939:
390:
362:
246:
203:
2848:
322:
2585:
414:
Tokunaga M. (1932). "A remarkable dipterous insect from Japan, Nymphomyia alba, gen. et sp. nov".
2790:
2409:
977:
111:
2944:
314:
656:
Ide, F. P. (2012). "A Fly of the Archaic Family Nymphomyiidae Found in New Brunswick in 1961".
532:
Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness
2926:
2835:
1262:
595:
275:
2892:
2637:
2543:
2526:
2404:
1846:
1449:
1206:
1117:
719:
692:
665:
622:
587:
578:
558:
511:
484:
218:
2931:
2254:
2249:
2114:
2013:
1962:
1799:
1480:
1345:
1171:
1163:
985:
944:
723:
475:
alba(Diptera: Nymphomyiidae) from Japan with notes on the larvae and the micro-habitat".
245:, the type species for this family, was discovered in a fast-flowing stream in Japan by
2978:
2535:
2461:
2375:
2312:
2228:
2163:
2145:
2130:
2036:
2031:
1880:
1658:
1579:
1535:
1496:
1270:
1189:
1130:
1125:
993:
891:
434:
257:
was described from the Himalayas in 1970. Hennig examined the pupal characteristics of
3021:
2676:
2668:
2659:
2551:
2485:
2477:
2430:
2396:
2212:
2168:
2073:
2065:
1987:
1977:
1921:
1893:
1783:
1642:
1595:
1564:
1316:
1308:
1219:
1099:
1091:
1019:
920:
383:
70:
2840:
2707:
2699:
2690:
2632:
2593:
2576:
2451:
2351:
2288:
2196:
2173:
2104:
2049:
2026:
1982:
1972:
1967:
1957:
1936:
1851:
1812:
1775:
1745:
1703:
1682:
1650:
1551:
1501:
1299:
1198:
1109:
1057:
1049:
952:
928:
866:
808:
298:
235:
2966:
2874:
2784:
2624:
2603:
2334:
2317:
2293:
2280:
2204:
2183:
2096:
2088:
2078:
2021:
1992:
1952:
1926:
1817:
1791:
1736:
1620:
1587:
1530:
1467:
1424:
1278:
1237:
1229:
1211:
1104:
1086:
899:
262:
45:
17:
2775:
696:
274:. Based on larval morphology, the family has been suggested to be close to the
2974:
2616:
2559:
2493:
2469:
2443:
2388:
2220:
2191:
2158:
2135:
2057:
1944:
1911:
1898:
1872:
1807:
1695:
1522:
1509:
1454:
1404:
1361:
1353:
1070:
1036:
1028:
875:
848:
840:
824:
515:
488:
301:
from the Jurassic of Asia, which are thought to have had a similar lifestyle.
178:
90:
55:
2642:
2598:
2435:
2425:
2359:
2343:
2267:
2259:
2241:
2153:
2140:
2122:
2109:
1888:
1838:
1825:
1762:
1754:
1666:
1569:
1556:
1543:
1488:
1433:
1369:
1329:
1138:
936:
796:
784:
626:
148:
128:
95:
599:
2801:
733:
2822:
2769:
2367:
2275:
2000:
1903:
1859:
1711:
1674:
1459:
1441:
907:
802:
85:
80:
65:
60:
50:
2827:
591:
562:
1633:
883:
669:
286:
100:
75:
2879:
2417:
1475:
790:
778:
230:
158:
138:
2866:
2746:
309:
Currently all species are treated as members of a single genus:
226:
2750:
1401:
821:
737:
297:
Nymphomyiidae are thought to be closely related to the extinct
765:
222:
168:
2853:
2982:
538:. Magnolia Press. Zootaxa, Vol. 3148. pp. 222–229.
278:
while others place them in a separate infraorder, the
234:
living representatives of a separate, suborder called
435:"Insect mist-swarming of Nymphomyia species in Japan"
2973:
This article related to members of the fly suborder
2759:
2698:
2689:
2667:
2658:
2615:
2584:
2575:
2534:
2525:
2460:
2387:
2342:
2333:
2305:
2240:
2182:
2087:
2048:
2012:
1935:
1871:
1837:
1774:
1753:
1744:
1735:
1694:
1641:
1632:
1619:
1578:
1521:
1432:
1423:
1328:
1307:
1298:
1261:
1228:
1197:
1188:
1162:
1069:
1048:
1027:
1018:
976:
919:
874:
865:
839:
1677:(scuttle flies, coffin flies, humpbacked flies)
1214:(minute black scavenger flies, or dung midges)
3002:
749:
471:Takemon, Y.; Tanida, K. (1994). "New data on
8:
225:(Diptera). Larvae are found among aquatic
3009:
2995:
2747:
2695:
2664:
2581:
2531:
2522:
2339:
1750:
1741:
1732:
1638:
1629:
1625:
1429:
1420:
1416:
1412:
1398:
1304:
1295:
1258:
1194:
1185:
1159:
1024:
1015:
973:
871:
862:
836:
832:
818:
756:
742:
734:
433:Saigusa, T.; T. Nakamura; S. Sato (2009).
31:
334:Rohdendorf & Kalugina, 1974 - Siberia
466:
464:
2399:(blow-flies: bluebottles, greenbottles)
406:
253:described by Ide in Quebec in 1965 and
221:of tiny (2 mm) slender, delicate
504:Arthropod Structure & Development
7:
2963:
2961:
724:10.1046/j.1365-3113.2000.025001115.x
615:Smithsonian Contributions to Zoology
1862:(cactus flies, banana stalk flies)
416:Annotationes Zoologicae Japonenses
25:
2965:
632:
115:
374:Courtney, 1994 - North America
249:in 1932. This was followed by
1:
1546:(dagger flies, balloon flies)
3038:Monogeneric Diptera families
2981:. You can help Knowledge by
261:and placed it in the family
2738:List of families of Diptera
2370:(house flies, stable flies)
358:Makarchenko, 1979 - Siberia
3059:
2960:
1141:(dark-winged fungus gnats)
1133:(long-winged fungus gnats)
1120:(long-beaked fungus gnats)
697:10.11646/zootaxa.4763.2.11
2735:
2521:
2215:(vinegar and fruit flies)
2125:(lekking, or druid flies)
1731:
1628:
1419:
1415:
1411:
1397:
1348:(long-bodied crane flies)
1294:
1257:
1184:
1158:
1014:
972:
861:
835:
831:
817:
773:
658:The Canadian Entomologist
551:The Canadian Entomologist
516:10.1016/j.asd.2012.01.002
489:10.1080/01650429409361544
350:Courtney 1994 - Hong Kong
342:(Kevan, 1970) - Himalayas
112:Scientific classification
110:
34:
1112:(predatory fungus gnats)
265:. Rohdendorf considered
2003:(black scavenger flies)
1364:(hairy-eyed craneflies)
1281:(primitive crane flies)
1039:(march flies, lovebugs)
627:10.5479/si.00810282.550
2645:(horse and deer flies)
1917:Strongylophthalmyiidae
1828:(picture-winged flies)
1356:(limoniid crane flies)
371:Nymphomyia dolichopeza
326:Tokunaga, 1932 - Japan
251:Palaeodipteron walkeri
2940:Paleobiology Database
2231:(frightful hairy fly)
2060:(beetle-backed flies)
1598:(tangle-veined flies)
1483:(hilarimorphid flies)
1444:(flower-loving flies)
1273:(phantom crane flies)
712:Systematic Entomology
255:Felicitomyia brundini
2562:(wood soldier flies)
2412:(New Zealand batfly)
2362:(little house flies)
1854:(stilt-legged flies)
1765:(thick-headed flies)
1661:(spear-winged flies)
1590:(small-headed flies)
1319:(winter crane flies)
894:(frog-biting midges)
366:(Ide, 1964) - Canada
3028:Nematocera families
2627:(water snipe flies)
2148:(upside-down flies)
1685:(flat-footed flies)
1669:(flat-footed flies)
1538:(long-legged flies)
1372:(large crane flies)
988:(net-winged midges)
955:(non-biting midges)
592:10.1038/nature11898
563:10.4039/Ent121603-7
379:Nymphomyia allissae
3033:Blephariceromorpha
2296:(freeloader flies)
2099:(leaf miner flies)
2039:(small dung flies)
1883:(stalk-eyed flies)
1706:(big-headed flies)
1164:Perissommatomorpha
994:Deuterophlebiidae
978:Blephariceromorpha
670:10.4039/Ent96119-1
391:Nymphomyia succina
285:Nymphomyiidae are
2990:
2989:
2955:
2954:
2927:Open Tree of Life
2753:Taxon identifiers
2744:
2743:
2731:
2730:
2727:
2726:
2723:
2722:
2719:
2718:
2711:
2685:
2684:
2654:
2653:
2646:
2628:
2607:
2571:
2570:
2563:
2555:
2547:
2544:Pantophthalmidae
2517:
2516:
2513:
2512:
2509:
2508:
2505:
2504:
2497:
2489:
2481:
2473:
2447:
2439:
2421:
2413:
2400:
2379:
2371:
2363:
2355:
2329:
2328:
2321:
2297:
2284:
2271:
2263:
2250:Acartophthalmidae
2232:
2224:
2216:
2208:
2200:
2199:(quasimodo flies)
2149:
2126:
2118:
2100:
2069:
2061:
2040:
2004:
1996:
1948:
1907:
1884:
1863:
1855:
1829:
1821:
1803:
1795:
1787:
1766:
1727:
1726:
1723:
1722:
1715:
1707:
1686:
1678:
1670:
1662:
1654:
1615:
1614:
1611:
1610:
1607:
1606:
1599:
1591:
1560:
1547:
1539:
1513:
1505:
1492:
1484:
1471:
1463:
1445:
1393:
1392:
1389:
1388:
1385:
1384:
1381:
1380:
1373:
1365:
1357:
1349:
1339:
1320:
1290:
1289:
1282:
1274:
1263:Ptychopteromorpha
1253:
1252:
1249:
1248:
1241:
1215:
1180:
1179:
1154:
1153:
1150:
1149:
1142:
1134:
1121:
1118:Lygistorrhinidae
1113:
1095:
1080:
1061:
1040:
1010:
1009:
997:
996:(mountain midges)
989:
968:
967:
964:
963:
956:
948:
940:
932:
931:(solitary midges)
911:
903:
895:
887:
886:(meniscus midges)
857:
856:
276:Deuterophlebiidae
211:
210:
207:
41:Cenomanian–Recent
16:(Redirected from
3050:
3043:Nematocera stubs
3011:
3004:
2997:
2969:
2962:
2948:
2947:
2935:
2934:
2922:
2921:
2909:
2908:
2896:
2895:
2883:
2882:
2870:
2869:
2857:
2856:
2844:
2843:
2831:
2830:
2818:
2817:
2805:
2804:
2795:
2794:
2793:
2780:
2779:
2778:
2748:
2709:
2696:
2665:
2660:Vermileonomorpha
2644:
2638:Pelecorhynchidae
2626:
2605:
2582:
2561:
2553:
2545:
2532:
2527:Stratiomyomorpha
2523:
2495:
2487:
2479:
2471:
2445:
2437:
2419:
2411:
2410:Mystacinobiidae
2405:Mesembrinellidae
2398:
2377:
2369:
2361:
2353:
2340:
2319:
2295:
2282:
2269:
2261:
2230:
2222:
2214:
2206:
2198:
2147:
2124:
2116:
2115:Aulacigastridae
2098:
2067:
2059:
2038:
2002:
1994:
1946:
1922:Syringogastridae
1905:
1882:
1861:
1853:
1847:Cypselosomatidae
1827:
1819:
1801:
1800:Platystomatidae
1793:
1785:
1764:
1751:
1742:
1733:
1713:
1705:
1684:
1676:
1668:
1660:
1652:
1639:
1630:
1626:
1597:
1589:
1558:
1545:
1537:
1512:(stiletto flies)
1511:
1503:
1490:
1482:
1481:Hilarimorphidae
1469:
1461:
1450:Apsilocephalidae
1443:
1430:
1421:
1417:
1413:
1399:
1371:
1363:
1355:
1347:
1346:Cylindrotomidae
1338:
1337:
1333:
1318:
1305:
1296:
1280:
1272:
1259:
1239:
1213:
1207:Canthyloscelidae
1195:
1186:
1160:
1140:
1132:
1119:
1111:
1093:
1079:
1078:
1074:
1059:
1038:
1025:
1016:
995:
987:
986:Blephariceridae
974:
954:
946:
945:Ceratopogonidae
938:
930:
909:
902:(phantom midges)
901:
893:
885:
872:
863:
837:
833:
819:
758:
751:
744:
735:
728:
727:
707:
701:
700:
680:
674:
673:
664:(1–2): 119–120.
653:
647:
644:
638:
637:
636:
630:
610:
604:
603:
573:
567:
566:
546:
540:
539:
537:
526:
520:
519:
499:
493:
492:
468:
459:
456:
450:
449:
439:
430:
424:
423:
411:
394:† (Baltic amber)
280:Nymphomyiomorpha
247:Masaaki Tokunaga
229:in small, rapid
202:
189:Nymphomyiomorpha
120:
119:
105:
42:
38:Temporal range:
32:
21:
18:Nymphomyiomorpha
3058:
3057:
3053:
3052:
3051:
3049:
3048:
3047:
3018:
3017:
3016:
3015:
2958:
2956:
2951:
2943:
2938:
2930:
2925:
2917:
2912:
2904:
2899:
2891:
2886:
2878:
2873:
2865:
2860:
2852:
2847:
2839:
2834:
2826:
2821:
2813:
2808:
2800:
2798:
2789:
2788:
2783:
2774:
2773:
2768:
2755:
2745:
2740:
2715:
2691:Xylophagomorpha
2681:
2650:
2611:
2567:
2554:(soldier flies)
2501:
2456:
2446:(tachina flies)
2383:
2376:Scathophagidae
2354:(cabbage flies)
2325:
2301:
2255:Australimyzidae
2236:
2229:Mormotomyiidae
2178:
2146:Neurochaetidae
2083:
2044:
2037:Sphaeroceridae
2014:Sphaeroceroidea
2008:
1968:Heterocheilidae
1963:Helosciomyzidae
1931:
1867:
1833:
1820:(peacock flies)
1786:(flutter flies)
1770:
1719:
1690:
1659:Lonchopteridae
1603:
1574:
1552:Homalocnemiidae
1536:Dolichopodidae
1517:
1407:
1377:
1335:
1334:
1332:
1324:
1286:
1271:Ptychopteridae
1245:
1224:
1176:
1172:Perissommatidae
1146:
1131:Rangomaramidae
1076:
1075:
1073:
1065:
1044:
1006:
960:
947:(biting midges)
915:
892:Corethrellidae
853:
827:
813:
769:
762:
732:
731:
709:
708:
704:
682:
681:
677:
655:
654:
650:
645:
641:
631:
612:
611:
607:
586:(7439): 94–97.
575:
574:
570:
548:
547:
543:
535:
528:
527:
523:
501:
500:
496:
477:Aquatic Insects
470:
469:
462:
457:
453:
437:
432:
431:
427:
413:
412:
408:
403:
318:Tokunaga, 1932
307:
243:Nymphomyia alba
201:
114:
106:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
40:
39:
36:
28:
27:Family of flies
23:
22:
15:
12:
11:
5:
3056:
3054:
3046:
3045:
3040:
3035:
3030:
3020:
3019:
3014:
3013:
3006:
2999:
2991:
2988:
2987:
2970:
2953:
2952:
2950:
2949:
2936:
2923:
2910:
2897:
2884:
2871:
2858:
2845:
2832:
2819:
2806:
2796:
2781:
2765:
2763:
2757:
2756:
2751:
2742:
2741:
2736:
2733:
2732:
2729:
2728:
2725:
2724:
2721:
2720:
2717:
2716:
2714:
2713:
2704:
2702:
2693:
2687:
2686:
2683:
2682:
2680:
2679:
2673:
2671:
2669:Vermileonoidea
2662:
2656:
2655:
2652:
2651:
2649:
2648:
2640:
2635:
2630:
2621:
2619:
2613:
2612:
2610:
2609:
2601:
2596:
2594:Austroleptidae
2590:
2588:
2579:
2573:
2572:
2569:
2568:
2566:
2565:
2557:
2552:Stratiomyidae
2549:
2546:(timber flies)
2540:
2538:
2536:Stratiomyoidea
2529:
2519:
2518:
2515:
2514:
2511:
2510:
2507:
2506:
2503:
2502:
2500:
2499:
2491:
2486:Nycteribiidae
2483:
2478:Hippoboscidae
2475:
2472:(tsetse flies)
2466:
2464:
2462:Hippoboscoidea
2458:
2457:
2455:
2454:
2449:
2441:
2436:Sarcophagidae
2433:
2428:
2423:
2415:
2407:
2402:
2397:Calliphoridae
2393:
2391:
2385:
2384:
2382:
2381:
2373:
2365:
2357:
2348:
2346:
2337:
2331:
2330:
2327:
2326:
2324:
2323:
2315:
2313:Cryptochetidae
2309:
2307:
2303:
2302:
2300:
2299:
2291:
2286:
2278:
2273:
2265:
2257:
2252:
2246:
2244:
2238:
2237:
2235:
2234:
2226:
2218:
2213:Drosophilidae
2210:
2202:
2194:
2188:
2186:
2180:
2179:
2177:
2176:
2171:
2166:
2164:Periscelididae
2161:
2156:
2151:
2143:
2138:
2133:
2131:Fergusoninidae
2128:
2120:
2112:
2107:
2102:
2093:
2091:
2085:
2084:
2082:
2081:
2076:
2071:
2066:Chamaemyiidae
2063:
2054:
2052:
2046:
2045:
2043:
2042:
2034:
2032:Nannodastiidae
2029:
2024:
2018:
2016:
2010:
2009:
2007:
2006:
1998:
1990:
1985:
1980:
1975:
1970:
1965:
1960:
1955:
1950:
1941:
1939:
1933:
1932:
1930:
1929:
1924:
1919:
1914:
1909:
1901:
1896:
1891:
1886:
1877:
1875:
1869:
1868:
1866:
1865:
1857:
1849:
1843:
1841:
1835:
1834:
1832:
1831:
1823:
1815:
1810:
1805:
1802:(signal flies)
1797:
1794:(cheese flies)
1789:
1784:Pallopteridae
1780:
1778:
1772:
1771:
1769:
1768:
1759:
1757:
1748:
1739:
1729:
1728:
1725:
1724:
1721:
1720:
1718:
1717:
1709:
1700:
1698:
1692:
1691:
1689:
1688:
1680:
1672:
1664:
1656:
1653:(ironic flies)
1647:
1645:
1636:
1623:
1617:
1616:
1613:
1612:
1609:
1608:
1605:
1604:
1602:
1601:
1596:Nemestrinidae
1593:
1584:
1582:
1580:Nemestrinoidea
1576:
1575:
1573:
1572:
1567:
1562:
1554:
1549:
1541:
1533:
1527:
1525:
1519:
1518:
1516:
1515:
1507:
1504:(window flies)
1499:
1497:Mythicomyiidae
1494:
1486:
1478:
1473:
1465:
1462:(robber flies)
1457:
1452:
1447:
1438:
1436:
1427:
1409:
1408:
1402:
1395:
1394:
1391:
1390:
1387:
1386:
1383:
1382:
1379:
1378:
1376:
1375:
1367:
1359:
1351:
1342:
1340:
1326:
1325:
1323:
1322:
1317:Trichoceridae
1313:
1311:
1309:Trichoceroidea
1302:
1292:
1291:
1288:
1287:
1285:
1284:
1276:
1267:
1265:
1255:
1254:
1251:
1250:
1247:
1246:
1244:
1243:
1234:
1232:
1226:
1225:
1223:
1222:
1217:
1209:
1203:
1201:
1192:
1190:Psychodomorpha
1182:
1181:
1178:
1177:
1175:
1174:
1168:
1166:
1156:
1155:
1152:
1151:
1148:
1147:
1145:
1144:
1136:
1128:
1126:Mycetophilidae
1123:
1115:
1107:
1102:
1097:
1092:Cecidomyiidae
1089:
1087:Bolitophilidae
1083:
1081:
1077:(fungus gnats)
1067:
1066:
1064:
1063:
1054:
1052:
1046:
1045:
1043:
1042:
1033:
1031:
1022:
1012:
1011:
1008:
1007:
1005:
1004:
999:
991:
982:
980:
970:
969:
966:
965:
962:
961:
959:
958:
950:
942:
934:
925:
923:
917:
916:
914:
913:
905:
897:
889:
880:
878:
869:
859:
858:
855:
854:
852:
851:
845:
843:
829:
828:
822:
815:
814:
812:
811:
805:
799:
793:
787:
781:
774:
771:
770:
763:
761:
760:
753:
746:
738:
730:
729:
702:
691:(2): 287–293.
675:
648:
639:
605:
568:
541:
521:
494:
460:
451:
425:
405:
404:
402:
399:
398:
397:
396:
395:
387:
375:
367:
359:
355:N. rohdendorfi
351:
343:
335:
331:N. levanidovae
327:
306:
303:
209:
208:
196:
192:
191:
186:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
146:
142:
141:
136:
132:
131:
126:
122:
121:
108:
107:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
43:
37:
26:
24:
14:
13:
10:
9:
6:
4:
3:
2:
3055:
3044:
3041:
3039:
3036:
3034:
3031:
3029:
3026:
3025:
3023:
3012:
3007:
3005:
3000:
2998:
2993:
2992:
2986:
2984:
2980:
2976:
2971:
2968:
2964:
2959:
2946:
2941:
2937:
2933:
2928:
2924:
2920:
2915:
2911:
2907:
2902:
2898:
2894:
2889:
2885:
2881:
2876:
2872:
2868:
2863:
2859:
2855:
2850:
2846:
2842:
2837:
2833:
2829:
2824:
2820:
2816:
2811:
2807:
2803:
2797:
2792:
2791:Nymphomyiidae
2786:
2782:
2777:
2771:
2767:
2766:
2764:
2762:
2761:Nymphomyiidae
2758:
2754:
2749:
2739:
2734:
2712:
2708:Xylophagidae
2706:
2705:
2703:
2701:
2700:Xylophagoidea
2697:
2694:
2692:
2688:
2678:
2677:Vermileonidae
2675:
2674:
2672:
2670:
2666:
2663:
2661:
2657:
2647:
2641:
2639:
2636:
2634:
2631:
2629:
2623:
2622:
2620:
2618:
2614:
2608:
2606:(snipe flies)
2602:
2600:
2597:
2595:
2592:
2591:
2589:
2587:
2583:
2580:
2578:
2574:
2564:
2558:
2556:
2550:
2548:
2542:
2541:
2539:
2537:
2533:
2530:
2528:
2524:
2520:
2498:
2492:
2490:
2484:
2482:
2480:(louse flies)
2476:
2474:
2468:
2467:
2465:
2463:
2459:
2453:
2450:
2448:
2442:
2440:
2438:(flesh flies)
2434:
2432:
2431:Rhinophoridae
2429:
2427:
2424:
2422:
2416:
2414:
2408:
2406:
2403:
2401:
2395:
2394:
2392:
2390:
2386:
2380:
2374:
2372:
2366:
2364:
2358:
2356:
2352:Anthomyiidae
2350:
2349:
2347:
2345:
2341:
2338:
2336:
2332:
2322:
2320:(lance flies)
2316:
2314:
2311:
2310:
2308:
2304:
2298:
2292:
2290:
2287:
2285:
2279:
2277:
2274:
2272:
2270:(beach flies)
2266:
2264:
2258:
2256:
2253:
2251:
2248:
2247:
2245:
2243:
2239:
2233:
2227:
2225:
2223:(shore flies)
2219:
2217:
2211:
2209:
2203:
2201:
2197:Curtonotidae
2195:
2193:
2190:
2189:
2187:
2185:
2181:
2175:
2172:
2170:
2169:Teratomyzidae
2167:
2165:
2162:
2160:
2157:
2155:
2152:
2150:
2144:
2142:
2139:
2137:
2134:
2132:
2129:
2127:
2121:
2119:
2113:
2111:
2108:
2106:
2103:
2101:
2095:
2094:
2092:
2090:
2086:
2080:
2077:
2075:
2074:Cremifaniidae
2072:
2070:
2068:(aphid flies)
2064:
2062:
2056:
2055:
2053:
2051:
2047:
2041:
2035:
2033:
2030:
2028:
2025:
2023:
2020:
2019:
2017:
2015:
2011:
2005:
1999:
1997:
1995:(marsh flies)
1991:
1989:
1988:Ropalomeridae
1986:
1984:
1981:
1979:
1978:Natalimyzidae
1976:
1974:
1971:
1969:
1966:
1964:
1961:
1959:
1956:
1954:
1951:
1949:
1943:
1942:
1940:
1938:
1934:
1928:
1925:
1923:
1920:
1918:
1915:
1913:
1910:
1908:
1902:
1900:
1897:
1895:
1894:Megamerinidae
1892:
1890:
1887:
1885:
1879:
1878:
1876:
1874:
1870:
1864:
1858:
1856:
1852:Micropezidae
1850:
1848:
1845:
1844:
1842:
1840:
1836:
1830:
1824:
1822:
1816:
1814:
1811:
1809:
1806:
1804:
1798:
1796:
1790:
1788:
1782:
1781:
1779:
1777:
1773:
1767:
1761:
1760:
1758:
1756:
1752:
1749:
1747:
1743:
1740:
1738:
1734:
1730:
1716:
1710:
1708:
1704:Pipunculidae
1702:
1701:
1699:
1697:
1693:
1687:
1683:Platypezidae
1681:
1679:
1673:
1671:
1665:
1663:
1657:
1655:
1651:Ironomyiidae
1649:
1648:
1646:
1644:
1643:Platypezoidea
1640:
1637:
1635:
1631:
1627:
1624:
1622:
1618:
1600:
1594:
1592:
1586:
1585:
1583:
1581:
1577:
1571:
1568:
1566:
1565:Oreogetonidae
1563:
1561:
1559:(dance flies)
1555:
1553:
1550:
1548:
1542:
1540:
1534:
1532:
1529:
1528:
1526:
1524:
1520:
1514:
1508:
1506:
1502:Scenopinidae
1500:
1498:
1495:
1493:
1491:(mydas flies)
1487:
1485:
1479:
1477:
1474:
1472:
1466:
1464:
1458:
1456:
1455:Apystomyiidae
1453:
1451:
1448:
1446:
1440:
1439:
1437:
1435:
1431:
1428:
1426:
1422:
1418:
1414:
1410:
1406:
1400:
1396:
1374:
1368:
1366:
1360:
1358:
1352:
1350:
1344:
1343:
1341:
1336:(crane flies)
1331:
1327:
1321:
1315:
1314:
1312:
1310:
1306:
1303:
1301:
1297:
1293:
1283:
1277:
1275:
1269:
1268:
1266:
1264:
1260:
1256:
1242:
1236:
1235:
1233:
1231:
1227:
1221:
1220:Valeseguyidae
1218:
1216:
1210:
1208:
1205:
1204:
1202:
1200:
1196:
1193:
1191:
1187:
1183:
1173:
1170:
1169:
1167:
1165:
1161:
1157:
1143:
1137:
1135:
1129:
1127:
1124:
1122:
1116:
1114:
1110:Keroplatidae
1108:
1106:
1103:
1101:
1100:Diadocidiidae
1098:
1096:
1094:(gall midges)
1090:
1088:
1085:
1084:
1082:
1072:
1068:
1062:
1058:Anisopodidae
1056:
1055:
1053:
1051:
1050:Anisopodoidea
1047:
1041:
1035:
1034:
1032:
1030:
1026:
1023:
1021:
1020:Bibionomorpha
1017:
1013:
1003:
1002:Nymphomyiidae
1000:
998:
992:
990:
984:
983:
981:
979:
975:
971:
957:
953:Chironomidae
951:
949:
943:
941:
939:(black flies)
935:
933:
929:Thaumaleidae
927:
926:
924:
922:
921:Chironomoidea
918:
912:
906:
904:
898:
896:
890:
888:
882:
881:
879:
877:
873:
870:
868:
864:
860:
850:
847:
846:
844:
842:
841:Axymyiomorpha
838:
834:
830:
826:
820:
816:
810:
806:
804:
800:
798:
794:
792:
788:
786:
782:
780:
776:
775:
772:
767:
759:
754:
752:
747:
745:
740:
739:
736:
725:
721:
717:
713:
706:
703:
698:
694:
690:
686:
679:
676:
671:
667:
663:
659:
652:
649:
643:
640:
635:
628:
624:
621:(550): 1–41.
620:
616:
609:
606:
601:
597:
593:
589:
585:
581:
580:
572:
569:
564:
560:
556:
552:
545:
542:
534:
533:
525:
522:
517:
513:
509:
505:
498:
495:
490:
486:
482:
478:
474:
467:
465:
461:
455:
452:
447:
443:
436:
429:
426:
421:
417:
410:
407:
400:
393:
392:
388:
385:
384:Burmese amber
381:
380:
376:
373:
372:
368:
365:
364:
360:
357:
356:
352:
349:
348:
344:
341:
340:
336:
333:
332:
328:
325:
324:
320:
319:
317:
316:
312:
311:
310:
304:
302:
300:
295:
292:
288:
283:
281:
277:
273:
268:
264:
260:
256:
252:
248:
244:
239:
237:
232:
228:
224:
220:
216:
215:Nymphomyiidae
205:
200:
199:Nymphomyiidae
197:
194:
193:
190:
187:
184:
183:
180:
177:
174:
173:
170:
167:
164:
163:
160:
157:
154:
153:
150:
147:
144:
143:
140:
137:
134:
133:
130:
127:
124:
123:
118:
113:
109:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
35:Nymphomyiidae
33:
30:
19:
2983:expanding it
2972:
2957:
2760:
2633:Oreoleptidae
2625:Athericidae
2604:Rhagionidae
2599:Bolbomyiidae
2586:Rhagionoidea
2577:Tabanomorpha
2470:Glossinidae
2452:Ulurumyiidae
2378:(dung flies)
2318:Lonchaeidae
2306:Lonchaeoidea
2294:Milichiidae
2289:Inbiomyiidae
2283:(frit flies)
2281:Chloropidae
2205:Diastatidae
2174:Xenasteiidae
2105:Anthomyzidae
2097:Agromyzidae
2050:Lauxanioidea
2027:Heleomyzidae
1993:Sciomyzidae
1983:Phaeomyiidae
1973:Huttoninidae
1958:Helcomyzidae
1947:(kelp flies)
1937:Sciomyzoidea
1906:(rust flies)
1818:Tephritidae
1813:Richardiidae
1792:Piophilidae
1776:Tephritoidea
1746:Acalyptratae
1714:(hoverflies)
1588:Acroceridae
1468:Bombyliidae
1442:Apioceridae
1300:Tipulomorpha
1279:Tanyderidae
1240:(moth flies)
1238:Psychodidae
1230:Psychodoidea
1212:Scatopsidae
1199:Scatopsoidea
1060:(wood gnats)
1001:
910:(mosquitoes)
900:Chaoboridae
867:Culicomorpha
809:Holometabola
807:Superorder:
801:Infraclass:
715:
711:
705:
688:
684:
678:
661:
657:
651:
642:
618:
614:
608:
583:
577:
571:
554:
550:
544:
531:
524:
507:
503:
497:
480:
476:
472:
454:
445:
441:
428:
419:
415:
409:
389:
377:
369:
361:
353:
347:N. holoptica
345:
337:
329:
321:
313:
308:
299:Strashilidae
296:
290:
284:
279:
271:
266:
258:
254:
250:
242:
240:
236:Archidiptera
214:
212:
198:
188:
185:Infraorder:
29:
2875:iNaturalist
2785:Wikispecies
2710:(awl flies)
2560:Xylomyidae
2496:(bat flies)
2494:Streblidae
2488:(bat flies)
2444:Tachinidae
2335:Calyptratae
2221:Ephydridae
2207:(bog flies)
2184:Ephydroidea
2117:(sap flies)
2089:Opomyzoidea
2079:Lauxaniidae
2058:Celyphidae
2022:Chyromyidae
1953:Dryomyzidae
1945:Coelopidae
1927:Tanypezidae
1737:Schizophora
1621:Muscomorpha
1531:Atelestidae
1510:Therevidae
1470:(bee flies)
1425:Asilomorpha
1362:Pediciidae
1354:Limoniidae
1105:Ditomyiidae
1037:Bibionidae
1029:Bibionoidea
937:Simuliidae
718:: 115–120.
339:N. brundini
263:Psychodidae
3022:Categories
2975:Nematocera
2643:Tabanidae
2617:Tabanoidea
2420:(botflies)
2418:Oestridae
2389:Oestroidea
2360:Fanniidae
2268:Canacidae
2262:(bee lice)
2260:Braulidae
2192:Camillidae
2159:Opomyzidae
2136:Marginidae
2123:Clusiidae
1912:Somatiidae
1899:Nothybidae
1881:Diopsidae
1873:Diopsoidea
1826:Ulidiidae
1808:Pyrgotidae
1763:Conopidae
1755:Conopoidea
1712:Syrphidae
1696:Syrphoidea
1667:Opetiidae
1557:Hybotidae
1544:Empididae
1523:Empidoidea
1405:Brachycera
1370:Tipulidae
1330:Tipuloidea
1139:Sciaridae
1071:Sciaroidea
908:Culicidae
876:Culicoidea
849:Axymyiidae
825:Nematocera
795:Subclass:
785:Arthropoda
557:(7): 603.
510:(3): 293.
483:(2): 119.
473:Nymphomyia
422:: 559–569.
401:References
363:N. walkeri
315:Nymphomyia
291:Nymphomyia
272:Nymphomyia
267:Nymphomyia
259:Nymphomyia
179:Nematocera
175:Suborder:
149:Arthropoda
2426:Rhiniidae
2368:Muscidae
2344:Muscoidea
2242:Carnoidea
2154:Odiniidae
2141:Neminidae
2110:Asteiidae
2001:Sepsidae
1904:Psilidae
1889:Gobryidae
1860:Neriidae
1839:Nerioidea
1675:Phoridae
1570:Ragadidae
1476:Evocoidae
1460:Asilidae
1434:Asiloidea
1403:Suborder
823:Suborder
797:Pterygota
777:Kingdom:
442:Fly Times
135:Kingdom:
129:Eukaryota
2823:BugGuide
2799:BioLib:
2776:Q1936442
2770:Wikidata
2276:Carnidae
1489:Mydidae
884:Dixidae
803:Neoptera
783:Phylum:
779:Animalia
768:families
600:23426262
287:neotenic
204:Tokunaga
195:Family:
145:Phylum:
139:Animalia
125:Domain:
1634:Aschiza
791:Insecta
789:Class:
766:Diptera
764:Extant
685:Zootaxa
323:N. alba
305:Species
231:streams
169:Diptera
165:Order:
159:Insecta
155:Class:
2945:140835
2919:560809
2906:121292
2893:109702
2880:557991
2828:172895
2815:449536
2802:779144
598:
579:Nature
448:: 2–8.
227:mosses
219:family
217:are a
206:, 1932
2977:is a
2932:53947
2888:IRMNG
2841:626HL
536:(PDF)
438:(PDF)
223:flies
2979:stub
2914:NCBI
2901:ITIS
2867:5574
2862:GBIF
2854:8976
2810:BOLD
689:4763
596:PMID
213:The
46:PreꞒ
2849:EoL
2836:CoL
720:doi
693:doi
666:doi
623:doi
619:550
588:doi
584:495
559:doi
555:121
512:doi
485:doi
382:† (
3024::
2942::
2929::
2916::
2903::
2890::
2877::
2864::
2851::
2838::
2825::
2812::
2787::
2772::
716:25
714:.
687:.
662:96
660:.
617:.
594:.
582:.
553:.
508:41
506:.
481:16
479:.
463:^
446:43
444:.
440:.
420:13
418:.
282:.
96:Pg
3010:e
3003:t
2996:v
2985:.
757:e
750:t
743:v
726:.
722::
699:.
695::
672:.
668::
629:.
625::
602:.
590::
565:.
561::
518:.
514::
491:.
487::
386:)
101:N
91:K
86:J
81:T
76:P
71:C
66:D
61:S
56:O
51:Ꞓ
20:)
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