474:. The antennae are generally moniliform in structure, thus having an appearance similar to a string of beads. They are composed of 12 segments (flagellomeres), which gradually increase in diameter from the base to the antenna tip. This is the lowest number of antennal segments found in the Termitidae, with some modern Syntermitinae genera coming close, having 13 segments. The presence of three pairs of spurs on the tibia is considered unusual for the Termitinae, leading Engel and Grimaldi to leave the fossil's placement among the Termitinae tentative.
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to a light brown, and to a white strip along the labrum's apical margin. The holotype has a length of 2.0 mm (0.079 in) and sports wings that are 2.6 mm (0.10 in). Adults have a slightly oval head which is longer than wide with a sparse distribution of
352:, India. The holotype is composed of a mostly complete adult of indeterminate sex and minute overall size. Cambay amber dates to between 50 and 52 million years old, placing it in the Early to Mid
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in a transparent chunk of Cambay amber. The amber specimen, "Tad-262", is currently housed in the fossil collection of the
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423:. This is in reference to the size of the adult, which is noted as possibly the smallest termite adult known. The
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563:"Biogeographic and evolutionary implications of a diverse paleobiota in amber from the early Eocene of India"
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is the oldest member of the
Termitidae to be described to date, with the prior oldest being from the Late
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is in honor of
Charlotte Isaac, who discovered the holotype and "many other interesting inclusions".
379:, during a collecting trip on 17β22 January 2010. The fossil was first studied by paleoentomologists
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508:"The termites of Early Eocene Cambay amber, with the earliest record of the Termitidae (Isoptera)"
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which ranges from dark brown on the majority of the head and abdomen, lightening to brown on the
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Proceedings of the
National Academy of Sciences of the United States of America
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Engel, M.S.; Grimaldi, D.A.; Nascimbene, P.C.; Singh, H. (2011).
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of the new genus and species was published in the online journal
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315:, India). The genus contains a single described species,
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438:, 20 million years younger than the Cambay ambers.
364:, which is produced mainly by trees in the family
368:. The holotype specimen was recovered from the
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360:shore environment. The amber formed from a
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356:age of the Eocene, and was preserved in a
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725:8A3AF869-D486-4A22-94A5-C8F2DDC66D5D
321:placed tentatively in the subfamily
405:was coined as a combination of the
336:is known from a single fossil, the
389:American Museum of Natural History
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773:Taxa named by Michael S. Engel
561:Rust, J.; et al. (2010).
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763:Fossil taxa described in 2011
391:. Engel and Grimaldi's 2011
348:Institute of Palaeobotany in
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329:History and classification
758:Insects described in 2011
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129:Scientific classification
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16:Extinct genus of termites
768:Cenozoic insects of Asia
529:10.3897/zookeys.148.1797
419:, the type genus of the
588:10.1073/pnas.1007407107
707:Paleobiology Database
413:meaning "small" and
299:known from only one
579:2010PNAS..10718360R
573:(43): 18360β18365.
520:2011ZooK..148..105E
340:adult, which is an
631:Nanotermes isaacae
401:. The genus name
318:Nanotermes isaacae
255:Nanotermes isaacae
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694:Open Tree of Life
623:Taxon identifiers
375:mine, located in
362:dammar type resin
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514:(148): 105β123.
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425:specific epithet
393:type description
381:Michael S. Engel
366:Dipterocarpaceae
303:fossil found in
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29:Temporal range:
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192:Infraorder:
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463:postclypeus
442:Description
742:Categories
478:References
451:coloration
447:Nanotermes
432:Nanotermes
421:Termitidae
403:Nanotermes
370:Tadkeshwar
334:Nanotermes
323:Termitinae
297:Termitidae
277:Nanotermes
220:Nanotermes
206:Termitidae
166:Arthropoda
121:N. isaacae
25:Nanotermes
436:Oligocene
342:inclusion
227:Species:
186:Blattodea
152:Kingdom:
146:Eukaryota
123:holotype
673:27326377
646:Q6964160
640:Wikidata
607:20974929
548:22287892
467:antennae
455:pronotum
358:brackish
354:Ypresian
338:holotype
293:Isoptera
289:termites
266:Grimaldi
202:Family:
196:Isoptera
162:Phylum:
156:Animalia
142:Domain:
33:Ypresian
720:ZooBank
699:6291754
686:8583139
598:2972964
575:Bibcode
539:3264413
516:Bibcode
512:ZooKeys
428:isaacae
398:ZooKeys
373:lignite
350:Lucknow
313:Gujarat
307:of the
295:family
291:in the
282:extinct
252:†
231:†
212:Genus:
182:Order:
176:Insecta
172:Class:
100:↓
712:220285
605:
595:
546:
536:
461:, and
459:labrum
416:Termes
301:Eocene
280:is an
268:, 2011
264:&
660:744N6
472:setae
411:nanos
409:word
407:Greek
305:amber
285:genus
262:Engel
681:GBIF
603:PMID
544:PMID
383:and
39:Preκ
668:EoL
655:CoL
593:PMC
583:doi
571:107
534:PMC
524:doi
287:of
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510:.
486:^
457:,
325:.
89:Pg
609:.
585::
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550:.
526::
518::
311:(
216:β
94:N
84:K
79:J
74:T
69:P
64:C
59:D
54:S
49:O
44:κ
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