Knowledge (XXG)

Natal shyshark

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and greatly expanded, triangular nasal skin flaps that reach the mouth. There are a pair of deep grooves, covered by the nasal flaps, that run from the excurrent (outflow) openings of the nostrils to the mouth. The mouth has furrows at the corners on both jaws, and contains teeth with 3–5
426:. The back is brown, with a distinctive pattern of H-shaped, dark brown saddles with well-defined margins, along with numerous small white dots and darker mottling between the saddles. This species attains a length of 50 cm (20 in). 366:. As opposed to the puffadder shyshark, which inhabits deeper, cooler waters in the northeastern parts of its range, in those areas the Natal shyshark occurs in warm inshore waters at a depth of 0–30 m (0–98 ft). 438:
at around 50 cm (20 in) long, and females at 48 cm (19 in) long. The shysharks are so named because they curl into a ring and cover their eyes with their tails when threatened.
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are of roughly equal size. The dorsal fins are placed far back on the body, with the first originating behind the pelvic fin origins and the second originating behind the anal fin origins. The
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habits. Reaching 50 cm (20 in) in length, the Natal shyshark is similar to the puffadder shyshark in appearance but has a stockier body, less flattened head, a compressed
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where industrial and tourism sectors have undergone rapid expansion. Furthermore, the region is subject to heavy fishing pressure, in particular from the commercial
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The Natal shyshark closely resembles the puffadder shyshark, but has a more robust body, a less flattened head, and a laterally compressed
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Pollom, R.; Gledhill, K.; Da Silva, C.; Ebert, D.A.; Leslie, R.; McCord, M.E.; Sink, K.; Winker, H. (2019).
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The Natal shyshark is rarer than the puffadder shyshark and little is known of its habits. Males are
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has a ventral notch on the upper lobe and an indistinct lower lobe. The skin is thick, bearing well-
196: 2097: 2088: 1632: 1461: 1443: 1425: 1371: 1317: 1248: 1104: 1017: 823: 798: 764: 757: 447: 292: 30: 2229: 1980: 1926: 1893: 1815: 1578: 1470: 1407: 1389: 1059: 981: 972: 956: 938: 839: 805: 748: 462: 354:. Its total range is estimated to encompass an area of under 100 km (39 sq mi). A 324: 316: 300: 296: 256: 60: 45: 570: 2397: 2444: 2340: 2211: 2139: 1998: 1953: 1737: 1668: 1569: 1512: 1434: 1275: 990: 947: 920: 884: 875: 857: 848: 728: 606: 551: 210: 137: 2449: 2410: 2115: 2070: 2055: 1989: 1686: 1266: 1230: 1086: 866: 814: 789: 639: 510: 249: 200: 2265: 2184: 2106: 2022: 1962: 1782: 1308: 1284: 603:
Sharks of the World: An Annotated and Illustrated Catalogue of Shark Species Known to Date
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formally described this shark as a new species, in an article published in the
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The distribution of the Natal shyshark is restricted to the waters off the
653: 2353: 2312: 704: 407: 264: 252: 245: 147: 97: 2358: 2371: 458: 379: 355: 284: 241: 2384: 630:, a new catshark (Chondrichthyes: Scyliorhinidae) from South Africa". 1560: 698: 451: 388: 87: 2289: 446:
The highly restricted range of the Natal shyshark is threatened by
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This species was once considered to be the "Natal" form of the
605:. Rome: Food and Agricultural Organization. pp. 332–333. 486: 484: 482: 291:, and a different color pattern. Rare and under threat from 626:
Human, B.A. & Compagno, L.J.V. (2006). "Description of
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from coastal development, particularly in the vicinity of
2302: 2128: 2068: 2035: 2011: 1915: 1855: 1804: 1726: 1558: 1525: 1501: 1306: 1219: 1186: 1126: 1039: 737: 546:Compagno, L.J.V.; M. Dando & S. Fowler (2005). 378:. The snout is broad and rounded, with very large 255:. It was once regarded as the "Natal" form of the 550:. Princeton University Press. pp. 234–235. 541: 539: 537: 535: 533: 516:10.2305/IUCN.UK.2019-3.RLTS.T161667A124524866.en 383:points. The eyes are large, with a rudimentary 467:International Union for Conservation of Nature 395:are positioned somewhat dorsally on the body. 305:International Union for Conservation of Nature 283:to a depth of 30 m (98 ft), and has 669: 8: 279:. It is found close to the coast, from the 2290: 676: 662: 654: 596: 594: 592: 590: 414:are moderately large. The short and broad 391:) and a prominent ridge beneath. The five 209: 39: 20: 514: 571:"UCT Scientists Honour Shark Researcher" 478: 2529:Critically endangered fauna of Africa 7: 2463:EC93734C-29CE-F43C-1D03-0E1DD10FFFE1 502:IUCN Red List of Threatened Species 1363:Australian reticulate swellshark ( 1207:Australian blackspotted catshark ( 14: 2499:IUCN Red List vulnerable species 1327:Circle-blotch pygmy swellshark ( 358:species, it can be found in the 64: 584:. Retrieved on August 31, 2009. 1765:Indonesian speckled catshark ( 1: 1714:Springer's sawtail catshark ( 1537:Australian sawtail catshark ( 1156:Australian marbled catshark ( 1051:Australian spotted catshark ( 2514:Endemic fish of South Africa 1399:Leopard-spotted swellshark ( 218:Range of the Natal shyshark 2509:Marine fish of South Africa 1696:Blacktip sawtail catshark ( 1660:Broadfin sawtail catshark ( 1642:Southern sawtail catshark ( 1624:Longnose sawtail catshark ( 1588:Atlantic sawtail catshark ( 1546:Northern sawtail catshark ( 912:Black roughscale catshark ( 2550: 2524:Critically endangered fish 2304:Haploblepharus kistnasamyi 1678:African sawtail catshark ( 1615:Slender sawtail catshark ( 1597:Longfin sawtail catshark ( 1417:Sarawak pygmy swellshark ( 1069:Western spotted catshark ( 628:Haploblepharus kistnasamyi 495:Haploblepharus kistnasamyi 299:, it has been assessed as 238:Haploblepharus kistnasamyi 190:Haploblepharus kistnasamyi 1165:Eastern banded catshark ( 1096:Orange spotted catshark ( 693: 644:10.11646/zootaxa.1318.1.2 601:Compagno, L.J.V. (1984). 217: 208: 186: 179: 61:Scientific classification 59: 37: 28: 23: 2203:Cinder cloudy catshark ( 2167:Yellowspotted catshark ( 2158:Small-spotted catshark ( 2080:Narrowmouthed catshark ( 1936:White-clasper catshark ( 1705:Dwarf sawtail catshark ( 1345:Reticulated swellshark ( 1198:New Caledonia catshark ( 582:The Times (South Africa) 342:, and possibly also the 334:Distribution and habitat 2221:Whitesaddled catshark ( 2194:Brownspotted catshark ( 2176:West African catshark ( 1747:Blackspotted catshark ( 1381:Australian swellshark ( 1078:Pale spotted catshark ( 1027:Panama ghost catshark ( 964:Spatulasnout catshark ( 930:Smalldorsal cat shark ( 509:: e.T161667A124524866. 2534:Fish described in 2006 1489:Narrowbar swellshark ( 1258:New Zealand catshark ( 1147:Banded sand catshark ( 1009:South China catshark ( 894:Broadmouth cat shark ( 348:Eastern Cape Provinces 2098:Narrowtail catshark ( 2089:Redspotted catshark ( 1972:McMillan's catshark ( 1633:Blackmouth catshark ( 1462:Blotchy swell shark ( 1444:Speckled swellshark ( 1426:Flagtail swellshark ( 1372:Draughtsboard shark ( 1318:Whitefin swellshark ( 1249:Broadhead cat shark ( 1105:Variegated catshark ( 1018:Spongehead catshark ( 824:Smallbelly catshark ( 362:and on shallow rocky 321:Leonard J.V. Compagno 1981:Blackgill catshark ( 1927:White-tip catshark ( 1894:White-spotted Izak ( 1816:Puffadder shyshark ( 1579:Roughtail catshark ( 1471:Saddled swellshark ( 1408:Painted swellshark ( 1390:Spotted swellshark ( 1354:Formosa swellshark ( 982:Broadgill catshark ( 973:Deepwater catshark ( 939:Largenose catshark ( 840:Broadnose catshark ( 806:Humpback cat shark ( 749:Flatnose cat shark ( 580:(January 18, 2009). 385:nictitating membrane 267:to a small area off 2230:Blotched catshark ( 2212:Freckled catshark ( 2140:Polkadot catshark ( 1999:Filetail catshark ( 1954:Campeche catshark ( 1738:Speckled catshark ( 1669:Peppered catshark ( 1570:Antilles catshark ( 1513:Lollipop catshark ( 1435:Indian swellshark ( 1336:Cook's swellshark ( 1276:Spotless catshark ( 1060:Blotched catshark ( 991:Saldanha catshark ( 948:Smallfin catshark ( 921:Smalleye catshark ( 885:Flathead catshark ( 876:Longhead catshark ( 858:Longnose catshark ( 849:Japanese catshark ( 548:Sharks of the World 448:habitat degradation 430:Biology and ecology 293:habitat degradation 248:, belonging to the 172:H. kistnasamyi 31:Conservation status 2116:Slender catshark ( 2056:Leopard catshark ( 1990:Salamander shark ( 1687:Phallic catshark ( 1267:Bristly catshark ( 1231:Arabian catshark ( 867:Iceland catshark ( 815:Longfin catshark ( 790:Flaccid catshark ( 576:2009-09-20 at the 442:Human interactions 387:(protective third 325:scientific journal 317:puffadder shyshark 297:commercial fishing 257:puffadder shyshark 2486: 2485: 2445:Open Tree of Life 2296:Taxon identifiers 2287: 2286: 2266:Cloudy catshark ( 2185:Comoro catshark ( 2107:Lizard catshark ( 2023:Onefin catshark ( 1983:P. melanobranchus 1963:Velvet catshark ( 1929:P. albimarginatus 1783:Quagga catshark ( 1285:Sombre catshark ( 932:A. micropterygeus 842:A. investigatoris 729:Carcharhiniformes 557:978-0-691-12072-0 263:). This shark is 222: 221: 138:Carcharhiniformes 54: 2541: 2479: 2478: 2466: 2465: 2453: 2452: 2440: 2439: 2427: 2426: 2414: 2413: 2401: 2400: 2388: 2387: 2375: 2374: 2362: 2361: 2349: 2348: 2336: 2335: 2323: 2322: 2321: 2291: 2275:Dwarf catshark ( 2239:Chain catshark ( 2071:Schroederichthys 2025:P. profundicolus 1956:P. campechiensis 1945:Beige catshark ( 1867:Honeycomb Izak ( 1834:Natal shyshark ( 1825:Brown shyshark ( 1792:Rusty catshark ( 1774:Tiger catshark ( 1756:Lined catshark ( 1651:Mouse catshark ( 1606:Gecko catshark ( 1329:C. circulopullum 1240:Dusky catshark ( 1174:Coral catshark ( 1087:Dwarf catshark ( 966:A. platyrhynchus 903:Ghost catshark ( 887:A. macrorhynchus 878:A. longicephalus 781:Hoary catshark ( 772:Brown catshark ( 678: 671: 664: 655: 648: 647: 623: 617: 616: 598: 585: 568: 562: 561: 543: 528: 527: 525: 523: 518: 488: 424:dermal denticles 230:eastern shyshark 213: 192: 69: 68: 48: 43: 42: 21: 16:Species of shark 2549: 2548: 2544: 2543: 2542: 2540: 2539: 2538: 2489: 2488: 2487: 2482: 2474: 2469: 2461: 2456: 2448: 2443: 2435: 2430: 2422: 2417: 2409: 2404: 2396: 2391: 2383: 2378: 2370: 2365: 2357: 2352: 2344: 2339: 2331: 2326: 2317: 2316: 2311: 2298: 2288: 2283: 2205:S. hachijoensis 2124: 2109:S. saurisqualus 2064: 2031: 2007: 1911: 1903:Izak catshark ( 1887:H. melanostigma 1876:Grinning Izak ( 1851: 1843:Dark shyshark ( 1800: 1722: 1626:G. longirostris 1554: 1521: 1497: 1453:Balloon shark ( 1419:C. sarawakensis 1309:Cephaloscyllium 1302: 1215: 1182: 1138:Bali catshark ( 1122: 1114:Gulf catshark ( 1107:A. submaculatus 1035: 1000:Pale catshark ( 733: 689: 682: 652: 651: 625: 624: 620: 613: 600: 599: 588: 578:Wayback Machine 569: 565: 558: 545: 544: 531: 521: 519: 490: 489: 480: 475: 444: 436:sexually mature 432: 376:caudal peduncle 372: 336: 313: 289:caudal peduncle 204: 194: 188: 175: 63: 55: 44: 40: 33: 24:Natal shyshark 17: 12: 11: 5: 2547: 2545: 2537: 2536: 2531: 2526: 2521: 2516: 2511: 2506: 2504:Haploblepharus 2501: 2491: 2490: 2484: 2483: 2481: 2480: 2467: 2454: 2441: 2428: 2415: 2402: 2389: 2376: 2363: 2350: 2337: 2324: 2308: 2306: 2300: 2299: 2294: 2285: 2284: 2282: 2281: 2272: 2263: 2257:Izu catshark ( 2254: 2245: 2236: 2227: 2218: 2209: 2200: 2191: 2187:S. comoroensis 2182: 2173: 2164: 2155: 2149:Boa catshark ( 2146: 2136: 2134: 2126: 2125: 2123: 2122: 2113: 2104: 2095: 2086: 2076: 2074: 2066: 2065: 2063: 2062: 2058:P. pantherinum 2053: 2047:Pyjama shark ( 2043: 2041: 2033: 2032: 2030: 2029: 2019: 2017: 2009: 2008: 2006: 2005: 1996: 1987: 1978: 1969: 1960: 1951: 1942: 1933: 1923: 1921: 1913: 1912: 1910: 1909: 1900: 1891: 1882: 1873: 1863: 1861: 1858:Holohalaelurus 1853: 1852: 1850: 1849: 1840: 1836:H. kistnasamyi 1831: 1822: 1812: 1810: 1807:Haploblepharus 1802: 1801: 1799: 1798: 1789: 1780: 1771: 1762: 1753: 1744: 1734: 1732: 1724: 1723: 1721: 1720: 1711: 1702: 1693: 1684: 1675: 1666: 1662:G. nipponensis 1657: 1648: 1639: 1630: 1621: 1612: 1603: 1594: 1585: 1576: 1572:G. antillensis 1566: 1564: 1556: 1555: 1553: 1552: 1543: 1533: 1531: 1523: 1522: 1520: 1519: 1509: 1507: 1499: 1498: 1496: 1495: 1486: 1477: 1468: 1459: 1450: 1441: 1432: 1423: 1414: 1405: 1396: 1387: 1378: 1369: 1360: 1351: 1342: 1333: 1324: 1314: 1312: 1304: 1303: 1301: 1300: 1294:Mud catshark ( 1291: 1282: 1278:B. immaculatus 1273: 1264: 1255: 1246: 1237: 1227: 1225: 1217: 1216: 1214: 1213: 1204: 1194: 1192: 1189:Aulohalaelurus 1184: 1183: 1181: 1180: 1171: 1162: 1153: 1144: 1134: 1132: 1124: 1123: 1121: 1120: 1111: 1102: 1098:A. rubiginosus 1093: 1084: 1075: 1066: 1057: 1047: 1045: 1037: 1036: 1034: 1033: 1024: 1015: 1006: 997: 988: 979: 975:A. profundorum 970: 961: 954: 950:A. parvipinnis 945: 936: 927: 918: 914:A. melanoasper 909: 900: 896:A. macrostomus 891: 882: 873: 869:A. laurussonii 864: 855: 846: 837: 830: 821: 812: 803: 796: 787: 778: 769: 762: 755: 745: 743: 735: 734: 732: 731: 725: 719: 717:Elasmobranchii 713: 711:Chondrichthyes 707: 701: 694: 691: 690: 683: 681: 680: 673: 666: 658: 650: 649: 618: 611: 586: 563: 556: 529: 477: 476: 474: 471: 443: 440: 431: 428: 371: 368: 335: 332: 312: 309: 253:Scyliorhinidae 226:Natal shyshark 220: 219: 215: 214: 206: 205: 195: 184: 183: 177: 176: 169: 167: 163: 162: 159:Haploblepharus 155: 151: 150: 148:Scyliorhinidae 145: 141: 140: 135: 131: 130: 125: 121: 120: 118:Elasmobranchii 115: 111: 110: 108:Chondrichthyes 105: 101: 100: 95: 91: 90: 85: 81: 80: 75: 71: 70: 57: 56: 38: 35: 34: 29: 26: 25: 15: 13: 10: 9: 6: 4: 3: 2: 2546: 2535: 2532: 2530: 2527: 2525: 2522: 2520: 2519:KwaZulu-Natal 2517: 2515: 2512: 2510: 2507: 2505: 2502: 2500: 2497: 2496: 2494: 2477: 2472: 2468: 2464: 2459: 2455: 2451: 2446: 2442: 2438: 2433: 2429: 2425: 2420: 2416: 2412: 2407: 2403: 2399: 2394: 2390: 2386: 2381: 2377: 2373: 2368: 2364: 2360: 2355: 2351: 2347: 2342: 2338: 2334: 2329: 2325: 2320: 2314: 2310: 2309: 2307: 2305: 2301: 2297: 2292: 2280: 2278: 2273: 2271: 2269: 2264: 2262: 2260: 2255: 2253: 2251: 2246: 2244: 2242: 2237: 2235: 2233: 2228: 2226: 2224: 2219: 2217: 2215: 2210: 2208: 2206: 2201: 2199: 2197: 2192: 2190: 2188: 2183: 2181: 2179: 2174: 2172: 2170: 2165: 2163: 2161: 2156: 2154: 2152: 2147: 2145: 2143: 2138: 2137: 2135: 2133: 2132: 2127: 2121: 2119: 2114: 2112: 2110: 2105: 2103: 2101: 2096: 2094: 2092: 2087: 2085: 2083: 2078: 2077: 2075: 2073: 2072: 2067: 2061: 2059: 2054: 2052: 2050: 2045: 2044: 2042: 2040: 2039: 2034: 2028: 2026: 2021: 2020: 2018: 2016: 2015: 2010: 2004: 2002: 1997: 1995: 1993: 1988: 1986: 1984: 1979: 1977: 1975: 1974:P. macmillani 1970: 1968: 1966: 1961: 1959: 1957: 1952: 1950: 1948: 1943: 1941: 1939: 1934: 1932: 1930: 1925: 1924: 1922: 1920: 1919: 1914: 1908: 1906: 1901: 1899: 1897: 1892: 1890: 1888: 1885:Crying Izak ( 1883: 1881: 1879: 1874: 1872: 1870: 1865: 1864: 1862: 1860: 1859: 1854: 1848: 1846: 1841: 1839: 1837: 1832: 1830: 1828: 1823: 1821: 1819: 1814: 1813: 1811: 1809: 1808: 1803: 1797: 1795: 1790: 1788: 1786: 1781: 1779: 1777: 1776:H. natalensis 1772: 1770: 1768: 1763: 1761: 1759: 1754: 1752: 1750: 1745: 1743: 1741: 1736: 1735: 1733: 1731: 1730: 1725: 1719: 1717: 1712: 1710: 1708: 1703: 1701: 1699: 1694: 1692: 1690: 1685: 1683: 1681: 1676: 1674: 1672: 1667: 1665: 1663: 1658: 1656: 1654: 1649: 1647: 1645: 1644:G. mincaronei 1640: 1638: 1636: 1635:G. melastomus 1631: 1629: 1627: 1622: 1620: 1618: 1613: 1611: 1609: 1604: 1602: 1600: 1595: 1593: 1591: 1590:G. atlanticus 1586: 1584: 1582: 1577: 1575: 1573: 1568: 1567: 1565: 1563: 1562: 1557: 1551: 1549: 1544: 1542: 1540: 1535: 1534: 1532: 1530: 1529: 1524: 1518: 1516: 1511: 1510: 1508: 1506: 1505: 1500: 1494: 1492: 1487: 1485: 1483: 1482:C. ventriosum 1478: 1476: 1474: 1473:C. variegatum 1469: 1467: 1465: 1460: 1458: 1456: 1451: 1449: 1447: 1442: 1440: 1438: 1433: 1431: 1429: 1424: 1422: 1420: 1415: 1413: 1411: 1406: 1404: 1402: 1401:C. pardelotum 1397: 1395: 1393: 1388: 1386: 1384: 1379: 1377: 1375: 1370: 1368: 1366: 1365:C. hicosellum 1361: 1359: 1357: 1356:C. formosanum 1352: 1350: 1348: 1343: 1341: 1339: 1334: 1332: 1330: 1325: 1323: 1321: 1320:C. albipinnum 1316: 1315: 1313: 1311: 1310: 1305: 1299: 1297: 1292: 1290: 1288: 1283: 1281: 1279: 1274: 1272: 1270: 1265: 1263: 1261: 1256: 1254: 1252: 1247: 1245: 1243: 1238: 1236: 1234: 1229: 1228: 1226: 1224: 1223: 1218: 1212: 1210: 1205: 1203: 1201: 1196: 1195: 1193: 1191: 1190: 1185: 1179: 1177: 1176:A. marmoratus 1172: 1170: 1168: 1163: 1161: 1159: 1154: 1152: 1150: 1145: 1143: 1141: 1136: 1135: 1133: 1131: 1130: 1129:Atelomycterus 1125: 1119: 1117: 1112: 1110: 1108: 1103: 1101: 1099: 1094: 1092: 1090: 1085: 1083: 1081: 1076: 1074: 1072: 1067: 1065: 1063: 1058: 1056: 1054: 1049: 1048: 1046: 1044: 1043: 1038: 1032: 1030: 1025: 1023: 1021: 1020:A. spongiceps 1016: 1014: 1012: 1007: 1005: 1003: 998: 996: 994: 989: 987: 985: 980: 978: 976: 971: 969: 967: 962: 960: 959: 955: 953: 951: 946: 944: 942: 937: 935: 933: 928: 926: 924: 919: 917: 915: 910: 908: 906: 901: 899: 897: 892: 890: 888: 883: 881: 879: 874: 872: 870: 865: 863: 861: 856: 854: 852: 847: 845: 843: 838: 836: 835: 834:A. internatus 831: 829: 827: 822: 820: 818: 813: 811: 809: 804: 802: 801: 797: 795: 793: 788: 786: 784: 779: 777: 775: 770: 768: 767: 763: 761: 760: 756: 754: 752: 747: 746: 744: 742: 741: 736: 730: 726: 724: 720: 718: 714: 712: 708: 706: 702: 700: 696: 695: 692: 687: 679: 674: 672: 667: 665: 660: 659: 656: 645: 641: 637: 633: 629: 622: 619: 614: 612:92-5-101384-5 608: 604: 597: 595: 593: 591: 587: 583: 579: 575: 572: 567: 564: 559: 553: 549: 542: 540: 538: 536: 534: 530: 517: 512: 508: 504: 503: 498: 496: 487: 485: 483: 479: 472: 470: 468: 464: 460: 457: 453: 449: 441: 439: 437: 429: 427: 425: 421: 417: 413: 412:pectoral fins 409: 405: 401: 396: 394: 390: 386: 381: 377: 369: 367: 365: 361: 357: 353: 349: 345: 341: 340:KwaZulu-Natal 333: 331: 329: 326: 322: 318: 310: 308: 306: 302: 298: 294: 290: 286: 282: 278: 277:KwaZulu-Natal 274: 270: 266: 262: 258: 254: 251: 247: 243: 239: 235: 234:happy chappie 231: 227: 216: 212: 207: 202: 198: 193: 191: 185: 182: 181:Binomial name 178: 174: 173: 168: 165: 164: 161: 160: 156: 153: 152: 149: 146: 143: 142: 139: 136: 133: 132: 129: 128:Selachimorpha 126: 124:Subdivision: 123: 122: 119: 116: 113: 112: 109: 106: 103: 102: 99: 96: 93: 92: 89: 86: 83: 82: 79: 76: 73: 72: 67: 62: 58: 52: 47: 36: 32: 27: 22: 19: 2303: 2276: 2267: 2258: 2250:S. stellaris 2249: 2248:Nursehound ( 2240: 2231: 2223:S. hesperius 2222: 2214:S. haeckelii 2213: 2204: 2195: 2186: 2178:S. cervigoni 2177: 2168: 2159: 2150: 2141: 2131:Scyliorhinus 2129: 2117: 2108: 2100:S. maculatus 2099: 2091:S. chilensis 2090: 2081: 2069: 2057: 2049:P. africanum 2048: 2036: 2024: 2012: 2000: 1991: 1982: 1973: 1964: 1955: 1946: 1938:P. albipenis 1937: 1928: 1916: 1904: 1896:H. punctatus 1895: 1886: 1877: 1868: 1856: 1844: 1835: 1833: 1826: 1818:H. edwardsii 1817: 1805: 1793: 1784: 1775: 1767:H. maculosus 1766: 1757: 1748: 1740:H. boesemani 1739: 1727: 1716:G. springeri 1715: 1706: 1697: 1688: 1679: 1671:G. piperatus 1670: 1661: 1652: 1643: 1634: 1625: 1616: 1607: 1598: 1589: 1580: 1571: 1559: 1547: 1539:F. boardmani 1538: 1526: 1514: 1502: 1490: 1481: 1480:Swellshark ( 1472: 1464:C. umbratile 1463: 1454: 1445: 1436: 1428:C. signourum 1427: 1418: 1409: 1400: 1392:C. maculatum 1391: 1382: 1374:C. isabellum 1373: 1364: 1355: 1347:C. fasciatum 1346: 1337: 1328: 1319: 1307: 1295: 1286: 1277: 1268: 1259: 1250: 1242:B. canescens 1241: 1232: 1220: 1208: 1200:A. kanakorum 1199: 1187: 1175: 1167:A. marnkalha 1166: 1157: 1149:A. fasciatus 1148: 1140:A. baliensis 1139: 1127: 1115: 1106: 1097: 1088: 1079: 1070: 1061: 1052: 1040: 1028: 1019: 1010: 1001: 992: 983: 974: 965: 957: 949: 940: 931: 922: 913: 904: 895: 886: 877: 868: 859: 851:A. japonicus 850: 841: 833: 825: 817:A. herklotsi 816: 807: 799: 792:A. exsanguis 791: 782: 773: 765: 758: 750: 738: 721:Subdivision 635: 631: 627: 621: 602: 581: 566: 547: 520:. Retrieved 506: 500: 494: 445: 433: 422:leaf-shaped 397: 373: 352:South Africa 344:Western Cape 337: 327: 314: 273:Western Cape 269:South Africa 261:H. edwardsii 260: 237: 233: 229: 225: 223: 189: 187: 171: 170: 158: 18: 2380:iNaturalist 2268:S. torazame 2169:S. capensis 2160:S. canicula 2142:S. besnardi 2001:P. xaniurus 1878:H. grennian 1758:H. lineatus 1749:H. buergeri 1707:G. schultzi 1617:G. gracilis 1608:G. eastmani 1599:G. cadenati 1548:F. striatus 1515:C. cephalus 1455:C. sufflans 1383:C. laticeps 1296:B. lutarius 1269:B. hispidus 1233:B. alcockii 1222:Bythaelurus 1209:A. labiosus 1158:A. macleayi 1116:A. vincenti 1080:A. pallidus 1071:A. occiduus 1062:A. funebris 1029:A. stenseni 1011:A. sinensis 993:A. saldanha 808:A. gibbosus 800:A. fedorovi 774:A. brunneus 766:A. aphyodes 759:A. albisoma 751:A. acanutus 522:19 November 370:Description 2493:Categories 2259:S. tokubee 2241:S. retifer 2196:S. garmani 2014:Pentanchus 1992:P. pilosus 1965:P. lanatus 1918:Parmaturus 1729:Halaelurus 1698:G. sauteri 1689:G. priapus 1653:G. murinus 1504:Cephalurus 1446:C. speccum 1287:B. incanus 1260:B. dawsoni 1002:A. sibogae 958:A. pinguis 941:A. nasutus 923:A. microps 826:A. indicus 783:A. canutus 740:Apristurus 473:References 463:Vulnerable 416:caudal fin 393:gill slits 301:Vulnerable 114:Subclass: 46:Vulnerable 2277:S. torrei 2118:S. tenuis 2082:S. bivius 2038:Poroderma 1905:H. regani 1845:H. pictus 1827:H. fuscus 1794:H. sellus 1785:H. quagga 1491:C. zebrum 1437:C. silasi 1410:C. pictum 1251:B. clevai 1089:A. parvus 1053:A. analis 1042:Asymbolus 984:A. riveri 860:A. kampae 715:Subclass 638:: 41–58. 420:calcified 408:anal fins 360:surf zone 281:surf zone 271:from the 166:Species: 84:Kingdom: 78:Eukaryota 2398:11682396 2354:FishBase 2319:Q1570451 2313:Wikidata 2232:S. meadi 1947:P. bigus 1869:H. favus 1680:G. polli 1338:C. cooki 905:A. manis 723:Selachii 705:Chordata 699:Animalia 697:Kingdom 686:catshark 574:Archived 469:(IUCN). 380:nostrils 311:Taxonomy 307:(IUCN). 246:catshark 201:Compagno 144:Family: 98:Chordata 94:Phylum: 88:Animalia 74:Domain: 51:IUCN 3.1 2424:1003958 2372:2418606 1581:G. arae 703:Phylum 688:species 684:Extant 632:Zootaxa 465:by the 459:fishery 356:benthic 328:Zootaxa 303:by the 285:benthic 265:endemic 242:species 240:) is a 154:Genus: 134:Order: 104:Class: 49: ( 2476:295747 2450:104315 2437:295747 2411:161667 2385:102103 2333:275055 2151:S. boa 1561:Galeus 1528:Figaro 727:Order 709:Class 609:  554:  452:Durban 406:, and 404:pelvic 400:dorsal 389:eyelid 250:family 203:, 2006 199:& 2471:WoRMS 2458:Plazi 2393:IRMNG 2359:63795 2346:3JL88 456:prawn 364:reefs 197:Human 2432:OBIS 2419:NCBI 2406:IUCN 2367:GBIF 2328:BOLD 636:1318 607:ISBN 552:ISBN 524:2021 507:2019 398:The 346:and 295:and 224:The 2341:CoL 640:doi 511:doi 350:of 275:to 244:of 232:or 2495:: 2473:: 2460:: 2447:: 2434:: 2421:: 2408:: 2395:: 2382:: 2369:: 2356:: 2343:: 2330:: 2315:: 634:. 589:^ 532:^ 505:. 499:. 481:^ 402:, 228:, 2279:) 2270:) 2261:) 2252:) 2243:) 2234:) 2225:) 2216:) 2207:) 2198:) 2189:) 2180:) 2171:) 2162:) 2153:) 2144:) 2120:) 2111:) 2102:) 2093:) 2084:) 2060:) 2051:) 2027:) 2003:) 1994:) 1985:) 1976:) 1967:) 1958:) 1949:) 1940:) 1931:) 1907:) 1898:) 1889:) 1880:) 1871:) 1847:) 1838:) 1829:) 1820:) 1796:) 1787:) 1778:) 1769:) 1760:) 1751:) 1742:) 1718:) 1709:) 1700:) 1691:) 1682:) 1673:) 1664:) 1655:) 1646:) 1637:) 1628:) 1619:) 1610:) 1601:) 1592:) 1583:) 1574:) 1550:) 1541:) 1517:) 1493:) 1484:) 1475:) 1466:) 1457:) 1448:) 1439:) 1430:) 1421:) 1412:) 1403:) 1394:) 1385:) 1376:) 1367:) 1358:) 1349:) 1340:) 1331:) 1322:) 1298:) 1289:) 1280:) 1271:) 1262:) 1253:) 1244:) 1235:) 1211:) 1202:) 1178:) 1169:) 1160:) 1151:) 1142:) 1118:) 1109:) 1100:) 1091:) 1082:) 1073:) 1064:) 1055:) 1031:) 1022:) 1013:) 1004:) 995:) 986:) 977:) 968:) 952:) 943:) 934:) 925:) 916:) 907:) 898:) 889:) 880:) 871:) 862:) 853:) 844:) 828:) 819:) 810:) 794:) 785:) 776:) 753:) 677:e 670:t 663:v 646:. 642:: 615:. 560:. 526:. 513:: 497:" 493:" 259:( 236:( 53:)

Index

Conservation status
Vulnerable
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Chondrichthyes
Elasmobranchii
Selachimorpha
Carcharhiniformes
Scyliorhinidae
Haploblepharus
Binomial name
Human
Compagno

species
catshark
family
Scyliorhinidae
puffadder shyshark
endemic
South Africa
Western Cape
KwaZulu-Natal
surf zone
benthic
caudal peduncle

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