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Nemonychidae

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64: 42: 260:, and the maxillary palps are long and projecting. Nemonychidae have all ventrites free, while Anthribidae have ventrites 1-4 connate or partially fused. Nemonychidae lack lateral carinae on the pronotum, while these are usually present, though may be short, in Anthribidae. 1214:
Shin S, Clarke DJ, Lemmon AR, Moriarty Lemmon E, Aitken AL, Haddad S, Farrell BD, Marvaldi AE, Oberprieler RG, McKenna DD (2018) Phylogenomic data yield new and robust insights into the phylogeny and evolution of weevils. Mol. Biol. Evol. 35: 823-836.
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Both Nemonychidae adults and larvae feed primarily on pollen. When mature, the larvae will drop to the ground from the male cones or flowers they resided in to pupate in the soil less than five centimeters from the surface.
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ICZN (2005) Opinion 2111 (Case 3093). Nemonychidae Bedel, November 1882 (Insecta, Coleoptera): given precedence over Cimberididae Gozis, March 1882; and
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Riedel, A. (2010). A new tribe, genus and species of Nemonychidae from Baltic amber (Coleoptera: Curculionoidea: Nemonychidae: Cimberidinae).
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Anderson, R.S, Oberprieler, R.G., Marvaldi, A.E. 2014. 3.1 Nemonychidae Bedel, 1882. In: Leschen R.A.B. & Beutel, R.G. (Eds)
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Nemonychidae are historically divided into three subfamilies: Nemonychinae of the palearctic region with the single genus
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Kuschel, G. & Leschen, R. A. (2011). Phylogeny and taxonomy of the Rhinorhynchinae (Coleoptera: Nemonychidae).
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There exists a fairly extensive fossil record of Nemonychidae reaching from the upper Jurassic to tertiary
3279: 1085: 3367: 816: 656: 3589: 3541: 1080: 1015: 1010: 777: 670: 647: 577: 492: 901: 843: 598: 1050: 990: 958: 906: 582: 499: 3520: 3401: 3326: 2516: 1137: 1093: 553: 548: 379: 286: 197: 1026: 772: 626: 568: 485: 2735: 1363: 985: 976: 888: 871: 755: 746: 705: 683: 614: 531: 415: 1146: 1001: 733: 692: 364: 321: 297:. Recent phylogenetic research indicates that the Cimberidinae are sister to all of the remaining 282: 192: 186: 1316: 1064: 1031: 953: 928: 714: 665: 436: 408: 394: 206: 58: 1112: 914: 513: 506: 457: 401: 387: 357: 3546: 3317: 857: 471: 464: 450: 443: 3528: 3388: 3004: 2421: 1308: 1269: 1251: 1228:
McKenna, Duane D.; Sequeira, Andrea S.; Marvaldi, Adriana E.; Farrell, Brian D. (2009-04-13).
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Handbook of Zoology, Coleoptera, Beetles Volume 3: Morphology and Systematics (Phytophaga)
697: 3427: 3414: 3193: 3144: 2999: 2806: 2474: 2458: 2397: 2292: 2263: 2191: 2155: 2139: 2055: 2000: 1905: 1802: 1651: 1613: 1593: 1489: 1264: 1229: 738: 298: 3573: 3331: 3175: 2979: 2970: 2711: 2682: 2674: 2666: 2640: 2615: 2511: 2498: 2437: 2355: 2258: 2253: 2134: 2068: 2039: 2023: 1987: 1957: 1910: 1864: 1831: 1747: 1739: 1710: 1548: 1117: 478: 343: 294: 290: 278: 3344: 1320: 3393: 3152: 3076: 2822: 2814: 2769: 2753: 2703: 2624: 2580: 2532: 2450: 2232: 2219: 2199: 2160: 2060: 2047: 2018: 1952: 1940: 1836: 1793: 1776: 1768: 1718: 1556: 1526: 1457: 1446: 1430: 135: 1230:"Temporal lags and overlap in the diversification of weevils and flowering plants" 1289:"Past and present of the relict family nemonychidae (Coleoptera, Curculionoidea)" 1179:
Gozis, 1881: usage conserved. Bulletin of Zoological Nomenclature 62(2): 101–103.
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Gratshev and Zherikhin 1995 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Gratshev and Zherikhin 1995 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Gratshev and Zherikhin 1995 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Gratshev and Zherikhin 2000 La Pedrera de Rúbies Formation, Spain, Barremian
639: 635: 586: 557: 329: 125: 95: 75: 1273: 3354: 3375: 3296: 3134: 3084: 3028: 2987: 2936: 2593: 2524: 2405: 2117: 1923: 1897: 1755: 1588: 1535: 1514: 1424: 760: 336: 32: 1288: 1216: 3380: 2851: 2843: 2695: 2429: 2350: 2334: 1304: 274: 233: 3458: 2442: 2302: 1935: 1412: 1400: 1387: 781: 630:
Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian,
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Gratshev and Legalov 2009 Sharin-Gol Formation, Mongolia, Barremian
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Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Martynov 1926 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Legalov 2003 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Legalov 2010 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
838:
Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
820:
Legalov 2009 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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group because they have straight rather than geniculate (elbowed)
301:, and it has been proposed to elevate the group to family rank. 273:. Most species of the other two subfamilies are associated with 3277: 1686: 1637: 1511: 1443: 1359: 1141:
Clarke and Oberprieler 2018 Burmese amber, Myanmar, Cenomanian
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occur largely in the Southern hemisphere, especially found on
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Legalov 2003 La Pedrera de Rúbies Formation, Spain, Barremian
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Zherikhin and Gratshev 2004 Crato Formation, Brazil, Aptian
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Scudder 1876 Green River Formation, United States, Eocene
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Scudder 1878 Green River Formation, United States, Eocene
1758:(furniture beetles, death watch beetles, spider beetles) 750:
Gratshev and Legalov 2009 Zaza Formation, Russia, Aptian
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Davis and Engel 2014 Burmese amber, Myanmar, Cenomanian
725:†subfamily Cretonemonychinae Gratshev and Legalov 2009 527: 3286: 3192: 3143: 3109: 3100: 2978: 2969: 2922: 2897: 2781: 2752: 2649: 2623: 2614: 2396: 2375: 2301: 2108: 1999: 1855: 1801: 1792: 1767: 1709: 1700: 1525: 1075:†tribe Selengarhynchini Gratshev and Legalov 2009 728:†tribe Cretonemonychini Gratshev and Legalov 2009 2635:(jewel beetles, or metallic wood-boring beetles) 1234:Proceedings of the National Academy of Sciences 1005:Legalov 2009 Emanra Formation, Russia, Turonian 2358:(true weevils, bark beetles, ambrosia beetles) 1371: 1351:Images of Nemonychidae species in New Zealand 1150:Legalov 2009 Yixian Formation, China, Aptian 8: 2601:(ironclad beetles, cylindrical bark beetles) 980:Legalov 2009 Crato Formation, Brazil, Aptian 285:are found in the Northern hemisphere, while 3274: 3106: 2975: 2620: 1798: 1706: 1697: 1683: 1634: 1522: 1508: 1440: 1378: 1364: 1356: 593:†subtribe Brenthorrhinoidina Legalov 2003 40: 20: 3265:List of subgroups of the order Coleoptera 2345:(straight snout weevils, New York weevil) 1263: 1245: 1045:†subtribe Nebrenthorrhinina Legalov 2007 537:†subfamily Brenthorrhininae Arnoldi 1977 2772:(cicada beetle, cicada parasite beetles) 618:Legalov 2010 Khasurty locality, Russia, 1168: 767:†tribe Eocaenonemonychini Legalov 2013 543:†subtribe Brenthorrhinina Arnoldi 1977 2862:(false fireflies, long-lipped beetles) 968:†subfamily Paleocartinae Legalov 2003 1217:https://doi.org/10.1093/molbev/msx324 256:appears as a separate segment to the 7: 3521:9f96fec3-e2bc-44b9-bb9a-8393cea68708 3433:e5a66d51-3437-4b8c-acb8-d0088b94b944 2888:(false metallic wood-boring beetles) 1104:Subfamily Rhinorhynchinae Voss 1922 1059:†subtribe Paleocartina Legalov 2003 971:†tribe Metrioxenoidini Legalov 2009 759:Legalov 2014 Kzyl-Zhar, Kazakhstan, 609:†tribe Distenorrhinini Arnoldi 1977 540:†tribe Brenthorrhinini Arnoldi 1977 267:and an unusual host, the angiosperm 866:†tribe Oxycorynoidini Arnoldi 1977 678:†tribe Eccoptarthrini Arnoldi 1977 1335:Insect Systematics & Evolution 852:†tribe Karataucarini Legalov 2009 277:feeding on the pollen of the male 14: 1192:. DeGruyter, Berlin, pp. 301–398. 1042:†tribe Paleocartini Legalov 2003 669:Oberprieler and Oberprieler 2012 642:Khasurty locality, Russia, Aptian 2499:Perimylopidae, or Promecheilidae 2084:(minute brown scavenger beetles) 1107:Tribe Mecomacerini Kuschel 1994 62: 2714:(variegated mud-loving beetles) 923:†tribe Probelini Legalov 2009 632:La Pedrera de Rúbies Formation 1: 2543:(narrow-waisted bark beetles) 2235:(parasitic flat bark beetles) 2186:(short-winged flower beetles) 2042:(lady beetles, or God's cows) 1779:(tooth-necked fungus beetles) 811:†tribe Eobelini Arnoldi 1977 3055:(sand-loving scarab beetles) 2445:(minute tree-fungus beetles) 2271:(silvanid flat bark beetles) 879:, United Kingdom, Barremian 3204:(primitive carrion beetles) 3160:(minute mud-loving beetles) 3023:(bumble bee scarab beetles) 2793:(soft-bodied plant beetles) 2764:(soft bodied plant beetles) 1567:(predaceous diving beetles) 3606: 3039:(scavenger scarab beetles) 3031:(enigmatic scarab beetles) 2204:Lamingtonium binnaberrense 1918:(soft-wing flower beetles) 1014:Gratshev and Legalov 2009 737:Gratshev and Legalov 2009 581:Gratshev and Legalov 2009 3262: 3178:(water scavenger beetles) 2911:Rhinorhipus tamborinensis 2690:(long-toed water beetles) 2583:(polypore fungus beetles) 2485:(palm and flower beetles) 2469:(tumbling flower beetles) 2416:(ant-like flower beetles) 2194:(lined flat bark beetles) 2168:(pleasing fungus beetles) 2063:(handsome fungus beetles) 1696: 1682: 1647: 1633: 1608:(burrowing water beetles) 1521: 1507: 1453: 1439: 1395: 212: 205: 183: 178: 59:Scientific classification 57: 48: 39: 23: 3225:(feather-winged beetles) 2559:(false longhorn beetles) 2506:(jugular-horned beetles) 2453:(false darkling beetles) 2424:(cryptic fungus beetles) 2243:(shining flower beetles) 1583:(crawling water beetles) 1482:Sikhotealinia zhiltzovae 1203:Invertebrate Systematics 244:. They are often called 51:Doydirhynchus austriacus 2706:(forest stream beetles) 2567:(synchroa bark beetles) 2527:(dead log bark beetles) 2519:(fire-coloured beetles) 2408:(ant-like leaf beetles) 2142:(silken fungus beetles) 2076:(well polished beetles) 2050:(minute fungus beetles) 1869:Acanthocnemus nigricans 1247:10.1073/pnas.0810618106 3217:(round fungus beetles) 2551:(false flower beetles) 2535:(wedge-shaped beetles) 2493:(false blister beetle) 2477:(hairy fungus beetles) 2432:(conifer bark beetles) 2329:(leaf-rolling weevils) 1990:(bark-gnawing beetles) 1750:(wounded-tree beetles) 1616:(false ground beetles) 1538:(trout-stream beetles) 788:Eugnamptus" decemsatus 673:, Australia, Tithonian 3542:Paleobiology Database 3129:(false clown beetles) 2951:(plate-thigh beetles) 2833:(false click beetles) 2738:(water-penny beetles) 2387:(ship-timber beetles) 2366:(pine flower weevils) 2214:(root-eating beetles) 2034:(minute bark beetles) 1945:Phloiophilus edwardsi 1472:(reticulated beetles) 1088:, Mongolia, Barremian 1030:Gratshev et al. 1998 1016:Tsagaantsav Formation 881:Gurvan-Eren Formation 778:Green River Formation 232:is a small family of 3428:Fauna Europaea (new) 3007:(false stag beetles) 2992:Belohina inexpectata 2854:(net-winged beetles) 2817:(rare click beetles) 2730:(travertine beetles) 2722:(minute mud beetles) 2287:(dry-fungus beetles) 2122:Agapytho foveicollis 1879:(false skin beetles) 1742:(Jacobson's beetles) 1462:Crowsoniella relicta 1086:Sharin-Gol Formation 671:Talbragar Fossil bed 554:Karabastau Formation 236:, placed within the 3005:Diphyllostomatidae 2337:(primitive weevils) 2150:(flat bark beetles) 1900:(checkered beetles) 1887:(fruitworm beetles) 1575:(whirligig beetles) 1287:Kusche, G. (1983). 817:Archaeorrhynchoides 657:Microbrenthorrhinus 246:pine flower weevils 3585:Polyphaga families 2870:(glowworm beetles) 2575:(darkling beetles) 2422:Archeocrypticidae 2279:(palmetto beetles) 2026:(dry bark beetles) 1813:(longhorn beetles) 1305:10.1007/bf00218522 1081:Selengarhynchoides 1034:, United Kingdom, 1032:Lulworth Formation 1011:Megametrioxenoides 909:, Russia, Turonian 883:, Mongolia, Aptian 795:Sitona" grandaevus 648:Megabrenthorrhinus 578:Gobibrenthorrhinus 493:Rhynchitomacerinus 3567: 3566: 3529:Open Tree of Life 3280:Taxon identifiers 3271: 3270: 3258: 3257: 3254: 3253: 3250: 3249: 3242: 3234: 3233:(carrion beetles) 3226: 3218: 3205: 3179: 3161: 3130: 3122: 3096: 3095: 3088: 3080: 3072: 3064: 3056: 3048: 3040: 3032: 3024: 3016: 3008: 2995: 2965: 2964: 2952: 2944: 2914: 2889: 2871: 2863: 2855: 2847: 2834: 2826: 2818: 2810: 2809:(soldier beetles) 2802: 2799:Brachypsectridae 2794: 2773: 2765: 2739: 2731: 2723: 2715: 2707: 2699: 2691: 2678: 2670: 2662: 2636: 2610: 2609: 2602: 2584: 2576: 2568: 2560: 2557:Stenotrachelidae 2552: 2544: 2536: 2528: 2520: 2507: 2494: 2486: 2478: 2470: 2462: 2461:(blister beetles) 2454: 2446: 2433: 2425: 2417: 2409: 2388: 2367: 2359: 2346: 2338: 2330: 2322: 2312: 2288: 2280: 2272: 2244: 2236: 2228: 2215: 2207: 2195: 2187: 2169: 2151: 2143: 2125: 2085: 2077: 2064: 2051: 2043: 2035: 2027: 2009:Akalyptoischiidae 1991: 1948: 1931: 1919: 1901: 1888: 1880: 1872: 1822: 1814: 1788: 1787: 1780: 1759: 1751: 1743: 1730: 1722: 1678: 1677: 1655: 1629: 1628: 1625: 1624: 1617: 1609: 1601: 1584: 1576: 1568: 1560: 1552: 1539: 1503: 1502: 1485: 1473: 1465: 1240:(17): 7083–7088. 1205:, 24(6), 573-615. 902:Khetoxycorynoides 844:Martynovirhynchus 780:, United States, 599:Brenthorrhinoides 227: 226: 222: 217: 174: 16:Family of beetles 3597: 3560: 3559: 3550: 3549: 3537: 3536: 3524: 3523: 3514: 3513: 3501: 3500: 3498:NHMSYS0020152782 3488: 3487: 3475: 3474: 3462: 3461: 3449: 3448: 3436: 3435: 3423: 3422: 3410: 3409: 3397: 3396: 3384: 3383: 3371: 3370: 3358: 3357: 3348: 3347: 3335: 3334: 3322: 3321: 3320: 3307: 3306: 3305: 3275: 3240: 3232: 3224: 3216: 3203: 3177: 3159: 3128: 3120: 3107: 3102:Staphyliniformia 3086: 3078: 3070: 3062: 3054: 3046: 3038: 3030: 3022: 3014: 3006: 2989: 2976: 2950: 2941:Declinia relicta 2938: 2908: 2887: 2881:Sinopyrophoridae 2876:Rhagophthalmidae 2869: 2861: 2853: 2845: 2832: 2824: 2816: 2808: 2800: 2792: 2791:Artematopodidae 2771: 2763: 2737: 2729: 2721: 2713: 2705: 2698:(riffle beetles) 2697: 2689: 2677:(turtle beetles) 2676: 2668: 2660: 2634: 2621: 2600: 2582: 2574: 2566: 2558: 2550: 2542: 2534: 2526: 2518: 2505: 2492: 2484: 2476: 2468: 2460: 2452: 2444: 2431: 2423: 2415: 2407: 2386: 2365: 2357: 2344: 2336: 2328: 2321:(fungus weevils) 2320: 2311: 2310: 2306: 2293:Tasmosalpingidae 2286: 2278: 2270: 2242: 2234: 2226: 2213: 2201: 2193: 2185: 2167: 2149: 2141: 2119: 2083: 2075: 2062: 2049: 2041: 2033: 2025: 1989: 1942: 1925: 1917: 1899: 1886: 1878: 1866: 1865:Acanthocnemidae 1820: 1812: 1799: 1778: 1757: 1749: 1741: 1728: 1720: 1707: 1698: 1684: 1653: 1635: 1615: 1607: 1595: 1582: 1574: 1566: 1558: 1551:(ground beetles) 1550: 1537: 1523: 1509: 1479: 1471: 1459: 1458:Crowsoniellidae 1441: 1380: 1373: 1366: 1357: 1338: 1331: 1325: 1324: 1284: 1278: 1277: 1267: 1249: 1225: 1219: 1212: 1206: 1199: 1193: 1186: 1180: 1173: 1051:Nebrenthorrhinus 991:Huhtyk Formation 959:Dalazi Formation 907:Emanra Formation 894:Yixian Formation 892:Liu et al. 2006 826:Archaeorrhynchus 741:, Russia, Aptian 700:, Brazil, Aptian 583:Ulugei Formation 529: 500:Rhynchitoplesius 238:primitive weevil 220: 215: 172: 67: 66: 44: 27:Temporal range: 21: 3605: 3604: 3600: 3599: 3598: 3596: 3595: 3594: 3570: 3569: 3568: 3563: 3555: 3553: 3545: 3540: 3532: 3527: 3519: 3517: 3509: 3504: 3496: 3491: 3483: 3478: 3470: 3465: 3457: 3452: 3444: 3439: 3431: 3426: 3418: 3413: 3405: 3400: 3392: 3387: 3379: 3374: 3366: 3361: 3353: 3351: 3343: 3338: 3330: 3325: 3316: 3315: 3310: 3301: 3300: 3295: 3282: 3272: 3267: 3246: 3188: 3139: 3121:(clown beetles) 3092: 3063:(betsy beetles) 2971:Scarabaeiformia 2961: 2918: 2893: 2825:(click beetles) 2801:(Texas beetles) 2777: 2748: 2744:Ptilodactylidae 2669:(cedar beetles) 2645: 2606: 2589:Trictenotomidae 2475:Mycetophagidae 2392: 2371: 2308: 2307: 2305: 2297: 2249:Phloeostichidae 2200:Lamingtoniidae 2192:Laemophloeidae 2140:Cryptophagidae 2104: 1995: 1978:Thanerocleridae 1941:Phloiophilidae 1928:Metaxina ornata 1893:Chaetosomatidae 1851: 1784: 1763: 1721:(auger beetles) 1702:Bostrichiformia 1692: 1674: 1670:Torridincolidae 1654:(skiff beetles) 1652:Hydroscaphidae 1643: 1621: 1614:Trachypachidae 1559:(tiger beetles) 1517: 1499: 1449: 1435: 1391: 1384: 1347: 1342: 1341: 1332: 1328: 1286: 1285: 1281: 1227: 1226: 1222: 1213: 1209: 1200: 1196: 1187: 1183: 1174: 1170: 1165: 1138:Guillermorhinus 1094:Selengarhynchus 961:, China, Aptian 896:, China, Aptian 698:Crato Formation 549:Abrenthorrhinus 524: 519: 380:Rhinorhynchinae 318: 287:Rhinorhynchinae 221:Auctt. (missp.) 218: 198:Rhinorhynchinae 171: 61: 35: 25: 17: 12: 11: 5: 3603: 3601: 3593: 3592: 3587: 3582: 3572: 3571: 3565: 3564: 3562: 3561: 3551: 3538: 3525: 3515: 3502: 3489: 3476: 3463: 3450: 3437: 3424: 3415:Fauna Europaea 3411: 3398: 3385: 3372: 3359: 3349: 3336: 3323: 3308: 3292: 3290: 3284: 3283: 3278: 3269: 3268: 3263: 3260: 3259: 3256: 3255: 3252: 3251: 3248: 3247: 3245: 3244: 3241:(rove beetles) 3239:Staphylinidae 3236: 3228: 3220: 3212: 3207: 3198: 3196: 3194:Staphylinoidea 3190: 3189: 3187: 3186: 3181: 3176:Hydrophilidae 3173: 3168: 3163: 3155: 3149: 3147: 3145:Hydrophiloidea 3141: 3140: 3138: 3137: 3132: 3124: 3115: 3113: 3104: 3098: 3097: 3094: 3093: 3091: 3090: 3087:(hide beetles) 3082: 3074: 3071:(rain beetles) 3066: 3058: 3050: 3047:(stag beetles) 3042: 3034: 3026: 3018: 3010: 3002: 3000:Bolboceratidae 2997: 2984: 2982: 2973: 2967: 2966: 2963: 2962: 2960: 2959: 2954: 2946: 2934: 2928: 2926: 2920: 2919: 2917: 2916: 2907:Rhinorhipidae 2903: 2901: 2899:Rhinorhipoidea 2895: 2894: 2892: 2891: 2883: 2878: 2873: 2865: 2857: 2849: 2841: 2836: 2828: 2820: 2812: 2804: 2796: 2787: 2785: 2779: 2778: 2776: 2775: 2767: 2758: 2756: 2750: 2749: 2747: 2746: 2741: 2733: 2725: 2717: 2712:Heteroceridae 2709: 2701: 2693: 2685: 2680: 2675:Chelonariidae 2672: 2667:Callirhipidae 2664: 2661:(pill beetles) 2655: 2653: 2647: 2646: 2644: 2643: 2638: 2629: 2627: 2618: 2612: 2611: 2608: 2607: 2605: 2604: 2596: 2591: 2586: 2578: 2573:Tenebrionidae 2570: 2562: 2554: 2546: 2538: 2530: 2522: 2514: 2509: 2501: 2496: 2488: 2480: 2472: 2464: 2456: 2448: 2440: 2435: 2427: 2419: 2411: 2402: 2400: 2398:Tenebrionoidea 2394: 2393: 2391: 2390: 2381: 2379: 2373: 2372: 2370: 2369: 2361: 2356:Curculionidae 2353: 2348: 2340: 2332: 2324: 2315: 2313: 2303:Curculionoidea 2299: 2298: 2296: 2295: 2290: 2282: 2274: 2266: 2264:Protocucujidae 2261: 2256: 2251: 2246: 2238: 2230: 2222: 2217: 2209: 2197: 2189: 2181: 2176: 2171: 2163: 2158: 2156:Cybocephalidae 2153: 2145: 2137: 2132: 2127: 2114: 2112: 2106: 2105: 2103: 2102: 2097: 2092: 2087: 2079: 2071: 2066: 2058: 2056:Discolomatidae 2053: 2045: 2040:Coccinellidae 2037: 2029: 2024:Bothrideridae 2021: 2016: 2011: 2005: 2003: 2001:Coccinelloidea 1997: 1996: 1994: 1993: 1988:Trogossitidae 1985: 1980: 1975: 1970: 1965: 1960: 1955: 1950: 1938: 1933: 1921: 1913: 1908: 1906:Lophocateridae 1903: 1895: 1890: 1882: 1874: 1861: 1859: 1853: 1852: 1850: 1849: 1844: 1839: 1834: 1829: 1824: 1821:(leaf beetles) 1819:Chrysomelidae 1816: 1807: 1805: 1803:Chrysomeloidea 1796: 1790: 1789: 1786: 1785: 1783: 1782: 1773: 1771: 1765: 1764: 1762: 1761: 1753: 1748:Nosodendridae 1745: 1740:Jacobsoniidae 1737: 1732: 1729:(skin beetles) 1724: 1715: 1713: 1704: 1694: 1693: 1687: 1680: 1679: 1676: 1675: 1673: 1672: 1667: 1662: 1657: 1648: 1645: 1644: 1638: 1631: 1630: 1627: 1626: 1623: 1622: 1620: 1619: 1611: 1603: 1598:Meru phyllisae 1591: 1586: 1578: 1570: 1562: 1554: 1546: 1541: 1532: 1530: 1519: 1518: 1512: 1505: 1504: 1501: 1500: 1498: 1497: 1492: 1490:Micromalthidae 1487: 1475: 1467: 1454: 1451: 1450: 1444: 1437: 1436: 1434: 1433: 1427: 1421: 1415: 1409: 1403: 1396: 1393: 1392: 1385: 1383: 1382: 1375: 1368: 1360: 1354: 1353: 1346: 1345:External links 1343: 1340: 1339: 1326: 1279: 1220: 1207: 1194: 1181: 1167: 1166: 1164: 1161: 1160: 1159: 1158: 1157: 1156: 1155: 1154: 1153: 1142: 1133: 1124: 1102: 1101: 1100: 1099: 1098: 1089: 1073: 1072: 1071: 1070: 1069: 1057: 1056: 1055: 1040: 1039: 1038: 1027:Metrioxenoides 1022: 1006: 997: 981: 966: 965: 964: 963: 962: 947:Incertae sedis 944: 943: 942: 933: 921: 920: 919: 910: 897: 884: 864: 863: 862: 850: 849: 848: 839: 830: 821: 802: 801: 800: 799: 798: 791: 784: 773:Eocaenonemonyx 765: 764: 763: 751: 742: 739:Zaza Formation 723: 722: 721: 720: 719: 710: 701: 688: 676: 675: 674: 661: 652: 643: 627:Distenorrhinus 622: 607: 606: 605: 604: 603: 591: 590: 589: 573: 569:Brenthorrhinus 564: 556:, Kazakhstan, 535: 523: 520: 518: 517: 510: 503: 496: 489: 486:Rhynchitomacer 482: 475: 468: 461: 454: 447: 440: 433: 426: 419: 412: 405: 398: 391: 377: 376: 362: 361: 354: 347: 340: 333: 319: 317: 314: 299:Curculionoidea 279:inflorescences 225: 224: 210: 209: 203: 202: 201: 200: 195: 190: 189:(but see text) 181: 180: 176: 175: 166: 162: 161: 159:Curculionoidea 156: 152: 151: 146: 139: 138: 133: 129: 128: 123: 119: 118: 113: 109: 108: 103: 99: 98: 93: 89: 88: 83: 79: 78: 73: 69: 68: 55: 54: 46: 45: 37: 36: 26: 15: 13: 10: 9: 6: 4: 3: 2: 3602: 3591: 3588: 3586: 3583: 3581: 3578: 3577: 3575: 3558: 3552: 3548: 3543: 3539: 3535: 3530: 3526: 3522: 3516: 3512: 3507: 3503: 3499: 3494: 3490: 3486: 3481: 3477: 3473: 3468: 3464: 3460: 3455: 3451: 3447: 3442: 3438: 3434: 3429: 3425: 3421: 3416: 3412: 3408: 3403: 3399: 3395: 3390: 3386: 3382: 3377: 3373: 3369: 3364: 3360: 3356: 3350: 3346: 3341: 3337: 3333: 3328: 3324: 3319: 3313: 3309: 3304: 3298: 3294: 3293: 3291: 3289: 3285: 3281: 3276: 3266: 3261: 3243: 3237: 3235: 3229: 3227: 3221: 3219: 3213: 3211: 3208: 3206: 3200: 3199: 3197: 3195: 3191: 3185: 3182: 3180: 3174: 3172: 3169: 3167: 3164: 3162: 3156: 3154: 3151: 3150: 3148: 3146: 3142: 3136: 3133: 3131: 3127:Sphaeritidae 3125: 3123: 3117: 3116: 3114: 3112: 3108: 3105: 3103: 3099: 3089: 3083: 3081: 3077:Scarabaeidae 3075: 3073: 3067: 3065: 3059: 3057: 3051: 3049: 3043: 3041: 3035: 3033: 3027: 3025: 3019: 3017: 3015:(dor beetles) 3011: 3009: 3003: 3001: 2998: 2996: 2993: 2986: 2985: 2983: 2981: 2980:Scarabaeoidea 2977: 2974: 2972: 2968: 2958: 2955: 2953: 2947: 2945: 2942: 2935: 2933: 2930: 2929: 2927: 2925: 2921: 2915: 2912: 2905: 2904: 2902: 2900: 2896: 2890: 2884: 2882: 2879: 2877: 2874: 2872: 2866: 2864: 2858: 2856: 2850: 2848: 2842: 2840: 2837: 2835: 2829: 2827: 2821: 2819: 2815:Cerophytidae 2813: 2811: 2805: 2803: 2797: 2795: 2789: 2788: 2786: 2784: 2780: 2774: 2770:Rhipiceridae 2768: 2766: 2760: 2759: 2757: 2755: 2751: 2745: 2742: 2740: 2734: 2732: 2726: 2724: 2718: 2716: 2710: 2708: 2704:Eulichadidae 2702: 2700: 2694: 2692: 2686: 2684: 2683:Cneoglossidae 2681: 2679: 2673: 2671: 2665: 2663: 2657: 2656: 2654: 2652: 2648: 2642: 2641:Schizopodidae 2639: 2637: 2631: 2630: 2628: 2626: 2622: 2619: 2617: 2616:Elateriformia 2613: 2603: 2597: 2595: 2592: 2590: 2587: 2585: 2581:Tetratomidae 2579: 2577: 2571: 2569: 2563: 2561: 2555: 2553: 2547: 2545: 2539: 2537: 2533:Ripiphoridae 2531: 2529: 2523: 2521: 2517:Pyrochroidae 2515: 2513: 2512:Pterogeniidae 2510: 2508: 2502: 2500: 2497: 2495: 2489: 2487: 2481: 2479: 2473: 2471: 2465: 2463: 2457: 2455: 2451:Melandryidae 2449: 2447: 2441: 2439: 2438:Chalcodryidae 2436: 2434: 2428: 2426: 2420: 2418: 2412: 2410: 2404: 2403: 2401: 2399: 2395: 2389: 2383: 2382: 2380: 2378: 2374: 2368: 2364:Nemonychidae 2362: 2360: 2354: 2352: 2349: 2347: 2341: 2339: 2333: 2331: 2325: 2323: 2317: 2316: 2314: 2304: 2300: 2294: 2291: 2289: 2283: 2281: 2275: 2273: 2267: 2265: 2262: 2260: 2259:Propalticidae 2257: 2255: 2254:Priasilphidae 2252: 2250: 2247: 2245: 2239: 2237: 2233:Passandridae 2231: 2229: 2227:(sap beetles) 2223: 2221: 2218: 2216: 2210: 2208: 2205: 2198: 2196: 2190: 2188: 2182: 2180: 2177: 2175: 2172: 2170: 2164: 2162: 2159: 2157: 2154: 2152: 2146: 2144: 2138: 2136: 2135:Cavognathidae 2133: 2131: 2128: 2126: 2123: 2116: 2115: 2113: 2111: 2107: 2101: 2098: 2096: 2093: 2091: 2088: 2086: 2080: 2078: 2072: 2070: 2069:Eupsilobiidae 2067: 2065: 2061:Endomychidae 2059: 2057: 2054: 2052: 2048:Corylophidae 2046: 2044: 2038: 2036: 2030: 2028: 2022: 2020: 2017: 2015: 2012: 2010: 2007: 2006: 2004: 2002: 1998: 1992: 1986: 1984: 1981: 1979: 1976: 1974: 1971: 1969: 1966: 1964: 1963:Protopeltidae 1961: 1959: 1958:Prionoceridae 1956: 1954: 1951: 1949: 1946: 1939: 1937: 1934: 1932: 1929: 1922: 1920: 1914: 1912: 1911:Mauroniscidae 1909: 1907: 1904: 1902: 1896: 1894: 1891: 1889: 1883: 1881: 1875: 1873: 1870: 1863: 1862: 1860: 1858: 1854: 1848: 1845: 1843: 1840: 1838: 1835: 1833: 1832:Megalopodidae 1830: 1828: 1825: 1823: 1817: 1815: 1811:Cerambycidae 1809: 1808: 1806: 1804: 1800: 1797: 1795: 1791: 1781: 1777:Derodontidae 1775: 1774: 1772: 1770: 1769:Derodontoidea 1766: 1760: 1754: 1752: 1746: 1744: 1738: 1736: 1735:Endecatomidae 1733: 1731: 1725: 1723: 1719:Bostrichidae 1717: 1716: 1714: 1712: 1711:Bostrichoidea 1708: 1705: 1703: 1699: 1695: 1691: 1685: 1681: 1671: 1668: 1666: 1665:Sphaeriusidae 1663: 1661: 1658: 1656: 1650: 1649: 1646: 1642: 1636: 1632: 1618: 1612: 1610: 1604: 1602: 1599: 1592: 1590: 1587: 1585: 1579: 1577: 1571: 1569: 1563: 1561: 1557:Cicindelidae 1555: 1553: 1547: 1545: 1542: 1540: 1534: 1533: 1531: 1528: 1524: 1520: 1516: 1510: 1506: 1496: 1493: 1491: 1488: 1486: 1483: 1476: 1474: 1468: 1466: 1463: 1456: 1455: 1452: 1448: 1442: 1438: 1432: 1428: 1426: 1422: 1420: 1416: 1414: 1410: 1408: 1404: 1402: 1398: 1397: 1394: 1389: 1381: 1376: 1374: 1369: 1367: 1362: 1361: 1358: 1352: 1349: 1348: 1344: 1336: 1330: 1327: 1322: 1318: 1314: 1310: 1306: 1302: 1298: 1294: 1290: 1283: 1280: 1275: 1271: 1266: 1261: 1257: 1253: 1248: 1243: 1239: 1235: 1231: 1224: 1221: 1218: 1211: 1208: 1204: 1198: 1195: 1191: 1185: 1182: 1178: 1172: 1169: 1162: 1152: 1151: 1149: 1148: 1143: 1140: 1139: 1134: 1131: 1130: 1125: 1123: 1119: 1118:Burmese amber 1116:Legalov 2018 1115: 1114: 1109: 1108: 1106: 1105: 1103: 1096: 1095: 1090: 1087: 1084:Legalov 2010 1083: 1082: 1077: 1076: 1074: 1067: 1066: 1061: 1060: 1058: 1053: 1052: 1047: 1046: 1044: 1043: 1041: 1037: 1033: 1029: 1028: 1023: 1021: 1017: 1013: 1012: 1007: 1004: 1003: 998: 996: 992: 989:Legalov 2010 988: 987: 986:Cretoxenoides 982: 979: 978: 977:Brasilnemonyx 973: 972: 970: 969: 967: 960: 956: 955: 950: 949: 948: 945: 940: 939: 934: 931: 930: 925: 924: 922: 917: 916: 911: 908: 905:Legalov 2011 904: 903: 898: 895: 891: 890: 889:Microprobelus 885: 882: 878: 874: 873: 872:Oxycorynoides 868: 867: 865: 860: 859: 854: 853: 851: 846: 845: 840: 837: 836: 831: 828: 827: 822: 819: 818: 813: 812: 810: 809: 808:Arnoldi 1977 807: 803: 796: 792: 789: 785: 783: 779: 776:Legalov 2013 775: 774: 769: 768: 766: 762: 758: 757: 756:Turononemonyx 752: 749: 748: 747:Pseudonemonyx 743: 740: 736: 735: 730: 729: 727: 726: 724: 717: 716: 711: 708: 707: 706:Eccoptarthrus 702: 699: 696:Legalov 2014 695: 694: 689: 686: 685: 684:Astenorrhinus 680: 679: 677: 672: 668: 667: 662: 659: 658: 653: 650: 649: 644: 641: 637: 633: 629: 628: 623: 621: 617: 616: 615:Buryatnemonyx 611: 610: 608: 601: 600: 595: 594: 592: 588: 584: 580: 579: 574: 571: 570: 565: 563: 559: 555: 552:Legalov 2009 551: 550: 545: 544: 542: 541: 539: 538: 536: 534: 533: 532:Kuschelomacer 526: 525: 521: 516: 515: 511: 509: 508: 504: 502: 501: 497: 495: 494: 490: 488: 487: 483: 481: 480: 479:Rhinorhynchus 476: 474: 473: 469: 467: 466: 462: 460: 459: 455: 453: 452: 448: 446: 445: 441: 439: 438: 434: 432: 431: 427: 425: 424: 420: 418: 417: 416:Basiliorhinus 413: 411: 410: 406: 404: 403: 399: 397: 396: 392: 390: 389: 385: 384: 383: 381: 375: 374: 370: 369: 368: 366: 360: 359: 355: 353: 352: 348: 346: 345: 344:Doydirhynchus 341: 339: 338: 334: 332: 331: 327: 326: 325: 323: 315: 313: 311: 306: 302: 300: 296: 295:Araucariaceae 292: 291:Podocarpaceae 288: 284: 280: 276: 272: 271: 266: 261: 259: 255: 251: 247: 243: 239: 235: 231: 223: 214:Cimberididae 211: 208: 204: 199: 196: 194: 191: 188: 185: 184: 182: 177: 170: 167: 164: 163: 160: 157: 155:Superfamily: 154: 153: 150: 147: 144: 141: 140: 137: 134: 131: 130: 127: 124: 121: 120: 117: 114: 111: 110: 107: 104: 101: 100: 97: 94: 91: 90: 87: 84: 81: 80: 77: 74: 71: 70: 65: 60: 56: 53: 52: 47: 43: 38: 34: 30: 22: 19: 3590:Nemonychidae 3345:Nemonychidae 3332:Nemonychidae 3318:Nemonychidae 3288:Nemonychidae 3287: 3166:Helophoridae 3158:Georissidae 3153:Epimetopidae 3069:Pleocomidae 3053:Ochodaeidae 3037:Hybosoridae 3021:Glaphyridae 3013:Geotrupidae 2991: 2988:Belohinidae 2949:Eucinetidae 2940: 2937:Decliniidae 2910: 2868:Phengodidae 2807:Cantharidae 2762:Dascillidae 2754:Dascilloidea 2736:Psephenidae 2728:Lutrochidae 2720:Limnichidae 2633:Buprestidae 2625:Buprestoidea 2565:Synchroidae 2549:Scraptiidae 2541:Salpingidae 2504:Prostomidae 2491:Oedemeridae 2467:Mordellidae 2385:Lymexylidae 2377:Lymexyloidea 2363: 2327:Attelabidae 2319:Anthribidae 2277:Smicripidae 2241:Phalacridae 2225:Nitidulidae 2220:Myraboliidae 2212:Monotomidae 2203: 2184:Kateretidae 2161:Cyclaxyridae 2121: 2118:Agapythidae 2082:Latridiidae 2032:Cerylonidae 2019:Anamorphidae 1953:Phycosecidae 1944: 1927: 1924:Metaxinidae 1877:Biphyllidae 1868: 1837:Orsodacnidae 1794:Cucujiformia 1727:Dermestidae 1597: 1536:Amphizoidae 1481: 1461: 1447:Archostemata 1431:Holometabola 1429:Superorder: 1423:Infraclass: 1337:, 41, 29–38. 1334: 1329: 1296: 1292: 1282: 1237: 1233: 1223: 1210: 1202: 1197: 1189: 1184: 1176: 1171: 1147:Renicimberis 1145: 1136: 1127: 1111: 1092: 1079: 1063: 1049: 1025: 1018:, Mongolia, 1009: 1002:Khetanamonyx 1000: 993:, Mongolia, 984: 975: 952: 946: 936: 927: 913: 900: 887: 870: 856: 842: 833: 824: 815: 794: 787: 771: 754: 745: 734:Cretonemonyx 732: 713: 704: 693:Cratonemonyx 691: 682: 664: 655: 646: 625: 613: 597: 585:, Mongolia, 576: 567: 547: 530: 522:Extinct taxa 512: 505: 498: 491: 484: 477: 470: 463: 456: 449: 442: 435: 428: 421: 414: 407: 400: 393: 386: 378: 371: 365:Nemonychinae 363: 356: 349: 342: 335: 328: 322:Cimberidinae 320: 307: 303: 283:Cimberidinae 268: 264: 262: 248:. As in the 245: 237: 230:Nemonychidae 229: 228: 213: 193:Nemonychinae 187:Cimberidinae 179:Subfamilies 169:Nemonychidae 168: 142: 136:Cucujiformia 132:Infraorder: 49: 24:Nemonychidae 18: 3454:iNaturalist 3312:Wikispecies 3210:Hydraenidae 3184:Spercheidae 3171:Hydrochidae 3135:Synteliidae 3119:Histeridae 3111:Histeroidea 3061:Passalidae 3029:Glaresidae 2886:Throscidae 2846:(fireflies) 2844:Lampyridae 2831:Eucnemidae 2823:Elateridae 2783:Elateroidea 2599:Zopheridae 2483:Mycteridae 2414:Anthicidae 2285:Sphindidae 2269:Silvanidae 2179:Hobartiidae 2166:Erotylidae 2095:Mycetaeidae 2090:Murmidiidae 2074:Euxestidae 1968:Rentoniidae 1842:Oxypeltidae 1827:Disteniidae 1660:Lepiceridae 1589:Hygrobiidae 1581:Haliplidae 1565:Dytiscidae 1544:Aspidytidae 1120:, Myanmar, 1065:Paleocartus 1020:Valanginian 995:Hauterivian 957:Zhang 1997 954:Longidorsum 929:Belonotaris 804:†subfamily 715:Procurculio 666:Talbragarus 437:Eutactobius 409:Basiliogeus 395:Araucomacer 351:Lecontellus 316:Extant taxa 250:Anthribidae 219:Cimberidae 216:Gozis, 1882 173:Bedel, 1882 3574:Categories 3231:Silphidae 3223:Ptiliidae 3215:Leiodidae 3202:Agyrtidae 3045:Lucanidae 2924:Scirtoidea 2860:Omethidae 2839:Jurasaidae 2688:Dryopidae 2659:Byrrhidae 2651:Byrrhoidea 2343:Brentidae 2148:Cucujidae 2130:Boganiidae 2110:Cucujoidea 1983:Thymalidae 1973:Rhadalidae 1916:Melyridae 1885:Byturidae 1847:Vesperidae 1756:Ptiniidae 1606:Noteridae 1573:Gyrinidae 1549:Carabidae 1478:Jurodidae 1470:Cupedidae 1417:Subclass: 1407:Arthropoda 1388:Coleoptera 1293:GeoJournal 1163:References 1122:Cenomanian 1113:Burmomacer 1036:Berriasian 915:Cratomacer 877:Weald Clay 514:Zimmiellus 507:Stenomacer 458:Nannomacer 402:Atopomacer 388:Aragomacer 358:Pityomacer 270:Delphinium 149:Phytophaga 122:Suborder: 116:Coleoptera 96:Arthropoda 3085:Trogidae 3079:(scarabs) 2957:Scirtidae 2932:Clambidae 2525:Pythidae 2459:Meloidae 2406:Aderidae 2309:(weevils) 2174:Helotidae 2100:Teredidae 2014:Alexiidae 1898:Cleridae 1857:Cleroidea 1690:Polyphaga 1688:Suborder 1641:Myxophaga 1639:Suborder 1594:Meruidae 1513:Suborder 1495:Ommatidae 1445:Suborder 1419:Pterygota 1399:Kingdom: 1313:0343-2521 1256:0027-8424 858:Ampliceps 806:Eobelinae 640:Barremian 638:, Spain, 636:Las Hoyas 587:Tithonian 562:Oxfordian 558:Callovian 472:Pagomacer 465:Notomacer 451:Mecomacer 444:Idiomacer 330:Acromacer 126:Polyphaga 82:Kingdom: 76:Eukaryota 29:Oxfordian 3376:BugGuide 3352:BioLib: 3297:Wikidata 2852:Lycidae 2696:Elmidae 2594:Ulodidae 2430:Boridae 2335:Belidae 1936:Peltidae 1529:families 1515:Adephaga 1425:Neoptera 1405:Phylum: 1401:Animalia 1390:families 1321:83680109 1274:19365072 1177:Cimberis 1129:Burmonyx 938:Probelus 761:Turonian 430:Bunyaeus 337:Cimberis 242:antennae 207:Synonyms 165:Family: 92:Phylum: 86:Animalia 72:Domain: 3580:Weevils 3303:Q134893 2443:Ciidae 2351:Caridae 1413:Insecta 1411:Class: 1386:Extant 1265:2678426 835:Eobelus 373:Nemonyx 275:Pinales 265:Nemonyx 258:clypeus 234:weevils 112:Order: 106:Insecta 102:Class: 33:Present 3557:360133 3554:uBio: 3534:149636 3518:NZOR: 3485:616722 3472:100991 3459:174369 3368:211306 3355:243767 1527:Extant 1319:  1311:  1272:  1262:  1254:  782:Eocene 620:Aptian 423:Brarus 254:labrum 252:, the 3547:69422 3511:27443 3467:IRMNG 3420:11171 3394:8HBR3 3381:38594 1317:S2CID 1299:(6). 310:amber 143:Clade 3506:NCBI 3480:ITIS 3446:8584 3441:GBIF 3407:5233 3363:BOLD 1309:ISSN 1270:PMID 1252:ISSN 293:and 3493:NBN 3402:EoL 3389:CoL 3340:AFD 3327:ADW 1301:doi 1260:PMC 1242:doi 1238:106 3576:: 3544:: 3531:: 3508:: 3495:: 3482:: 3469:: 3456:: 3443:: 3430:: 3417:: 3404:: 3391:: 3378:: 3365:: 3342:: 3329:: 3314:: 3299:: 1315:. 1307:. 1295:. 1291:. 1268:. 1258:. 1250:. 1236:. 1232:. 793:†" 786:†" 634:, 382:: 367:: 324:: 312:. 281:. 145:: 2994:) 2990:( 2943:) 2939:( 2913:) 2909:( 2206:) 2202:( 2124:) 2120:( 1947:) 1943:( 1930:) 1926:( 1871:) 1867:( 1600:) 1596:( 1484:) 1480:( 1464:) 1460:( 1379:e 1372:t 1365:v 1323:. 1303:: 1297:7 1276:. 1244:: 1144:† 1135:† 1126:† 1110:† 1091:† 1078:† 1062:† 1048:† 1024:† 1008:† 999:† 983:† 974:† 951:† 935:† 926:† 912:† 899:† 886:† 869:† 855:† 841:† 832:† 823:† 814:† 770:† 753:† 744:† 731:† 712:† 703:† 690:† 681:† 663:† 654:† 645:† 624:† 612:† 596:† 575:† 566:† 560:/ 546:† 528:† 31:-

Index

Oxfordian
Present

Doydirhynchus austriacus
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Coleoptera
Polyphaga
Cucujiformia
Phytophaga
Curculionoidea
Nemonychidae
Cimberidinae
Nemonychinae
Rhinorhynchinae
Synonyms
weevils
antennae
Anthribidae
labrum
clypeus
Delphinium
Pinales
inflorescences
Cimberidinae
Rhinorhynchinae

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