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Nemonychidae

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75: 53: 271:, and the maxillary palps are long and projecting. Nemonychidae have all ventrites free, while Anthribidae have ventrites 1-4 connate or partially fused. Nemonychidae lack lateral carinae on the pronotum, while these are usually present, though may be short, in Anthribidae. 1225:
Shin S, Clarke DJ, Lemmon AR, Moriarty Lemmon E, Aitken AL, Haddad S, Farrell BD, Marvaldi AE, Oberprieler RG, McKenna DD (2018) Phylogenomic data yield new and robust insights into the phylogeny and evolution of weevils. Mol. Biol. Evol. 35: 823-836.
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Both Nemonychidae adults and larvae feed primarily on pollen. When mature, the larvae will drop to the ground from the male cones or flowers they resided in to pupate in the soil less than five centimeters from the surface.
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ICZN (2005) Opinion 2111 (Case 3093). Nemonychidae Bedel, November 1882 (Insecta, Coleoptera): given precedence over Cimberididae Gozis, March 1882; and
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Riedel, A. (2010). A new tribe, genus and species of Nemonychidae from Baltic amber (Coleoptera: Curculionoidea: Nemonychidae: Cimberidinae).
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Anderson, R.S, Oberprieler, R.G., Marvaldi, A.E. 2014. 3.1 Nemonychidae Bedel, 1882. In: Leschen R.A.B. & Beutel, R.G. (Eds)
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Nemonychidae are historically divided into three subfamilies: Nemonychinae of the palearctic region with the single genus
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Kuschel, G. & Leschen, R. A. (2011). Phylogeny and taxonomy of the Rhinorhynchinae (Coleoptera: Nemonychidae).
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There exists a fairly extensive fossil record of Nemonychidae reaching from the upper Jurassic to tertiary
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McKenna, Duane D.; Sequeira, Andrea S.; Marvaldi, Adriana E.; Farrell, Brian D. (2009-04-13).
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Handbook of Zoology, Coleoptera, Beetles Volume 3: Morphology and Systematics (Phytophaga)
708: 3438: 3425: 3204: 3155: 3010: 2817: 2485: 2469: 2408: 2303: 2274: 2202: 2166: 2150: 2066: 2011: 1916: 1813: 1662: 1624: 1604: 1500: 1275: 1240: 749: 309: 3584: 3342: 3186: 2990: 2981: 2722: 2693: 2685: 2677: 2651: 2626: 2522: 2509: 2448: 2366: 2269: 2264: 2145: 2079: 2050: 2034: 1998: 1968: 1921: 1875: 1842: 1758: 1750: 1721: 1559: 1128: 489: 354: 305: 301: 289: 3355: 1331: 3404: 3163: 3087: 2833: 2825: 2780: 2764: 2714: 2635: 2591: 2543: 2461: 2243: 2230: 2210: 2171: 2071: 2058: 2029: 1963: 1951: 1847: 1804: 1787: 1779: 1729: 1567: 1537: 1468: 1457: 1441: 146: 1241:"Temporal lags and overlap in the diversification of weevils and flowering plants" 1300:"Past and present of the relict family nemonychidae (Coleoptera, Curculionoidea)" 1190:
Gozis, 1881: usage conserved. Bulletin of Zoological Nomenclature 62(2): 101–103.
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Gratshev and Zherikhin 1995 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Gratshev and Zherikhin 1995 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Gratshev and Zherikhin 1995 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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Gratshev and Zherikhin 2000 La Pedrera de Rúbies Formation, Spain, Barremian
650: 646: 597: 568: 340: 136: 106: 86: 17: 1284: 3365: 3386: 3307: 3145: 3095: 3039: 2998: 2947: 2604: 2535: 2416: 2128: 1934: 1908: 1766: 1599: 1546: 1525: 1435: 771: 347: 43: 1299: 1227: 3391: 2862: 2854: 2706: 2440: 2361: 2345: 1315: 285: 244: 3469: 2453: 2313: 1946: 1423: 1411: 1398: 792: 641:
Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian,
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Gratshev and Legalov 2009 Sharin-Gol Formation, Mongolia, Barremian
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Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
840:
Martynov 1926 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
1079:
Legalov 2003 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
952:
Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
943:
Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
872:
Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
858:
Legalov 2010 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
849:
Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
831:
Legalov 2009 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
729:
Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
720:
Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
583:
Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
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group because they have straight rather than geniculate (elbowed)
312:, and it has been proposed to elevate the group to family rank. 284:. Most species of the other two subfamilies are associated with 3288: 1697: 1648: 1522: 1454: 1370: 1152:
Clarke and Oberprieler 2018 Burmese amber, Myanmar, Cenomanian
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occur largely in the Southern hemisphere, especially found on
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Legalov 2003 La Pedrera de Rúbies Formation, Spain, Barremian
929:
Zherikhin and Gratshev 2004 Crato Formation, Brazil, Aptian
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Scudder 1876 Green River Formation, United States, Eocene
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Scudder 1878 Green River Formation, United States, Eocene
1769:(furniture beetles, death watch beetles, spider beetles) 761:
Gratshev and Legalov 2009 Zaza Formation, Russia, Aptian
1143:
Davis and Engel 2014 Burmese amber, Myanmar, Cenomanian
736:†subfamily Cretonemonychinae Gratshev and Legalov 2009 538: 3297: 3203: 3154: 3120: 3111: 2989: 2980: 2933: 2908: 2792: 2763: 2660: 2634: 2625: 2407: 2386: 2312: 2119: 2010: 1866: 1812: 1803: 1778: 1720: 1711: 1536: 1086:†tribe Selengarhynchini Gratshev and Legalov 2009 739:†tribe Cretonemonychini Gratshev and Legalov 2009 2646:(jewel beetles, or metallic wood-boring beetles) 1245:Proceedings of the National Academy of Sciences 1016:Legalov 2009 Emanra Formation, Russia, Turonian 2369:(true weevils, bark beetles, ambrosia beetles) 1382: 1362:Images of Nemonychidae species in New Zealand 1161:Legalov 2009 Yixian Formation, China, Aptian 8: 2612:(ironclad beetles, cylindrical bark beetles) 991:Legalov 2009 Crato Formation, Brazil, Aptian 296:are found in the Northern hemisphere, while 3285: 3117: 2986: 2631: 1809: 1717: 1708: 1694: 1645: 1533: 1519: 1451: 1389: 1375: 1367: 604:†subtribe Brenthorrhinoidina Legalov 2003 51: 31: 3276:List of subgroups of the order Coleoptera 2356:(straight snout weevils, New York weevil) 1274: 1256: 1056:†subtribe Nebrenthorrhinina Legalov 2007 548:†subfamily Brenthorrhininae Arnoldi 1977 2783:(cicada beetle, cicada parasite beetles) 629:Legalov 2010 Khasurty locality, Russia, 1179: 778:†tribe Eocaenonemonychini Legalov 2013 554:†subtribe Brenthorrhinina Arnoldi 1977 2873:(false fireflies, long-lipped beetles) 979:†subfamily Paleocartinae Legalov 2003 1228:https://doi.org/10.1093/molbev/msx324 267:appears as a separate segment to the 7: 3532:9f96fec3-e2bc-44b9-bb9a-8393cea68708 3444:e5a66d51-3437-4b8c-acb8-d0088b94b944 2899:(false metallic wood-boring beetles) 1115:Subfamily Rhinorhynchinae Voss 1922 1070:†subtribe Paleocartina Legalov 2003 982:†tribe Metrioxenoidini Legalov 2009 770:Legalov 2014 Kzyl-Zhar, Kazakhstan, 620:†tribe Distenorrhinini Arnoldi 1977 551:†tribe Brenthorrhinini Arnoldi 1977 278:and an unusual host, the angiosperm 877:†tribe Oxycorynoidini Arnoldi 1977 689:†tribe Eccoptarthrini Arnoldi 1977 1346:Insect Systematics & Evolution 863:†tribe Karataucarini Legalov 2009 288:feeding on the pollen of the male 25: 1203:. DeGruyter, Berlin, pp. 301–398. 1053:†tribe Paleocartini Legalov 2003 680:Oberprieler and Oberprieler 2012 653:Khasurty locality, Russia, Aptian 2510:Perimylopidae, or Promecheilidae 2095:(minute brown scavenger beetles) 1118:Tribe Mecomacerini Kuschel 1994 73: 2725:(variegated mud-loving beetles) 934:†tribe Probelini Legalov 2009 643:La Pedrera de Rúbies Formation 1: 2554:(narrow-waisted bark beetles) 2246:(parasitic flat bark beetles) 2197:(short-winged flower beetles) 2053:(lady beetles, or God's cows) 1790:(tooth-necked fungus beetles) 822:†tribe Eobelini Arnoldi 1977 3066:(sand-loving scarab beetles) 2456:(minute tree-fungus beetles) 2282:(silvanid flat bark beetles) 890:, United Kingdom, Barremian 3215:(primitive carrion beetles) 3171:(minute mud-loving beetles) 3034:(bumble bee scarab beetles) 2804:(soft-bodied plant beetles) 2775:(soft bodied plant beetles) 1578:(predaceous diving beetles) 3617: 3050:(scavenger scarab beetles) 3042:(enigmatic scarab beetles) 2215:Lamingtonium binnaberrense 1929:(soft-wing flower beetles) 1025:Gratshev and Legalov 2009 748:Gratshev and Legalov 2009 592:Gratshev and Legalov 2009 3273: 3189:(water scavenger beetles) 2922:Rhinorhipus tamborinensis 2701:(long-toed water beetles) 2594:(polypore fungus beetles) 2496:(palm and flower beetles) 2480:(tumbling flower beetles) 2427:(ant-like flower beetles) 2205:(lined flat bark beetles) 2179:(pleasing fungus beetles) 2074:(handsome fungus beetles) 1707: 1693: 1658: 1644: 1619:(burrowing water beetles) 1532: 1518: 1464: 1450: 1406: 223: 216: 194: 189: 70:Scientific classification 68: 59: 50: 34: 3236:(feather-winged beetles) 2570:(false longhorn beetles) 2517:(jugular-horned beetles) 2464:(false darkling beetles) 2435:(cryptic fungus beetles) 2254:(shining flower beetles) 1594:(crawling water beetles) 1493:Sikhotealinia zhiltzovae 1214:Invertebrate Systematics 255:. They are often called 62:Doydirhynchus austriacus 2717:(forest stream beetles) 2578:(synchroa bark beetles) 2538:(dead log bark beetles) 2530:(fire-coloured beetles) 2419:(ant-like leaf beetles) 2153:(silken fungus beetles) 2087:(well polished beetles) 2061:(minute fungus beetles) 1880:Acanthocnemus nigricans 1258:10.1073/pnas.0810618106 3228:(round fungus beetles) 2562:(false flower beetles) 2546:(wedge-shaped beetles) 2504:(false blister beetle) 2488:(hairy fungus beetles) 2443:(conifer bark beetles) 2340:(leaf-rolling weevils) 2001:(bark-gnawing beetles) 1761:(wounded-tree beetles) 1627:(false ground beetles) 1549:(trout-stream beetles) 799:Eugnamptus" decemsatus 684:, Australia, Tithonian 3553:Paleobiology Database 3140:(false clown beetles) 2962:(plate-thigh beetles) 2844:(false click beetles) 2749:(water-penny beetles) 2398:(ship-timber beetles) 2377:(pine flower weevils) 2225:(root-eating beetles) 2045:(minute bark beetles) 1956:Phloiophilus edwardsi 1483:(reticulated beetles) 1099:, Mongolia, Barremian 1041:Gratshev et al. 1998 1027:Tsagaantsav Formation 892:Gurvan-Eren Formation 789:Green River Formation 243:is a small family of 3439:Fauna Europaea (new) 3018:(false stag beetles) 3003:Belohina inexpectata 2865:(net-winged beetles) 2828:(rare click beetles) 2741:(travertine beetles) 2733:(minute mud beetles) 2298:(dry-fungus beetles) 2133:Agapytho foveicollis 1890:(false skin beetles) 1753:(Jacobson's beetles) 1473:Crowsoniella relicta 1097:Sharin-Gol Formation 682:Talbragar Fossil bed 565:Karabastau Formation 247:, placed within the 3016:Diphyllostomatidae 2348:(primitive weevils) 2161:(flat bark beetles) 1911:(checkered beetles) 1898:(fruitworm beetles) 1586:(whirligig beetles) 1298:Kusche, G. (1983). 828:Archaeorrhynchoides 668:Microbrenthorrhinus 257:pine flower weevils 3596:Polyphaga families 2881:(glowworm beetles) 2586:(darkling beetles) 2433:Archeocrypticidae 2290:(palmetto beetles) 2037:(dry bark beetles) 1824:(longhorn beetles) 1316:10.1007/bf00218522 1092:Selengarhynchoides 1045:, United Kingdom, 1043:Lulworth Formation 1022:Megametrioxenoides 920:, Russia, Turonian 894:, Mongolia, Aptian 806:Sitona" grandaevus 659:Megabrenthorrhinus 589:Gobibrenthorrhinus 504:Rhynchitomacerinus 3578: 3577: 3540:Open Tree of Life 3291:Taxon identifiers 3282: 3281: 3269: 3268: 3265: 3264: 3261: 3260: 3253: 3245: 3244:(carrion beetles) 3237: 3229: 3216: 3190: 3172: 3141: 3133: 3107: 3106: 3099: 3091: 3083: 3075: 3067: 3059: 3051: 3043: 3035: 3027: 3019: 3006: 2976: 2975: 2963: 2955: 2925: 2900: 2882: 2874: 2866: 2858: 2845: 2837: 2829: 2821: 2820:(soldier beetles) 2813: 2810:Brachypsectridae 2805: 2784: 2776: 2750: 2742: 2734: 2726: 2718: 2710: 2702: 2689: 2681: 2673: 2647: 2621: 2620: 2613: 2595: 2587: 2579: 2571: 2568:Stenotrachelidae 2563: 2555: 2547: 2539: 2531: 2518: 2505: 2497: 2489: 2481: 2473: 2472:(blister beetles) 2465: 2457: 2444: 2436: 2428: 2420: 2399: 2378: 2370: 2357: 2349: 2341: 2333: 2323: 2299: 2291: 2283: 2255: 2247: 2239: 2226: 2218: 2206: 2198: 2180: 2162: 2154: 2136: 2096: 2088: 2075: 2062: 2054: 2046: 2038: 2020:Akalyptoischiidae 2002: 1959: 1942: 1930: 1912: 1899: 1891: 1883: 1833: 1825: 1799: 1798: 1791: 1770: 1762: 1754: 1741: 1733: 1689: 1688: 1666: 1640: 1639: 1636: 1635: 1628: 1620: 1612: 1595: 1587: 1579: 1571: 1563: 1550: 1514: 1513: 1496: 1484: 1476: 1251:(17): 7083–7088. 1216:, 24(6), 573-615. 913:Khetoxycorynoides 855:Martynovirhynchus 791:, United States, 610:Brenthorrhinoides 238: 237: 233: 228: 185: 27:Family of beetles 16:(Redirected from 3608: 3571: 3570: 3561: 3560: 3548: 3547: 3535: 3534: 3525: 3524: 3512: 3511: 3509:NHMSYS0020152782 3499: 3498: 3486: 3485: 3473: 3472: 3460: 3459: 3447: 3446: 3434: 3433: 3421: 3420: 3408: 3407: 3395: 3394: 3382: 3381: 3369: 3368: 3359: 3358: 3346: 3345: 3333: 3332: 3331: 3318: 3317: 3316: 3286: 3251: 3243: 3235: 3227: 3214: 3188: 3170: 3139: 3131: 3118: 3113:Staphyliniformia 3097: 3089: 3081: 3073: 3065: 3057: 3049: 3041: 3033: 3025: 3017: 3000: 2987: 2961: 2952:Declinia relicta 2949: 2919: 2898: 2892:Sinopyrophoridae 2887:Rhagophthalmidae 2880: 2872: 2864: 2856: 2843: 2835: 2827: 2819: 2811: 2803: 2802:Artematopodidae 2782: 2774: 2748: 2740: 2732: 2724: 2716: 2709:(riffle beetles) 2708: 2700: 2688:(turtle beetles) 2687: 2679: 2671: 2645: 2632: 2611: 2593: 2585: 2577: 2569: 2561: 2553: 2545: 2537: 2529: 2516: 2503: 2495: 2487: 2479: 2471: 2463: 2455: 2442: 2434: 2426: 2418: 2397: 2376: 2368: 2355: 2347: 2339: 2332:(fungus weevils) 2331: 2322: 2321: 2317: 2304:Tasmosalpingidae 2297: 2289: 2281: 2253: 2245: 2237: 2224: 2212: 2204: 2196: 2178: 2160: 2152: 2130: 2094: 2086: 2073: 2060: 2052: 2044: 2036: 2000: 1953: 1936: 1928: 1910: 1897: 1889: 1877: 1876:Acanthocnemidae 1831: 1823: 1810: 1789: 1768: 1760: 1752: 1739: 1731: 1718: 1709: 1695: 1664: 1646: 1626: 1618: 1606: 1593: 1585: 1577: 1569: 1562:(ground beetles) 1561: 1548: 1534: 1520: 1490: 1482: 1470: 1469:Crowsoniellidae 1452: 1391: 1384: 1377: 1368: 1349: 1342: 1336: 1335: 1295: 1289: 1288: 1278: 1260: 1236: 1230: 1223: 1217: 1210: 1204: 1197: 1191: 1184: 1062:Nebrenthorrhinus 1002:Huhtyk Formation 970:Dalazi Formation 918:Emanra Formation 905:Yixian Formation 903:Liu et al. 2006 837:Archaeorrhynchus 752:, Russia, Aptian 711:, Brazil, Aptian 594:Ulugei Formation 540: 511:Rhynchitoplesius 249:primitive weevil 231: 226: 183: 78: 77: 55: 38:Temporal range: 32: 21: 3616: 3615: 3611: 3610: 3609: 3607: 3606: 3605: 3581: 3580: 3579: 3574: 3566: 3564: 3556: 3551: 3543: 3538: 3530: 3528: 3520: 3515: 3507: 3502: 3494: 3489: 3481: 3476: 3468: 3463: 3455: 3450: 3442: 3437: 3429: 3424: 3416: 3411: 3403: 3398: 3390: 3385: 3377: 3372: 3364: 3362: 3354: 3349: 3341: 3336: 3327: 3326: 3321: 3312: 3311: 3306: 3293: 3283: 3278: 3257: 3199: 3150: 3132:(clown beetles) 3103: 3074:(betsy beetles) 2982:Scarabaeiformia 2972: 2929: 2904: 2836:(click beetles) 2812:(Texas beetles) 2788: 2759: 2755:Ptilodactylidae 2680:(cedar beetles) 2656: 2617: 2600:Trictenotomidae 2486:Mycetophagidae 2403: 2382: 2319: 2318: 2316: 2308: 2260:Phloeostichidae 2211:Lamingtoniidae 2203:Laemophloeidae 2151:Cryptophagidae 2115: 2006: 1989:Thanerocleridae 1952:Phloiophilidae 1939:Metaxina ornata 1904:Chaetosomatidae 1862: 1795: 1774: 1732:(auger beetles) 1713:Bostrichiformia 1703: 1685: 1681:Torridincolidae 1665:(skiff beetles) 1663:Hydroscaphidae 1654: 1632: 1625:Trachypachidae 1570:(tiger beetles) 1528: 1510: 1460: 1446: 1402: 1395: 1358: 1353: 1352: 1343: 1339: 1297: 1296: 1292: 1238: 1237: 1233: 1224: 1220: 1211: 1207: 1198: 1194: 1185: 1181: 1176: 1149:Guillermorhinus 1105:Selengarhynchus 972:, China, Aptian 907:, China, Aptian 709:Crato Formation 560:Abrenthorrhinus 535: 530: 391:Rhinorhynchinae 329: 298:Rhinorhynchinae 232:Auctt. (missp.) 229: 209:Rhinorhynchinae 182: 72: 46: 36: 28: 23: 22: 15: 12: 11: 5: 3614: 3612: 3604: 3603: 3598: 3593: 3583: 3582: 3576: 3575: 3573: 3572: 3562: 3549: 3536: 3526: 3513: 3500: 3487: 3474: 3461: 3448: 3435: 3426:Fauna Europaea 3422: 3409: 3396: 3383: 3370: 3360: 3347: 3334: 3319: 3303: 3301: 3295: 3294: 3289: 3280: 3279: 3274: 3271: 3270: 3267: 3266: 3263: 3262: 3259: 3258: 3256: 3255: 3252:(rove beetles) 3250:Staphylinidae 3247: 3239: 3231: 3223: 3218: 3209: 3207: 3205:Staphylinoidea 3201: 3200: 3198: 3197: 3192: 3187:Hydrophilidae 3184: 3179: 3174: 3166: 3160: 3158: 3156:Hydrophiloidea 3152: 3151: 3149: 3148: 3143: 3135: 3126: 3124: 3115: 3109: 3108: 3105: 3104: 3102: 3101: 3098:(hide beetles) 3093: 3085: 3082:(rain beetles) 3077: 3069: 3061: 3058:(stag beetles) 3053: 3045: 3037: 3029: 3021: 3013: 3011:Bolboceratidae 3008: 2995: 2993: 2984: 2978: 2977: 2974: 2973: 2971: 2970: 2965: 2957: 2945: 2939: 2937: 2931: 2930: 2928: 2927: 2918:Rhinorhipidae 2914: 2912: 2910:Rhinorhipoidea 2906: 2905: 2903: 2902: 2894: 2889: 2884: 2876: 2868: 2860: 2852: 2847: 2839: 2831: 2823: 2815: 2807: 2798: 2796: 2790: 2789: 2787: 2786: 2778: 2769: 2767: 2761: 2760: 2758: 2757: 2752: 2744: 2736: 2728: 2723:Heteroceridae 2720: 2712: 2704: 2696: 2691: 2686:Chelonariidae 2683: 2678:Callirhipidae 2675: 2672:(pill beetles) 2666: 2664: 2658: 2657: 2655: 2654: 2649: 2640: 2638: 2629: 2623: 2622: 2619: 2618: 2616: 2615: 2607: 2602: 2597: 2589: 2584:Tenebrionidae 2581: 2573: 2565: 2557: 2549: 2541: 2533: 2525: 2520: 2512: 2507: 2499: 2491: 2483: 2475: 2467: 2459: 2451: 2446: 2438: 2430: 2422: 2413: 2411: 2409:Tenebrionoidea 2405: 2404: 2402: 2401: 2392: 2390: 2384: 2383: 2381: 2380: 2372: 2367:Curculionidae 2364: 2359: 2351: 2343: 2335: 2326: 2324: 2314:Curculionoidea 2310: 2309: 2307: 2306: 2301: 2293: 2285: 2277: 2275:Protocucujidae 2272: 2267: 2262: 2257: 2249: 2241: 2233: 2228: 2220: 2208: 2200: 2192: 2187: 2182: 2174: 2169: 2167:Cybocephalidae 2164: 2156: 2148: 2143: 2138: 2125: 2123: 2117: 2116: 2114: 2113: 2108: 2103: 2098: 2090: 2082: 2077: 2069: 2067:Discolomatidae 2064: 2056: 2051:Coccinellidae 2048: 2040: 2035:Bothrideridae 2032: 2027: 2022: 2016: 2014: 2012:Coccinelloidea 2008: 2007: 2005: 2004: 1999:Trogossitidae 1996: 1991: 1986: 1981: 1976: 1971: 1966: 1961: 1949: 1944: 1932: 1924: 1919: 1917:Lophocateridae 1914: 1906: 1901: 1893: 1885: 1872: 1870: 1864: 1863: 1861: 1860: 1855: 1850: 1845: 1840: 1835: 1832:(leaf beetles) 1830:Chrysomelidae 1827: 1818: 1816: 1814:Chrysomeloidea 1807: 1801: 1800: 1797: 1796: 1794: 1793: 1784: 1782: 1776: 1775: 1773: 1772: 1764: 1759:Nosodendridae 1756: 1751:Jacobsoniidae 1748: 1743: 1740:(skin beetles) 1735: 1726: 1724: 1715: 1705: 1704: 1698: 1691: 1690: 1687: 1686: 1684: 1683: 1678: 1673: 1668: 1659: 1656: 1655: 1649: 1642: 1641: 1638: 1637: 1634: 1633: 1631: 1630: 1622: 1614: 1609:Meru phyllisae 1602: 1597: 1589: 1581: 1573: 1565: 1557: 1552: 1543: 1541: 1530: 1529: 1523: 1516: 1515: 1512: 1511: 1509: 1508: 1503: 1501:Micromalthidae 1498: 1486: 1478: 1465: 1462: 1461: 1455: 1448: 1447: 1445: 1444: 1438: 1432: 1426: 1420: 1414: 1407: 1404: 1403: 1396: 1394: 1393: 1386: 1379: 1371: 1365: 1364: 1357: 1356:External links 1354: 1351: 1350: 1337: 1290: 1231: 1218: 1205: 1192: 1178: 1177: 1175: 1172: 1171: 1170: 1169: 1168: 1167: 1166: 1165: 1164: 1153: 1144: 1135: 1113: 1112: 1111: 1110: 1109: 1100: 1084: 1083: 1082: 1081: 1080: 1068: 1067: 1066: 1051: 1050: 1049: 1038:Metrioxenoides 1033: 1017: 1008: 992: 977: 976: 975: 974: 973: 958:Incertae sedis 955: 954: 953: 944: 932: 931: 930: 921: 908: 895: 875: 874: 873: 861: 860: 859: 850: 841: 832: 813: 812: 811: 810: 809: 802: 795: 784:Eocaenonemonyx 776: 775: 774: 762: 753: 750:Zaza Formation 734: 733: 732: 731: 730: 721: 712: 699: 687: 686: 685: 672: 663: 654: 638:Distenorrhinus 633: 618: 617: 616: 615: 614: 602: 601: 600: 584: 580:Brenthorrhinus 575: 567:, Kazakhstan, 546: 534: 531: 529: 528: 521: 514: 507: 500: 497:Rhynchitomacer 493: 486: 479: 472: 465: 458: 451: 444: 437: 430: 423: 416: 409: 402: 388: 387: 373: 372: 365: 358: 351: 344: 330: 328: 325: 310:Curculionoidea 290:inflorescences 236: 235: 221: 220: 214: 213: 212: 211: 206: 201: 200:(but see text) 192: 191: 187: 186: 177: 173: 172: 170:Curculionoidea 167: 163: 162: 157: 150: 149: 144: 140: 139: 134: 130: 129: 124: 120: 119: 114: 110: 109: 104: 100: 99: 94: 90: 89: 84: 80: 79: 66: 65: 57: 56: 48: 47: 37: 26: 24: 14: 13: 10: 9: 6: 4: 3: 2: 3613: 3602: 3599: 3597: 3594: 3592: 3589: 3588: 3586: 3569: 3563: 3559: 3554: 3550: 3546: 3541: 3537: 3533: 3527: 3523: 3518: 3514: 3510: 3505: 3501: 3497: 3492: 3488: 3484: 3479: 3475: 3471: 3466: 3462: 3458: 3453: 3449: 3445: 3440: 3436: 3432: 3427: 3423: 3419: 3414: 3410: 3406: 3401: 3397: 3393: 3388: 3384: 3380: 3375: 3371: 3367: 3361: 3357: 3352: 3348: 3344: 3339: 3335: 3330: 3324: 3320: 3315: 3309: 3305: 3304: 3302: 3300: 3296: 3292: 3287: 3277: 3272: 3254: 3248: 3246: 3240: 3238: 3232: 3230: 3224: 3222: 3219: 3217: 3211: 3210: 3208: 3206: 3202: 3196: 3193: 3191: 3185: 3183: 3180: 3178: 3175: 3173: 3167: 3165: 3162: 3161: 3159: 3157: 3153: 3147: 3144: 3142: 3138:Sphaeritidae 3136: 3134: 3128: 3127: 3125: 3123: 3119: 3116: 3114: 3110: 3100: 3094: 3092: 3088:Scarabaeidae 3086: 3084: 3078: 3076: 3070: 3068: 3062: 3060: 3054: 3052: 3046: 3044: 3038: 3036: 3030: 3028: 3026:(dor beetles) 3022: 3020: 3014: 3012: 3009: 3007: 3004: 2997: 2996: 2994: 2992: 2991:Scarabaeoidea 2988: 2985: 2983: 2979: 2969: 2966: 2964: 2958: 2956: 2953: 2946: 2944: 2941: 2940: 2938: 2936: 2932: 2926: 2923: 2916: 2915: 2913: 2911: 2907: 2901: 2895: 2893: 2890: 2888: 2885: 2883: 2877: 2875: 2869: 2867: 2861: 2859: 2853: 2851: 2848: 2846: 2840: 2838: 2832: 2830: 2826:Cerophytidae 2824: 2822: 2816: 2814: 2808: 2806: 2800: 2799: 2797: 2795: 2791: 2785: 2781:Rhipiceridae 2779: 2777: 2771: 2770: 2768: 2766: 2762: 2756: 2753: 2751: 2745: 2743: 2737: 2735: 2729: 2727: 2721: 2719: 2715:Eulichadidae 2713: 2711: 2705: 2703: 2697: 2695: 2694:Cneoglossidae 2692: 2690: 2684: 2682: 2676: 2674: 2668: 2667: 2665: 2663: 2659: 2653: 2652:Schizopodidae 2650: 2648: 2642: 2641: 2639: 2637: 2633: 2630: 2628: 2627:Elateriformia 2624: 2614: 2608: 2606: 2603: 2601: 2598: 2596: 2592:Tetratomidae 2590: 2588: 2582: 2580: 2574: 2572: 2566: 2564: 2558: 2556: 2550: 2548: 2544:Ripiphoridae 2542: 2540: 2534: 2532: 2528:Pyrochroidae 2526: 2524: 2523:Pterogeniidae 2521: 2519: 2513: 2511: 2508: 2506: 2500: 2498: 2492: 2490: 2484: 2482: 2476: 2474: 2468: 2466: 2462:Melandryidae 2460: 2458: 2452: 2450: 2449:Chalcodryidae 2447: 2445: 2439: 2437: 2431: 2429: 2423: 2421: 2415: 2414: 2412: 2410: 2406: 2400: 2394: 2393: 2391: 2389: 2385: 2379: 2375:Nemonychidae 2373: 2371: 2365: 2363: 2360: 2358: 2352: 2350: 2344: 2342: 2336: 2334: 2328: 2327: 2325: 2315: 2311: 2305: 2302: 2300: 2294: 2292: 2286: 2284: 2278: 2276: 2273: 2271: 2270:Propalticidae 2268: 2266: 2265:Priasilphidae 2263: 2261: 2258: 2256: 2250: 2248: 2244:Passandridae 2242: 2240: 2238:(sap beetles) 2234: 2232: 2229: 2227: 2221: 2219: 2216: 2209: 2207: 2201: 2199: 2193: 2191: 2188: 2186: 2183: 2181: 2175: 2173: 2170: 2168: 2165: 2163: 2157: 2155: 2149: 2147: 2146:Cavognathidae 2144: 2142: 2139: 2137: 2134: 2127: 2126: 2124: 2122: 2118: 2112: 2109: 2107: 2104: 2102: 2099: 2097: 2091: 2089: 2083: 2081: 2080:Eupsilobiidae 2078: 2076: 2072:Endomychidae 2070: 2068: 2065: 2063: 2059:Corylophidae 2057: 2055: 2049: 2047: 2041: 2039: 2033: 2031: 2028: 2026: 2023: 2021: 2018: 2017: 2015: 2013: 2009: 2003: 1997: 1995: 1992: 1990: 1987: 1985: 1982: 1980: 1977: 1975: 1974:Protopeltidae 1972: 1970: 1969:Prionoceridae 1967: 1965: 1962: 1960: 1957: 1950: 1948: 1945: 1943: 1940: 1933: 1931: 1925: 1923: 1922:Mauroniscidae 1920: 1918: 1915: 1913: 1907: 1905: 1902: 1900: 1894: 1892: 1886: 1884: 1881: 1874: 1873: 1871: 1869: 1865: 1859: 1856: 1854: 1851: 1849: 1846: 1844: 1843:Megalopodidae 1841: 1839: 1836: 1834: 1828: 1826: 1822:Cerambycidae 1820: 1819: 1817: 1815: 1811: 1808: 1806: 1802: 1792: 1788:Derodontidae 1786: 1785: 1783: 1781: 1780:Derodontoidea 1777: 1771: 1765: 1763: 1757: 1755: 1749: 1747: 1746:Endecatomidae 1744: 1742: 1736: 1734: 1730:Bostrichidae 1728: 1727: 1725: 1723: 1722:Bostrichoidea 1719: 1716: 1714: 1710: 1706: 1702: 1696: 1692: 1682: 1679: 1677: 1676:Sphaeriusidae 1674: 1672: 1669: 1667: 1661: 1660: 1657: 1653: 1647: 1643: 1629: 1623: 1621: 1615: 1613: 1610: 1603: 1601: 1598: 1596: 1590: 1588: 1582: 1580: 1574: 1572: 1568:Cicindelidae 1566: 1564: 1558: 1556: 1553: 1551: 1545: 1544: 1542: 1539: 1535: 1531: 1527: 1521: 1517: 1507: 1504: 1502: 1499: 1497: 1494: 1487: 1485: 1479: 1477: 1474: 1467: 1466: 1463: 1459: 1453: 1449: 1443: 1439: 1437: 1433: 1431: 1427: 1425: 1421: 1419: 1415: 1413: 1409: 1408: 1405: 1400: 1392: 1387: 1385: 1380: 1378: 1373: 1372: 1369: 1363: 1360: 1359: 1355: 1347: 1341: 1338: 1333: 1329: 1325: 1321: 1317: 1313: 1309: 1305: 1301: 1294: 1291: 1286: 1282: 1277: 1272: 1268: 1264: 1259: 1254: 1250: 1246: 1242: 1235: 1232: 1229: 1222: 1219: 1215: 1209: 1206: 1202: 1196: 1193: 1189: 1183: 1180: 1173: 1163: 1162: 1160: 1159: 1154: 1151: 1150: 1145: 1142: 1141: 1136: 1134: 1130: 1129:Burmese amber 1127:Legalov 2018 1126: 1125: 1120: 1119: 1117: 1116: 1114: 1107: 1106: 1101: 1098: 1095:Legalov 2010 1094: 1093: 1088: 1087: 1085: 1078: 1077: 1072: 1071: 1069: 1064: 1063: 1058: 1057: 1055: 1054: 1052: 1048: 1044: 1040: 1039: 1034: 1032: 1028: 1024: 1023: 1018: 1015: 1014: 1009: 1007: 1003: 1000:Legalov 2010 999: 998: 997:Cretoxenoides 993: 990: 989: 988:Brasilnemonyx 984: 983: 981: 980: 978: 971: 967: 966: 961: 960: 959: 956: 951: 950: 945: 942: 941: 936: 935: 933: 928: 927: 922: 919: 916:Legalov 2011 915: 914: 909: 906: 902: 901: 900:Microprobelus 896: 893: 889: 885: 884: 883:Oxycorynoides 879: 878: 876: 871: 870: 865: 864: 862: 857: 856: 851: 848: 847: 842: 839: 838: 833: 830: 829: 824: 823: 821: 820: 819:Arnoldi 1977 818: 814: 807: 803: 800: 796: 794: 790: 787:Legalov 2013 786: 785: 780: 779: 777: 773: 769: 768: 767:Turononemonyx 763: 760: 759: 758:Pseudonemonyx 754: 751: 747: 746: 741: 740: 738: 737: 735: 728: 727: 722: 719: 718: 717:Eccoptarthrus 713: 710: 707:Legalov 2014 706: 705: 700: 697: 696: 695:Astenorrhinus 691: 690: 688: 683: 679: 678: 673: 670: 669: 664: 661: 660: 655: 652: 648: 644: 640: 639: 634: 632: 628: 627: 626:Buryatnemonyx 622: 621: 619: 612: 611: 606: 605: 603: 599: 595: 591: 590: 585: 582: 581: 576: 574: 570: 566: 563:Legalov 2009 562: 561: 556: 555: 553: 552: 550: 549: 547: 545: 544: 543:Kuschelomacer 537: 536: 532: 527: 526: 522: 520: 519: 515: 513: 512: 508: 506: 505: 501: 499: 498: 494: 492: 491: 490:Rhinorhynchus 487: 485: 484: 480: 478: 477: 473: 471: 470: 466: 464: 463: 459: 457: 456: 452: 450: 449: 445: 443: 442: 438: 436: 435: 431: 429: 428: 427:Basiliorhinus 424: 422: 421: 417: 415: 414: 410: 408: 407: 403: 401: 400: 396: 395: 394: 392: 386: 385: 381: 380: 379: 377: 371: 370: 366: 364: 363: 359: 357: 356: 355:Doydirhynchus 352: 350: 349: 345: 343: 342: 338: 337: 336: 334: 326: 324: 322: 317: 313: 311: 307: 306:Araucariaceae 303: 302:Podocarpaceae 299: 295: 291: 287: 283: 282: 277: 272: 270: 266: 262: 258: 254: 250: 246: 242: 234: 225:Cimberididae 222: 219: 215: 210: 207: 205: 202: 199: 196: 195: 193: 188: 181: 178: 175: 174: 171: 168: 166:Superfamily: 165: 164: 161: 158: 155: 152: 151: 148: 145: 142: 141: 138: 135: 132: 131: 128: 125: 122: 121: 118: 115: 112: 111: 108: 105: 102: 101: 98: 95: 92: 91: 88: 85: 82: 81: 76: 71: 67: 64: 63: 58: 54: 49: 45: 41: 33: 30: 19: 3601:Nemonychidae 3356:Nemonychidae 3343:Nemonychidae 3329:Nemonychidae 3299:Nemonychidae 3298: 3177:Helophoridae 3169:Georissidae 3164:Epimetopidae 3080:Pleocomidae 3064:Ochodaeidae 3048:Hybosoridae 3032:Glaphyridae 3024:Geotrupidae 3002: 2999:Belohinidae 2960:Eucinetidae 2951: 2948:Decliniidae 2921: 2879:Phengodidae 2818:Cantharidae 2773:Dascillidae 2765:Dascilloidea 2747:Psephenidae 2739:Lutrochidae 2731:Limnichidae 2644:Buprestidae 2636:Buprestoidea 2576:Synchroidae 2560:Scraptiidae 2552:Salpingidae 2515:Prostomidae 2502:Oedemeridae 2478:Mordellidae 2396:Lymexylidae 2388:Lymexyloidea 2374: 2338:Attelabidae 2330:Anthribidae 2288:Smicripidae 2252:Phalacridae 2236:Nitidulidae 2231:Myraboliidae 2223:Monotomidae 2214: 2195:Kateretidae 2172:Cyclaxyridae 2132: 2129:Agapythidae 2093:Latridiidae 2043:Cerylonidae 2030:Anamorphidae 1964:Phycosecidae 1955: 1938: 1935:Metaxinidae 1888:Biphyllidae 1879: 1848:Orsodacnidae 1805:Cucujiformia 1738:Dermestidae 1608: 1547:Amphizoidae 1492: 1472: 1458:Archostemata 1442:Holometabola 1440:Superorder: 1434:Infraclass: 1348:, 41, 29–38. 1345: 1340: 1307: 1303: 1293: 1248: 1244: 1234: 1221: 1213: 1208: 1200: 1195: 1187: 1182: 1158:Renicimberis 1156: 1147: 1138: 1122: 1103: 1090: 1074: 1060: 1036: 1029:, Mongolia, 1020: 1013:Khetanamonyx 1011: 1004:, Mongolia, 995: 986: 963: 957: 947: 938: 924: 911: 898: 881: 867: 853: 844: 835: 826: 805: 798: 782: 765: 756: 745:Cretonemonyx 743: 724: 715: 704:Cratonemonyx 702: 693: 675: 666: 657: 636: 624: 608: 596:, Mongolia, 587: 578: 558: 541: 533:Extinct taxa 523: 516: 509: 502: 495: 488: 481: 474: 467: 460: 453: 446: 439: 432: 425: 418: 411: 404: 397: 389: 382: 376:Nemonychinae 374: 367: 360: 353: 346: 339: 333:Cimberidinae 331: 318: 314: 294:Cimberidinae 279: 275: 273: 259:. As in the 256: 248: 241:Nemonychidae 240: 239: 224: 204:Nemonychinae 198:Cimberidinae 190:Subfamilies 180:Nemonychidae 179: 153: 147:Cucujiformia 143:Infraorder: 60: 35:Nemonychidae 29: 18:Cimberididae 3465:iNaturalist 3323:Wikispecies 3221:Hydraenidae 3195:Spercheidae 3182:Hydrochidae 3146:Synteliidae 3130:Histeridae 3122:Histeroidea 3072:Passalidae 3040:Glaresidae 2897:Throscidae 2857:(fireflies) 2855:Lampyridae 2842:Eucnemidae 2834:Elateridae 2794:Elateroidea 2610:Zopheridae 2494:Mycteridae 2425:Anthicidae 2296:Sphindidae 2280:Silvanidae 2190:Hobartiidae 2177:Erotylidae 2106:Mycetaeidae 2101:Murmidiidae 2085:Euxestidae 1979:Rentoniidae 1853:Oxypeltidae 1838:Disteniidae 1671:Lepiceridae 1600:Hygrobiidae 1592:Haliplidae 1576:Dytiscidae 1555:Aspidytidae 1131:, Myanmar, 1076:Paleocartus 1031:Valanginian 1006:Hauterivian 968:Zhang 1997 965:Longidorsum 940:Belonotaris 815:†subfamily 726:Procurculio 677:Talbragarus 448:Eutactobius 420:Basiliogeus 406:Araucomacer 362:Lecontellus 327:Extant taxa 261:Anthribidae 230:Cimberidae 227:Gozis, 1882 184:Bedel, 1882 3585:Categories 3242:Silphidae 3234:Ptiliidae 3226:Leiodidae 3213:Agyrtidae 3056:Lucanidae 2935:Scirtoidea 2871:Omethidae 2850:Jurasaidae 2699:Dryopidae 2670:Byrrhidae 2662:Byrrhoidea 2354:Brentidae 2159:Cucujidae 2141:Boganiidae 2121:Cucujoidea 1994:Thymalidae 1984:Rhadalidae 1927:Melyridae 1896:Byturidae 1858:Vesperidae 1767:Ptiniidae 1617:Noteridae 1584:Gyrinidae 1560:Carabidae 1489:Jurodidae 1481:Cupedidae 1428:Subclass: 1418:Arthropoda 1399:Coleoptera 1304:GeoJournal 1174:References 1133:Cenomanian 1124:Burmomacer 1047:Berriasian 926:Cratomacer 888:Weald Clay 525:Zimmiellus 518:Stenomacer 469:Nannomacer 413:Atopomacer 399:Aragomacer 369:Pityomacer 281:Delphinium 160:Phytophaga 133:Suborder: 127:Coleoptera 107:Arthropoda 3096:Trogidae 3090:(scarabs) 2968:Scirtidae 2943:Clambidae 2536:Pythidae 2470:Meloidae 2417:Aderidae 2320:(weevils) 2185:Helotidae 2111:Teredidae 2025:Alexiidae 1909:Cleridae 1868:Cleroidea 1701:Polyphaga 1699:Suborder 1652:Myxophaga 1650:Suborder 1605:Meruidae 1524:Suborder 1506:Ommatidae 1456:Suborder 1430:Pterygota 1410:Kingdom: 1324:0343-2521 1267:0027-8424 869:Ampliceps 817:Eobelinae 651:Barremian 649:, Spain, 647:Las Hoyas 598:Tithonian 573:Oxfordian 569:Callovian 483:Pagomacer 476:Notomacer 462:Mecomacer 455:Idiomacer 341:Acromacer 137:Polyphaga 93:Kingdom: 87:Eukaryota 40:Oxfordian 3387:BugGuide 3363:BioLib: 3308:Wikidata 2863:Lycidae 2707:Elmidae 2605:Ulodidae 2441:Boridae 2346:Belidae 1947:Peltidae 1540:families 1526:Adephaga 1436:Neoptera 1416:Phylum: 1412:Animalia 1401:families 1332:83680109 1285:19365072 1188:Cimberis 1140:Burmonyx 949:Probelus 772:Turonian 441:Bunyaeus 348:Cimberis 253:antennae 218:Synonyms 176:Family: 103:Phylum: 97:Animalia 83:Domain: 3591:Weevils 3314:Q134893 2454:Ciidae 2362:Caridae 1424:Insecta 1422:Class: 1397:Extant 1276:2678426 846:Eobelus 384:Nemonyx 286:Pinales 276:Nemonyx 269:clypeus 245:weevils 123:Order: 117:Insecta 113:Class: 44:Present 3568:360133 3565:uBio: 3545:149636 3529:NZOR: 3496:616722 3483:100991 3470:174369 3379:211306 3366:243767 1538:Extant 1330:  1322:  1283:  1273:  1265:  793:Eocene 631:Aptian 434:Brarus 265:labrum 263:, the 3558:69422 3522:27443 3478:IRMNG 3431:11171 3405:8HBR3 3392:38594 1328:S2CID 1310:(6). 321:amber 154:Clade 3517:NCBI 3491:ITIS 3457:8584 3452:GBIF 3418:5233 3374:BOLD 1320:ISSN 1281:PMID 1263:ISSN 304:and 3504:NBN 3413:EoL 3400:CoL 3351:AFD 3338:ADW 1312:doi 1271:PMC 1253:doi 1249:106 3587:: 3555:: 3542:: 3519:: 3506:: 3493:: 3480:: 3467:: 3454:: 3441:: 3428:: 3415:: 3402:: 3389:: 3376:: 3353:: 3340:: 3325:: 3310:: 1326:. 1318:. 1306:. 1302:. 1279:. 1269:. 1261:. 1247:. 1243:. 804:†" 797:†" 645:, 393:: 378:: 335:: 323:. 292:. 156:: 3005:) 3001:( 2954:) 2950:( 2924:) 2920:( 2217:) 2213:( 2135:) 2131:( 1958:) 1954:( 1941:) 1937:( 1882:) 1878:( 1611:) 1607:( 1495:) 1491:( 1475:) 1471:( 1390:e 1383:t 1376:v 1334:. 1314:: 1308:7 1287:. 1255:: 1155:† 1146:† 1137:† 1121:† 1102:† 1089:† 1073:† 1059:† 1035:† 1019:† 1010:† 994:† 985:† 962:† 946:† 937:† 923:† 910:† 897:† 880:† 866:† 852:† 843:† 834:† 825:† 781:† 764:† 755:† 742:† 723:† 714:† 701:† 692:† 674:† 665:† 656:† 635:† 623:† 607:† 586:† 577:† 571:/ 557:† 539:† 42:- 20:)

Index

Cimberididae
Oxfordian
Present

Doydirhynchus austriacus
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Coleoptera
Polyphaga
Cucujiformia
Phytophaga
Curculionoidea
Nemonychidae
Cimberidinae
Nemonychinae
Rhinorhynchinae
Synonyms
weevils
antennae
Anthribidae
labrum
clypeus
Delphinium
Pinales
inflorescences
Cimberidinae

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