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Nipponopsalididae

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pronounced apophyses face outwards. Males also possess another apophysis on the second segment, which extends backwards and overlaps with the pronounced apophysis on the first segment. The penis in this species has a pair of soft bubble-like protuberances near the base of the glans, which are not found in other species. The glans is also longer than that of
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This species is absent from Urup Island in the central Kurils, and the populations from the Kurils north of Urup show clear morphological differences in the chelicerae and male pedipalps from the populations south of Urup into Hokkaido, suggesting that the Iturup Strait, between the islands of Iturup
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Nipponopsalididae belong to the superfamily Troguloidea. They have been regarded as a sister group to all remaining Troguloidea, though the most recent Opiliones phylogeny places them as the sister group to a clade consisting of Dicranolasmatidae and Trogulidae, with Nemastomatidae as sister to all
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The chelicerae of this species exhibit unique, conspicuous apophyses. Both male and female individuals possess apophyses on the first cheliceral segment, though they are considerably larger in males than in females. In males, the most pronounced apophysis faces inwards, but in females, the most
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This species exhibits unique male palpal morphology, with the tibia connected to the patella via a short stalk, and swollen basally; the tibia and tarsus are also densely covered in short hairs. Unlike the other species, male chelicerae in this species feature only small tubercles, rather than
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are longer than their body and heavily sclerotized. These chelicerae are superficially similar to those found in some Ischyropsalidioids, although this is a case of convergence. The cheliceral fingers exhibit 2 forms of teeth: narrow diaphanous teeth in the middle, and courser teeth distally.
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The penis shaft is long, slender, and gradually tapering, and the glans is three-branched. The two lateral branches of the glans appear as plates that are set with setae and shield the median branch, which houses the opening of the seminal duct.
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This subspecies is known from the island of Amami-oshima, one of the Satsunan Islands. It represents the southernmost distribution of the genus. This subspecies has distinctly narrower male cheliceral apophyses and considerably longer legs than
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with the first 5 opisthosomal tergites fused together. Further sexual dimorphisms include size, as females are larger than males, and the chelicerae, which are stouter and stronger in males, and which exhibit different sex-based armature.
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The male chelicerae in this species exhibit a unique, notable protuberance basally on the second segment. In addition, the penis shaft is shorter and broader than that of the other species, with a proportionally larger glans.
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This species was discovered in Akan National Park, in Hokkaido. Additional specimens have since been collected in the Kuril Islands of eastern Russia. This species represents the northernmost distribution of the genus.
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This subspecies is known from the three Japanese islands of Honshu, Kyoshu, and Shikoku. This subspecies has more swollen male cheliceral apophyses and considerably shorter legs than
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remaining Troguloidea. An internal phylogeny has not yet been conducted for this family. When originally described, the superficial morphological similarities between
944: 1962: 1975: 980:: International Symposium on Kuril Island Biodiversity, May 18th - 22nd, 2001 at Sapporo, Hokkaido Japan, The Hokkaido University Museum. Session 3 - Spiders. 913: 248:
is incomplete. The segmentation of the dorsum differs between males and females, as females generally are less heavily sclerotized than males, and exhibit a
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to be placed within that genus, though it is now known to be distinct, and morphological similarities between the two genera are a result of convergence.
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This species was discovered in Iwakuni, in the Yamagutchi prefecture in southern Japan. It was the first species of the genus to be described.
2021: 287:, by the complete absence of any spines on the second thoracic segment, though they are not known to be sympatric with any of those genera. 1136: 759: 232:
They range in body length from 2.3 to 4.1 mm. The carapace is domed, with a large, low ocularium, and rather large eyes. The
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Gorb, Stanislav N.; Taylor, Christopher K.; Wijnhoven, Hay; Martens, Jochen; Schönhofer, Axel L.; Wolff, Jonas O. (2016-07-01).
2016: 1980: 914:"Zur Systematischen Stellung Ostasiatischer Ischyropsalididen-Arten (Arachnoidea, Opiliones, Ischyropsalididae)" 1129: 720:
conspicuous armature. The penis shaft is proportionally longer than in other species, with a shorter glans.
1853: 866:"The evolution of pedipalps and glandular hairs as predatory devices in harvestmen (Arachnida, Opiliones)" 166: 1923: 1668: 689:
This species was discovered in Chungju, Korea, and was the first to be discovered outside of Japan.
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This family is very geographically conserved, and is known only from East Asia, primarily the
220:, meaning Japan, where the genus was first discovered, and the ending of the harvestman genus 1993: 1516: 1019: 1003: 877: 807: 1800: 1539: 1496: 1447: 1264: 1226: 990:
Fernández, Rosa; Sharma, Prashant P.; Tourinho, Ana Lúcia; Giribet, Gonzalo (2017-02-22).
308: 30: 1759: 1723: 1698: 1501: 1491: 1366: 1246: 1024: 991: 796:"A taxonomic catalogue of the Dyspnoi Hansen and Sørensen, 1904 (Arachnida: Opiliones)" 724:
and Urup, has served as an important geographical barrier separating these two groups.
2010: 1769: 1731: 1643: 1635: 1442: 1432: 1419: 1282: 1214: 1190: 795: 649:. Leg measurements (in millimeters) for males are : I 11, II 20, III 11, IV 15. 312: 224:, which comes from the Greek "psalis" (scissors), referring to the long chelicerae. 1915: 1795: 1777: 1713: 1703: 1688: 1683: 1678: 1615: 1529: 1483: 1474: 1437: 1427: 178: 158: 1094:"A remarkable new phalangid, Ischyropsalis yezoensis (Ischyropsalidae, Opiliones)" 1949: 1885: 1810: 1805: 1597: 1564: 1555: 1506: 1353: 1338: 1254: 667:. Leg measurements (in millimeters) for males are: I 22, II 40, III 23, IV 33. 300: 116: 1876: 1820: 1658: 1625: 1620: 1610: 1572: 1463: 1376: 1234: 812: 237: 195: 1015: 891: 1815: 1749: 1741: 1708: 1663: 1401: 1343: 1259: 1209: 1179: 1167: 1161: 1114: 304: 92: 68: 1902: 1033: 1007: 821: 975:
Diversity and Biogeography of Harvestmen (Opiliones) of the Kuril Islands.
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Sato et Suzuki (Opiliones) und seine weitere Verbreitung in Japan."
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Pinto-da-Rocha; Machado; Giribet, Ricardo; Glauco; Gonzalo (2007).
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They can be distinguished from other long-jawed Dyspnoi, like
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The opisthosoma is generally poorly sclerotized, and the
854:, Series B, Division 1 (Zoology), 7, 209–219 + pl. I. 1860: 1786: 1768: 1740: 1722: 1634: 1596: 1563: 1552: 1515: 1482: 1471: 1460: 1418: 1400: 1389: 1352: 1324: 1306: 1295: 1273: 1243: 1223: 1198: 1187: 1176: 952:Journal of Science of the Hiroshima University 945:"Opiliones from the South-west Islands, Japan" 852:Journal of Science of the Hiroshima University 1130: 8: 774:: CS1 maint: multiple names: authors list ( 236:and legs are very long and slender, and the 198:with three described species in one genus, 1848: 1560: 1479: 1468: 1397: 1303: 1195: 1184: 1173: 1137: 1123: 1115: 973:Tsurusaki, N. & Crawford, R.L. (2001) 912:Martens & Suzuki, J. & S. (1966). 29: 20: 1023: 881: 870:Zoological Journal of the Linnean Society 811: 303:, the more southern Japanese islands of 1065:"Four remarkable phalangids from Korea" 733: 619:, and the shaft is longer than that of 845:"Ăśber das Vorkommen des Männchens von 767: 1087: 1085: 581:Recovered by Fernández et al. (2017) 7: 969: 967: 965: 938: 936: 934: 907: 905: 903: 901: 839: 837: 789: 787: 785: 752:Harvestmen: The Biology of Opiliones 745: 743: 741: 739: 737: 1053:, Tokyo, 51, 523–528 (in Japanese). 519: 497: 477: 469: 459:Recovered by Giribet et al. (2002) 397: 377: 355: 347: 16:Family of harvestmen/daddy longlegs 1101:Annotationes Zoologicae Japonenses 1072:Annotationes Zoologicae Japonenses 921:Annotationes Zoologicae Japonenses 14: 204:, which is found in East Asia. 1: 2022:Monogeneric arachnid families 794:Schönhofer, Axel L. (2013). 754:. Harvard University Press. 654:Nipponopsalis abei longipes 299:. They are also known from 2038: 297:four main Japanese islands 152:Martens & Suzuki, 1966 1150: 1145:Extant Opiliones Families 813:10.11646/zootaxa.3679.1.1 631:(Sato & Suzuki, 1939) 597:(Sato & Suzuki, 1939) 537: 524: 517: 502: 495: 482: 475: 415: 402: 395: 382: 375: 360: 353: 268:In at least one species, 184: 177: 164: 157: 46:Scientific classification 44: 37: 28: 23: 708:Ischyropsalis yezoensis 698:Nipponopsalis yezoensis 641:Sato & Suzuki, 1939 628:Nipponopsalis abei abei 607:Sato & Suzuki, 1939 172:Sato & Suzuki, 1939 39:Nipponopsalis yezoensis 1008:10.1098/rspb.2016.2340 682:Ischyropsalis coreana 672:Nipponopsalis coreana 216:is a combination of 2017:Harvestman families 1092:Suzuki, S. (1958). 1063:Suzuki, S. (1966). 943:Suzuki, S. (1973). 843:Suzuki, S. (1940c) 185:1 genus, 3 species 1326:Ischyropsalidoidea 1308:Acropsopilionoidea 1051:Dobutsugaku Zasshi 1046:Suzuki, S. (1939) 1002:(1849): 20162340. 638:Ischyropsalis abei 604:Ischyropsalis abei 594:Nipponopsalis abei 526:Dicranolasmatidae 404:Dicranolasmatidae 311:, and most of the 167:Ischyropsalis abei 24:Nipponopsalididae 2004: 2003: 1989:Open Tree of Life 1854:Taxon identifiers 1845: 1844: 1841: 1840: 1837: 1836: 1833: 1832: 1829: 1828: 1548: 1547: 1535:Synthetonychiidae 1456: 1455: 1385: 1384: 1372:Nipponopsalididae 1362:Dicranolasmatidae 1334:Ischyropsalididae 1316:Acropsopilionidae 1291: 1290: 883:10.1111/zoj.12375 847:Ischropsalis abei 712: 702: 686: 676: 658: 642: 632: 608: 598: 585: 584: 575: 574: 566: 565: 557: 556: 548: 547: 505:Nipponopsalididae 463: 462: 453: 452: 444: 443: 435: 434: 426: 425: 363:Nipponopsalididae 192:Nipponopsalididae 189: 188: 129:Nipponopsalididae 2029: 1997: 1996: 1984: 1983: 1971: 1970: 1958: 1957: 1945: 1944: 1932: 1931: 1919: 1918: 1906: 1905: 1896: 1895: 1894: 1881: 1880: 1879: 1849: 1561: 1517:Triaenonychoidea 1480: 1469: 1398: 1304: 1196: 1185: 1174: 1139: 1132: 1125: 1116: 1109: 1108: 1098: 1089: 1080: 1079: 1069: 1060: 1054: 1044: 1038: 1037: 1027: 987: 981: 971: 960: 959: 949: 940: 929: 928: 918: 909: 896: 895: 885: 861: 855: 841: 832: 831: 829: 828: 815: 791: 780: 779: 773: 765: 747: 710: 700: 684: 674: 656: 647:N. abei longipes 640: 630: 606: 596: 520: 498: 478: 470: 465: 464: 398: 378: 356: 348: 343: 342: 254:scutum dissectum 250:scutum laminatum 33: 21: 2037: 2036: 2032: 2031: 2030: 2028: 2027: 2026: 2007: 2006: 2005: 2000: 1992: 1987: 1979: 1974: 1966: 1961: 1953: 1948: 1940: 1935: 1927: 1922: 1914: 1909: 1901: 1899: 1890: 1889: 1884: 1875: 1874: 1869: 1856: 1846: 1825: 1801:Fissiphalliidae 1782: 1764: 1736: 1718: 1669:Cryptogeobiidae 1630: 1592: 1554: 1544: 1540:Triaenonychidae 1511: 1497:Cryptomastridae 1473: 1462: 1452: 1448:Sclerosomatidae 1414: 1391: 1381: 1348: 1320: 1297: 1287: 1275: 1269: 1265:Troglosironidae 1245: 1239: 1227:Scopulophthalmi 1225: 1219: 1200: 1189: 1178: 1172: 1146: 1143: 1113: 1112: 1096: 1091: 1090: 1083: 1067: 1062: 1061: 1057: 1045: 1041: 996:Proc. R. Soc. B 989: 988: 984: 972: 963: 947: 942: 941: 932: 916: 911: 910: 899: 863: 862: 858: 842: 835: 826: 824: 793: 792: 783: 766: 762: 749: 748: 735: 730: 704: 678: 660: 634: 600: 590: 576: 567: 558: 549: 484:Nemastomatidae 454: 445: 436: 427: 384:Nemastomatidae 325: 293: 230: 212:The genus name 210: 194:is a family of 173: 170: 153: 150: 149: 135: 132: 131: 119: 107: 95: 83: 71: 59: 17: 12: 11: 5: 2035: 2033: 2025: 2024: 2019: 2009: 2008: 2002: 2001: 1999: 1998: 1985: 1972: 1959: 1946: 1933: 1920: 1907: 1897: 1882: 1866: 1864: 1858: 1857: 1852: 1843: 1842: 1839: 1838: 1835: 1834: 1831: 1830: 1827: 1826: 1824: 1823: 1818: 1813: 1808: 1803: 1798: 1792: 1790: 1784: 1783: 1781: 1780: 1774: 1772: 1766: 1765: 1763: 1762: 1760:Stygnommatidae 1757: 1752: 1746: 1744: 1738: 1737: 1735: 1734: 1728: 1726: 1724:Phalangodoidea 1720: 1719: 1717: 1716: 1711: 1706: 1701: 1699:Otilioleptidae 1696: 1691: 1686: 1681: 1676: 1671: 1666: 1661: 1656: 1651: 1646: 1640: 1638: 1632: 1631: 1629: 1628: 1623: 1618: 1613: 1608: 1602: 1600: 1594: 1593: 1591: 1590: 1588:Trionyxellidae 1585: 1580: 1575: 1569: 1567: 1558: 1550: 1549: 1546: 1545: 1543: 1542: 1537: 1532: 1527: 1521: 1519: 1513: 1512: 1510: 1509: 1504: 1502:Paranonychidae 1499: 1494: 1492:Cladonychiidae 1488: 1486: 1477: 1466: 1458: 1457: 1454: 1453: 1451: 1450: 1445: 1440: 1435: 1430: 1424: 1422: 1416: 1415: 1413: 1412: 1406: 1404: 1395: 1387: 1386: 1383: 1382: 1380: 1379: 1374: 1369: 1367:Nemastomatidae 1364: 1358: 1356: 1350: 1349: 1347: 1346: 1341: 1336: 1330: 1328: 1322: 1321: 1319: 1318: 1312: 1310: 1301: 1293: 1292: 1289: 1288: 1286: 1285: 1279: 1277: 1271: 1270: 1268: 1267: 1262: 1257: 1251: 1249: 1247:Sternophthalmi 1241: 1240: 1238: 1237: 1231: 1229: 1221: 1220: 1218: 1217: 1212: 1206: 1204: 1193: 1182: 1171: 1170: 1164: 1158: 1151: 1148: 1147: 1144: 1142: 1141: 1134: 1127: 1119: 1111: 1110: 1081: 1055: 1039: 982: 961: 930: 897: 876:(3): 558–601. 856: 833: 781: 760: 732: 731: 729: 726: 703: 701:(Suzuki, 1958) 695: 677: 675:(Suzuki, 1966) 669: 659: 651: 633: 625: 599: 591: 589: 586: 583: 582: 578: 577: 573: 572: 569: 568: 564: 563: 560: 559: 555: 554: 551: 550: 546: 545: 542: 541: 536: 533: 532: 529: 528: 523: 518: 516: 513: 512: 509: 508: 501: 496: 494: 491: 490: 487: 486: 481: 476: 474: 468: 461: 460: 456: 455: 451: 450: 447: 446: 442: 441: 438: 437: 433: 432: 429: 428: 424: 423: 420: 419: 414: 411: 410: 407: 406: 401: 396: 394: 391: 390: 387: 386: 381: 376: 374: 371: 370: 367: 366: 359: 354: 352: 346: 324: 321: 292: 289: 258:scutum parvum, 229: 226: 209: 206: 187: 186: 182: 181: 175: 174: 171: 162: 161: 155: 154: 151: 143: 141: 137: 136: 133: 127: 125: 121: 120: 115: 113: 109: 108: 103: 101: 97: 96: 91: 89: 85: 84: 79: 77: 73: 72: 67: 65: 61: 60: 55: 53: 49: 48: 42: 41: 35: 34: 26: 25: 15: 13: 10: 9: 6: 4: 3: 2: 2034: 2023: 2020: 2018: 2015: 2014: 2012: 1995: 1990: 1986: 1982: 1977: 1973: 1969: 1964: 1960: 1956: 1951: 1947: 1943: 1938: 1934: 1930: 1925: 1921: 1917: 1912: 1908: 1904: 1898: 1893: 1892:Nipponopsalis 1887: 1883: 1878: 1872: 1868: 1867: 1865: 1863: 1862:Nipponopsalis 1859: 1855: 1850: 1822: 1819: 1817: 1814: 1812: 1809: 1807: 1804: 1802: 1799: 1797: 1794: 1793: 1791: 1789: 1785: 1779: 1776: 1775: 1773: 1771: 1770:Sandokanoidea 1767: 1761: 1758: 1756: 1753: 1751: 1748: 1747: 1745: 1743: 1739: 1733: 1732:Phalangodidae 1730: 1729: 1727: 1725: 1721: 1715: 1712: 1710: 1707: 1705: 1702: 1700: 1697: 1695: 1694:Nomoclastidae 1692: 1690: 1687: 1685: 1682: 1680: 1677: 1675: 1672: 1670: 1667: 1665: 1662: 1660: 1657: 1655: 1652: 1650: 1647: 1645: 1644:Agoristenidae 1642: 1641: 1639: 1637: 1636:Gonyleptoidea 1633: 1627: 1624: 1622: 1619: 1617: 1614: 1612: 1609: 1607: 1604: 1603: 1601: 1599: 1595: 1589: 1586: 1584: 1581: 1579: 1578:Pyramidopidae 1576: 1574: 1571: 1570: 1568: 1566: 1562: 1559: 1557: 1551: 1541: 1538: 1536: 1533: 1531: 1528: 1526: 1525:Buemarinoidae 1523: 1522: 1520: 1518: 1514: 1508: 1505: 1503: 1500: 1498: 1495: 1493: 1490: 1489: 1487: 1485: 1481: 1478: 1476: 1470: 1467: 1465: 1459: 1449: 1446: 1444: 1443:Protolophidae 1441: 1439: 1436: 1434: 1433:Neopilionidae 1431: 1429: 1426: 1425: 1423: 1421: 1420:Phalangioidea 1417: 1411: 1408: 1407: 1405: 1403: 1399: 1396: 1394: 1388: 1378: 1375: 1373: 1370: 1368: 1365: 1363: 1360: 1359: 1357: 1355: 1351: 1345: 1342: 1340: 1337: 1335: 1332: 1331: 1329: 1327: 1323: 1317: 1314: 1313: 1311: 1309: 1305: 1302: 1300: 1294: 1284: 1283:Parasironidae 1281: 1280: 1278: 1272: 1266: 1263: 1261: 1258: 1256: 1253: 1252: 1250: 1248: 1242: 1236: 1233: 1232: 1230: 1228: 1222: 1216: 1215:Stylocellidae 1213: 1211: 1208: 1207: 1205: 1203: 1202:Boreophthalmi 1197: 1194: 1192: 1191:Cyphophthalmi 1186: 1183: 1181: 1175: 1169: 1165: 1163: 1159: 1157: 1153: 1152: 1149: 1140: 1135: 1133: 1128: 1126: 1121: 1120: 1117: 1106: 1102: 1095: 1088: 1086: 1082: 1077: 1073: 1066: 1059: 1056: 1052: 1049: 1043: 1040: 1035: 1031: 1026: 1021: 1017: 1013: 1009: 1005: 1001: 997: 993: 986: 983: 979: 976: 970: 968: 966: 962: 957: 953: 946: 939: 937: 935: 931: 926: 922: 915: 908: 906: 904: 902: 898: 893: 889: 884: 879: 875: 871: 867: 860: 857: 853: 850: 848: 840: 838: 834: 823: 819: 814: 809: 805: 801: 797: 790: 788: 786: 782: 777: 771: 763: 761:9780674023437 757: 753: 746: 744: 742: 740: 738: 734: 727: 725: 721: 717: 713: 709: 699: 696: 694: 690: 687: 683: 673: 670: 668: 666: 655: 652: 650: 648: 643: 639: 629: 626: 624: 622: 618: 612: 609: 605: 595: 592: 587: 580: 579: 571: 570: 562: 561: 553: 552: 544: 543: 540: 535: 534: 531: 530: 527: 522: 521: 515: 514: 511: 510: 507: 506: 500: 499: 493: 492: 489: 488: 485: 480: 479: 472: 471: 467: 466: 458: 457: 449: 448: 440: 439: 431: 430: 422: 421: 418: 413: 412: 409: 408: 405: 400: 399: 393: 392: 389: 388: 385: 380: 379: 373: 372: 369: 368: 365: 364: 358: 357: 350: 349: 345: 344: 341: 339: 338:Nipponopsalis 335: 334:Ischyropsalis 331: 330:Nipponopsalis 323:Relationships 322: 320: 318: 315:, except for 314: 313:Kuril Islands 310: 306: 302: 298: 290: 288: 286: 282: 278: 277:Ischyropsalis 273: 271: 266: 262: 259: 255: 251: 247: 246:corona analis 242: 239: 235: 227: 225: 223: 222:Ischyropsalis 219: 215: 214:Nipponopsalis 207: 205: 203: 202: 201:Nipponopsalis 197: 193: 183: 180: 176: 169: 168: 163: 160: 156: 148: 147: 146:Nipponopsalis 142: 139: 138: 134:Martens, 1976 130: 126: 123: 122: 118: 114: 112:Superfamily: 111: 110: 106: 102: 99: 98: 94: 90: 87: 86: 82: 78: 75: 74: 70: 66: 63: 62: 58: 54: 51: 50: 47: 43: 40: 36: 32: 27: 22: 19: 1861: 1796:Escadabiidae 1778:Sandokanidae 1714:Stygnopsidae 1704:Prostygnidae 1689:Metasarcidae 1684:Manaosbiidae 1679:Gonyleptidae 1654:Askawachidae 1616:Petrobunidae 1606:Beloniscidae 1530:Lomanellidae 1484:Travunioidea 1475:Insidiatores 1438:Phalangiidae 1428:Globipedidae 1371: 1104: 1100: 1075: 1071: 1058: 1050: 1042: 999: 995: 985: 977: 955: 951: 924: 920: 873: 869: 859: 851: 846: 825:. Retrieved 803: 799: 751: 722: 718: 714: 711:Suzuki, 1958 707: 705: 697: 691: 688: 685:Suzuki, 1966 681: 679: 671: 665:N. abei abei 664: 661: 657:Suzuki, 1973 653: 646: 644: 637: 635: 627: 620: 617:N. yezoensis 616: 613: 610: 603: 601: 593: 538: 525: 504: 503: 483: 473:Troguloidea 416: 403: 383: 362: 361: 351:Troguloidea 337: 333: 329: 326: 309:Amami ĹŚshima 294: 291:Distribution 284: 280: 276: 274: 269: 267: 263: 257: 253: 249: 245: 243: 231: 221: 217: 213: 211: 200: 199: 191: 190: 165: 159:Type species 145: 144: 128: 38: 18: 1950:iNaturalist 1886:Wikispecies 1811:Icaleptidae 1806:Guasiniidae 1788:Zalmoxoidea 1674:Gerdesiidae 1598:Epedanoidea 1583:Suthepiidae 1565:Assamioidea 1556:Grassatores 1507:Travuniidae 1354:Troguloidea 1339:Sabaconidae 1255:Neogoveidae 539:Trogulidae 417:Trogulidae 301:South Korea 228:Description 117:Troguloidea 2011:Categories 1821:Zalmoxidae 1659:Cosmetidae 1626:Tithaeidae 1621:Podoctidae 1611:Epedanidae 1573:Assamiidae 1553:Infraorder 1472:Infraorder 1464:Laniatores 1377:Trogulidae 1274:Infraorder 1244:Infraorder 1235:Pettalidae 1224:Infraorder 1199:Infraorder 1162:Arthropoda 1107:: 167–170. 958:: 205–279. 927:: 215–221. 827:2018-10-26 728:References 621:N. coreana 238:chelicerae 196:harvestmen 100:Suborder: 69:Arthropoda 1816:Kimulidae 1750:Biantidae 1742:Samooidea 1709:Stygnidae 1664:Cranaidae 1649:Ampycidae 1402:Caddoidea 1344:Taracidae 1260:Ogoveidae 1210:Sironidae 1180:Opiliones 1168:Arachnida 1154:Kingdom: 1078:: 95–106. 1016:0962-8452 892:0024-4082 770:cite book 706:Synonym: 680:Synonym: 636:Synonym: 602:Synonym: 305:Yakushima 234:pedipalps 179:Diversity 93:Opiliones 81:Arachnida 52:Kingdom: 1900:BioLib: 1877:Q3342114 1871:Wikidata 1755:Samoidae 1461:Suborder 1410:Caddidae 1390:Suborder 1296:Suborder 1276:(Indet.) 1188:Suborder 1160:Phylum: 1156:Animalia 1034:28228511 822:26146693 806:: 1–68. 124:Family: 64:Phylum: 57:Animalia 1968:1120561 1942:4661585 1929:9069564 1903:1044542 1299:Dyspnoi 1166:Class: 1025:5326524 800:Zootaxa 588:Species 285:Oskoron 281:Taracus 270:N. abei 140:Genus: 105:Dyspnoi 88:Order: 76:Class: 1994:235489 1981:109752 1955:525985 1393:Eupnoi 1177:Order 1032:  1022:  1014:  890:  820:  758:  283:, and 218:Nippon 1963:IRMNG 1916:7PQDC 1097:(PDF) 1068:(PDF) 948:(PDF) 917:(PDF) 1976:NCBI 1937:GBIF 1030:PMID 1012:ISSN 888:ISSN 818:PMID 804:3679 776:link 756:ISBN 336:led 332:and 317:Urup 307:and 208:Name 1924:EoL 1911:CoL 1020:PMC 1004:doi 1000:284 878:doi 874:177 808:doi 623:. 319:. 252:or 2013:: 1991:: 1978:: 1965:: 1952:: 1939:: 1926:: 1913:: 1888:: 1873:: 1105:31 1103:. 1099:. 1084:^ 1076:39 1074:. 1070:. 1028:. 1018:. 1010:. 998:. 994:. 978:In 964:^ 956:24 954:. 950:. 933:^ 925:39 923:. 919:. 900:^ 886:. 872:. 868:. 836:^ 816:. 802:. 798:. 784:^ 772:}} 768:{{ 736:^ 279:, 1138:e 1131:t 1124:v 1048:. 1036:. 1006:: 894:. 880:: 830:. 810:: 778:) 764:.

Index


Scientific classification
Animalia
Arthropoda
Arachnida
Opiliones
Dyspnoi
Troguloidea
Type species
Ischyropsalis abei
Diversity
harvestmen
pedipalps
chelicerae
four main Japanese islands
South Korea
Yakushima
Amami ĹŚshima
Kuril Islands
Urup





ISBN
9780674023437
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