565:, unique derived qualities. At the anterior back vertebra, the depression between the front edge of the side process and the lower edge of the side process is divided by two accessory ridges, one of which runs vertically while the other crosses the first horizontally. On the front side of the neural spines of the front tail vertebrae, run vertical ridges fusing at their lower ends, above the level of the front joint processes, creating a V-shaped structure. The upper inner corner of the humerus is strongly expanded, extending far beyond the inner side of the shaft. The humerus has in general a strongly expanded upper edge, 2.9 wider than the shaft diameter. On the humerus shaft, below the scar for the
2409:
2420:
2427:
3821:
476:, about eighty-six million years old. It consists of a partial skeleton lacking the skull. It contains an anterior back vertebra, an anterior tail vertebra, a right humerus and the upper part of a left pubic bone. Although the bones have not been discovered in articulation, they were considered to represent a single individual because they were found in close association on a surface of eight by eight metres. A second partial skeleton lacking the skull was referred to
149:
4553:
5788:
5782:
3826:
489:
547:
3283:
5777:
388:
3275:
5771:
4545:
2415:
3295:
3289:
4561:
571:, runs a diagonal ridge, from the upper and outer side to the inner and lower side. The first metatarsal has an upper width exceeding its length. The third metatarsal is relatively short with 1.2 times the length of the first metatarsal. The upper toe phalanges are 50% wider at their tops than their respective metatarsals are long. The toe claws are reduced, rough and truncated.
504:, and therefore one of the heaviest land animals, yet discovered. Although the incompleteness of the skeleton of the new sauropod has prevented scientists from making precise estimates of its size, its humerus, or upper arm bone, is 1.76 m (5.8 ft) in length, which is longer than that of any other titanosaur for which this bone is known, including other giants such as
936:
131:
480:: specimen UNCUYO-LD 302 representing a smaller individual. It contains a series of five anterior tail vertebrae and a complete right foot connected to an astragalus. It was recovered on a distance of 403 metres to the holotype from a surface of five by five metres.
526:
were comparable to those of better preserved titanosaurs, the new dinosaur may have weighed in the range of 44.9–75.9 tonnes (44.2–74.7 long tons; 49.5–83.7 short tons), most likely 60.4 tonnes (59.4 long tons; 66.6 short tons). In 2019
542:
which was estimated at 65-75 tonnes (71.6-82.7 short tons). This claim though, is considered unlikely. In 2020 Molina-Perez and
Larramendi estimated its length at 28 m (92 ft) and its weight at 40 tonnes (44 short tons).
1411:"Registro de un dinosaurio Aeolosaurini (Sauropoda, Titanosauria) en el Cretácico Superior (Formación Anacleto) del norte de la provincia de Neuquén, Argentina, y comentarios sobre aeolosaurinos sudamericanos"
5987:
590:, characteristics otherwise unknown in the Sauropoda. This parallels the vertical position of the metacarpals and the finger reduction in the hand of other titanosaurs.
1479:
1442:
1344:
1260:
522:. They also estimated the total femur length at 2.166 m (7.11 ft), by using an allometric equation. If, as is likely, the body proportions of
1511:
1179:
1313:
Leonardo S. Filippi; José Ignacio Canudo; Leonardo J. Salgado; Alberto C. Garrido; Rodolfo A. Garcia; Ignacio A. Cerda; Alejandro Otero (2011).
6007:
1223:"New Titanosauria (Dinosauria, Sauropoda) remains from the Upper Cretaceous (Plottier Fm) of the southern Neuquén Basin (Patagonia, Argentina)"
6027:
1053:"Determining the largest known land animal: A critical comparison of differing methods for restoring the volume and mass of extinct animals"
6022:
1172:"Osteology of the Late Cretaceous Argentinean sauropod dinosaur Mendozasaurus neguyelap: implications for basal titanosaur relationships"
535:
in the 45-55 tonne (49.6-60.6 short tons) range with the possibility that it might have been larger than that or approaching the mass of
6017:
6012:
1360:"Dinosaurs (Reptilia, Archosauria) at Museo de La Plata, Argentina: annotated catalogue of the type material and Antarctic specimens"
2419:
2408:
1122:
418:
400:
2426:
3820:
5997:
5992:
1504:
148:
4552:
582:, a skeleton part that is not often preserved in sauropods, shows a unique build. It contains a compact, homogeneous
986:
González Riga, Bernardo J.; Lamanna, Matthew C.; Ortiz David, Leonardo D.; Calvo, Jorge O.; Coria, Juan P. (2016).
5787:
5781:
6002:
567:
426:
5824:
3825:
1599:
1497:
5982:
5977:
5972:
450:
446:
3282:
5845:
3294:
3274:
5944:
5890:
5814:
5776:
5505:
1787:
1473:
1436:
1338:
1254:
1171:
631:
and found it to be outside of
Lithostrotia. In 2018, González Riga and colleagues found it to belong to
318:
5770:
4544:
3288:
4560:
2414:
5931:
5877:
1287:
1234:
1098:
999:
949:
921:
873:
295:
2490:
1315:"A new sauropod titanosaur from the Plottier Formation (Upper Cretaceous) of Patagonia (Argentina)"
442:
430:
5545:
2617:
1075:
857:
465:
143:
5949:
5895:
1276:"Cretaceous terrestrial beds from the Neuquén Basin (Argentina) and their tetrapod assemblages"
5681:
5222:
3969:
3610:
3339:
2527:
1391:
1149:
1033:
1015:
5552:
5397:
5337:
4979:
4871:
4777:
4668:
4470:
4450:
4389:
3508:
3361:
3208:
2925:
2879:
2859:
2803:
2106:
1883:
1863:
1422:
1381:
1371:
1326:
1295:
1242:
1196:
1188:
1139:
1131:
1067:
1023:
1007:
941:
861:
715:
512:
404:
372:
5819:
5689:
5422:
5315:
5011:
4991:
4972:
4930:
4857:
4843:
4726:
4477:
4397:
4044:
3580:
3571:
3432:
3424:
3351:
3111:
2989:
2810:
2666:
2497:
2226:
2219:
2211:
2198:
2115:
1661:
1648:
1611:
1584:
1559:
1520:
739:
537:
528:
222:
43:
1291:
1238:
1170:
Gonzalez Riga, B.J.; Mannion, P.D.; Poropat, S.F.; Ortiz David, L.; Coria, J.P. (2018).
1102:
1003:
5696:
5611:
5493:
5438:
5388:
5200:
5193:
5139:
5111:
5066:
5039:
4914:
4879:
4821:
4748:
4661:
4425:
4304:
4276:
4255:
4240:
4162:
4135:
3953:
3937:
3903:
3746:
3545:
3417:
3243:
2982:
2961:
2910:
2902:
2754:
2747:
2654:
2553:
2541:
2349:
2321:
2158:
2129:
2122:
2068:
1982:
1968:
1952:
1927:
1757:
1144:
1117:
1028:
987:
905:
898:
618:
562:
488:
546:
5966:
5739:
5724:
5703:
5619:
5579:
5567:
5537:
5486:
5458:
5431:
5352:
5329:
5277:
5208:
5132:
5098:
5087:
5025:
4944:
4907:
4886:
4836:
4814:
4806:
4799:
4792:
4733:
4698:
4675:
4617:
4484:
4463:
4331:
4322:
4311:
4290:
4269:
4247:
4190:
4067:
4004:
3888:
3711:
3687:
3653:
3624:
3617:
3596:
3531:
3524:
3517:
3482:
3473:
3410:
3402:
3395:
3325:
3312:
3229:
3153:
3096:
3075:
3024:
3017:
3010:
3003:
2996:
2953:
2932:
2851:
2828:
2675:
2632:
2534:
2511:
2483:
2314:
2307:
2255:
2151:
2088:
2079:
1961:
1835:
1803:
1771:
1732:
1714:
1693:
1458:"A new titanosaur sauropod from the Late Cretaceous of Neuquén, Patagonia, Argentina"
1457:
1410:
1314:
1275:
1222:
1079:
916:
691:
623:
586:, thought to be adapted for bearing extraordinary weight. It also presents truncated
575:
506:
81:
1118:"A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs"
988:"A gigantic new dinosaur from Argentina and the evolution of the sauropod hind foot"
5747:
5717:
5710:
5647:
5588:
5514:
5446:
5404:
5373:
5344:
5322:
5306:
5240:
5229:
5180:
5161:
5154:
5032:
5018:
5004:
4958:
4951:
4922:
4893:
4829:
4770:
4712:
4705:
4654:
4647:
4610:
4592:
4579:
4507:
4491:
4432:
4418:
4382:
4375:
4361:
4339:
4233:
4225:
4218:
4197:
4169:
4117:
4103:
4089:
4074:
4025:
3946:
3918:
3911:
3730:
3719:
3696:
3678:
3603:
3589:
3496:
3453:
3446:
3388:
3381:
3144:
3059:
2867:
2817:
2789:
2782:
2761:
2733:
2719:
2639:
2583:
2562:
2248:
2136:
2040:
1996:
1989:
1941:
1913:
1874:
1700:
1221:
Flavio
Bellardini, Mattia Antonio Baiano, Francisco Barrios, Borja Holgado (2018).
911:
797:
677:
667:
652:
647:
632:
628:
614:
422:
411:
270:
1299:
387:
5922:
5868:
920:
is also discovered in this formation, although it was originally assigned to the
864:. This formation is about 25 metres (82 ft) thick and composed of light red
5732:
5640:
5626:
5522:
5411:
5366:
5263:
5256:
5249:
5215:
5147:
5125:
5046:
4937:
4632:
4439:
4404:
4297:
4262:
4204:
4183:
4176:
4148:
4081:
4053:
4018:
4011:
3997:
3896:
3881:
3782:
3704:
3667:
3538:
3489:
3250:
3126:
3089:
3082:
3051:
3035:
2975:
2939:
2917:
2837:
2796:
2740:
2712:
2698:
2646:
2599:
2519:
2504:
2476:
2342:
2300:
2286:
2266:
2240:
2186:
2179:
2095:
2061:
2047:
1890:
1828:
1764:
1741:
1707:
1679:
1192:
889:
780:
641:
627:. A 2017 study by Carballido and colleagues recovered it as the sister taxon of
518:
56:
25:
453:, for having studied the dinosaur fossils of Mendoza province for two decades.
5633:
5560:
5529:
5291:
5118:
4965:
4900:
4864:
4850:
4785:
4756:
4719:
4682:
4625:
4211:
4155:
4110:
4060:
4032:
3990:
3926:
3874:
3856:
3843:
3767:
3760:
3753:
3739:
3646:
3639:
3632:
3552:
3464:
3439:
3169:
3119:
3103:
3067:
2968:
2844:
2768:
2682:
2625:
2576:
2328:
2144:
2033:
1897:
1856:
1821:
1550:
1246:
931:
583:
501:
368:
358:
254:
206:
101:
66:
20:
1395:
1019:
5596:
5478:
5284:
4763:
4741:
4690:
4411:
4368:
4354:
4096:
3789:
3775:
3660:
3374:
3222:
3215:
2946:
2895:
2458:
2445:
2360:
2335:
2293:
2172:
2012:
1779:
1721:
877:
869:
865:
659:
606:
473:
469:
376:
238:
160:
106:
50:
1201:
1153:
1135:
1052:
1037:
1427:
1330:
130:
5916:
5862:
5604:
5270:
5080:
4283:
4124:
3983:
3236:
2726:
2569:
2279:
2054:
2026:
1934:
1849:
1842:
1749:
1686:
1568:
1544:
1071:
881:
636:
457:
396:
364:
361:
193:
180:
96:
91:
76:
71:
61:
4640:
3044:
2888:
2775:
2233:
1386:
885:
111:
86:
1011:
5169:
4347:
2165:
1538:
587:
170:
5839:
884:
of the
Plottier formation are poorly known. They are represented by
433:
and Juan P. Coria. The generic name combines the Greek words νότος,
1095:
Dinosaur Facts and
Figures: The Sauropods and Other Sauropodomorphs
1489:
872:. The Plottier Formation can be differentiated from the overlying
602:
545:
487:
386:
354:
5936:
5882:
1376:
1359:
5471:
1975:
910:
and a specimen that appears to be an oldest member of the group
5843:
5803:
4577:
3841:
3310:
2443:
1646:
1531:
1493:
3133:
892:, and several titanosaurs. These titanosaurs, in addition to
605:
Titanosauria, a very diverse group of sauropods. In 2016, a
1409:
L. Filippi, Agustin G. Martinelli, Alberto
Garrido (2013).
981:
979:
977:
975:
973:
971:
969:
967:
965:
441:, "giant statue", in reference to the provenance from the
1456:
J. O. Calvo, B. J. González Riga, J. D. Porfiri (2007).
1097:. New Jersey: Princeton University Press. p. 267.
651:. This analysis also found Longkosauria to belong to
1358:
Reguero, Marcelo; Otero, Alejandro (March 8, 2013).
5906:
5852:
5673:
5578:
5504:
5457:
5421:
5387:
5305:
5239:
5179:
5097:
5064:
4990:
4590:
4506:
4449:
4321:
4134:
4043:
3967:
3936:
3854:
3729:
3686:
3677:
3579:
3570:
3507:
3472:
3463:
3360:
3349:
3323:
3200:
3168:
3152:
3143:
3034:
2878:
2827:
2696:
2665:
2597:
2552:
2456:
2359:
2265:
2197:
2105:
2078:
2010:
1951:
1911:
1873:
1801:
1731:
1659:
1610:
1583:
1165:
1163:
399:was discovered by the Argentine paleontologist Dr.
670:according to Gonzalez Riga and colleagues, 2018.
1216:
1214:
1212:
1505:
8:
1478:: CS1 maint: multiple names: authors list (
1441:: CS1 maint: multiple names: authors list (
1343:: CS1 maint: DOI inactive as of July 2024 (
1259:: CS1 maint: multiple names: authors list (
5988:Late Cretaceous dinosaurs of South America
5840:
5800:
5670:
5463:
5418:
5185:
5103:
5072:
4996:
4602:
4598:
4587:
4574:
4455:
4140:
3975:
3866:
3862:
3851:
3838:
3683:
3576:
3469:
3366:
3357:
3331:
3320:
3307:
3197:
3149:
2704:
2609:
2605:
2468:
2464:
2453:
2440:
2271:
2203:
2018:
1948:
1919:
1813:
1809:
1671:
1667:
1656:
1643:
1589:
1580:
1528:
1512:
1498:
1490:
1462:Arquivos do Museu Nacional, Rio de Janeiro
557:In 2016, several distinguishing traits of
129:
33:
19:Not to be confused with another Argentine
1426:
1385:
1375:
1200:
1143:
1116:Carballido, José L.; et al. (2017).
1027:
445:and the gigantic size of the animal. The
1274:Leanza, Héctor A.; et al. (2004).
961:
1471:
1434:
1336:
1252:
1093:Molina-Perez & Larramendi (2020).
7:
561:were determined. Nine of these were
1415:Revista Brasileira de Paleontologia
755:
731:
707:
683:
673:
876:by the former's higher content of
14:
5805:Topics in sauropodomorph research
5786:
5780:
5775:
5769:
4559:
4551:
4543:
3824:
3819:
3293:
3287:
3281:
3273:
2425:
2418:
2413:
2407:
934:
147:
1180:Journal of the Linnean Society
464:, was found in a layer of the
437:, "south wind", and κολοσσός,
1:
6008:Fossil taxa described in 2016
5908:Notocolossus gonzalezparejasi
1300:10.1016/j.cretres.2003.10.005
1060:Annals of the Carnegie Museum
552:Notocolossus gonzalesparejasi
419:Bernardo Javier González Riga
415:Notocolossus gonzalezparejasi
401:Bernardo Javier González Riga
331:Notocolossus gonzalezparejasi
6028:Taxa named by Jorge O. Calvo
6023:Taxa named by Rodolfo Coria
496:The evidence suggests that
427:Leonardo Daniel Ortiz David
417:was named and described by
6044:
6018:Taxa named by Matt Lamanna
6013:Reptiles described in 2016
1364:Palaeontologia Electronica
1227:Journal of Iberian Geology
18:
16:Extinct genus of dinosaurs
5810:
5799:
5767:
5669:
5466:
5188:
5106:
5075:
4999:
4605:
4601:
4586:
4573:
4540:
4458:
4143:
3978:
3869:
3865:
3850:
3837:
3817:
3369:
3334:
3319:
3306:
3270:
3196:
2707:
2612:
2608:
2471:
2467:
2452:
2439:
2405:
2274:
2206:
2021:
1922:
1816:
1812:
1674:
1670:
1655:
1642:
1592:
1579:
1527:
1247:10.1007/s41513-018-0047-5
1193:10.1093/zoolinnean/zlx103
1051:Paul, Gregory S. (2019).
794:
777:
760:
753:
736:
729:
712:
705:
688:
681:
568:musculus coracobrachialis
324:
317:
144:Scientific classification
142:
137:
128:
36:
23:discovered a year later,
5825:List of sauropod species
310:N. gonzalezparejasi
868:and thin bands of pink
1333:(inactive 2024-07-13).
1136:10.1098/rspb.2017.1219
856:was discovered in the
662:shows the position of
554:
531:estimated the mass of
500:was among the largest
493:
451:Jorge González Parejas
392:
5945:Paleobiology Database
5891:Paleobiology Database
5815:Sauropod neck posture
5506:Opisthocoelicaudiinae
1428:10.4072/rbp.2013.1.11
1331:10.1344/105.000001648
613:as a basal member of
549:
491:
390:
138:Skeletal restoration
5998:Cretaceous Argentina
5993:Fossils of Argentina
1072:10.2992/007.085.0403
950:2016 in paleontology
922:Portezuelo Formation
874:Portezuelo Formation
423:Matthew Carl Lamanna
395:A fossil of a large
383:Discovery and naming
5674:Dubious titanosaurs
2491:Chinshakiangosaurus
1292:2004CrRes..25...61L
1280:Cretaceous Research
1239:2018JIbG...44...75B
1103:2020dffs.book.....M
1004:2016NatSR...619165G
609:analysis recovered
443:Southern Hemisphere
431:Jorge Orlando Calvo
5546:Opisthocoelicaudia
2618:Archaeodontosaurus
1130:(1860): 20171219.
992:Scientific Reports
858:Plottier Formation
555:
550:Reconstruction of
494:
466:Plottier Formation
393:
5960:
5959:
5846:Taxon identifiers
5837:
5836:
5833:
5832:
5795:
5794:
5765:
5764:
5761:
5760:
5757:
5756:
5682:Bruhathkayosaurus
5665:
5664:
5661:
5660:
5657:
5656:
5383:
5382:
5301:
5300:
5223:Panamericansaurus
5060:
5059:
5056:
5055:
4569:
4568:
4538:
4537:
4534:
4533:
4530:
4529:
4526:
4525:
4502:
4501:
3970:Titanosauriformes
3963:
3962:
3833:
3832:
3815:
3814:
3811:
3810:
3807:
3806:
3803:
3802:
3799:
3798:
3611:Brachytrachelopan
3566:
3565:
3562:
3561:
3340:Haplocanthosaurus
3302:
3301:
3268:
3267:
3264:
3263:
3260:
3259:
3201:Dubious sauropods
3192:
3191:
3188:
3187:
3184:
3183:
2692:
2691:
2593:
2592:
2528:Protognathosaurus
2435:
2434:
2403:
2402:
2399:
2398:
2395:
2394:
2391:
2390:
2387:
2386:
2383:
2382:
2379:
2378:
2006:
2005:
1907:
1906:
1797:
1796:
1638:
1637:
1634:
1633:
1630:
1629:
1012:10.1038/srep19165
845:
844:
836:
835:
827:
826:
818:
817:
809:
808:
346:
345:
298:
6035:
5953:
5952:
5940:
5939:
5927:
5926:
5925:
5899:
5898:
5886:
5885:
5873:
5872:
5871:
5841:
5801:
5790:
5784:
5779:
5773:
5671:
5553:Pellegrinisaurus
5464:
5419:
5398:Bustingorrytitan
5338:Futalognkosaurus
5186:
5104:
5073:
4997:
4980:Diamantinasauria
4872:Pellegrinisaurus
4778:Jiangshanosaurus
4669:Baotianmansaurus
4603:
4599:
4588:
4575:
4564:
4563:
4556:
4555:
4548:
4547:
4471:Diamantinasaurus
4456:
4451:Diamantinasauria
4390:Jiangshanosaurus
4141:
3976:
3867:
3863:
3852:
3839:
3828:
3823:
3684:
3577:
3509:Rebbachisaurinae
3470:
3367:
3362:Rebbachisauridae
3358:
3332:
3321:
3308:
3297:
3291:
3286:
3285:
3278:
3277:
3209:Bothriospondylus
3198:
3150:
2926:Eomamenchisaurus
2880:Mamenchisauridae
2860:Lapparentosaurus
2804:Spinophorosaurus
2705:
2610:
2606:
2469:
2465:
2454:
2441:
2430:
2429:
2422:
2417:
2411:
2272:
2204:
2107:Massospondylidae
2019:
1949:
1920:
1884:Jaklapallisaurus
1864:Thecodontosaurus
1814:
1810:
1672:
1668:
1657:
1644:
1590:
1581:
1574:
1573:
1529:
1514:
1507:
1500:
1491:
1484:
1483:
1477:
1469:
1453:
1447:
1446:
1440:
1432:
1430:
1406:
1400:
1399:
1389:
1379:
1355:
1349:
1348:
1342:
1334:
1310:
1304:
1303:
1271:
1265:
1264:
1258:
1250:
1218:
1207:
1206:
1204:
1176:
1167:
1158:
1157:
1147:
1113:
1107:
1106:
1090:
1084:
1083:
1057:
1048:
1042:
1041:
1031:
983:
944:
942:Dinosaurs portal
939:
938:
937:
756:
732:
716:Futalognkosaurus
708:
684:
674:
513:Futalognkosaurus
468:dating from the
405:Mendoza province
373:Mendoza Province
333:
329:
289:
282:
269:
253:
237:
221:
152:
151:
133:
123:
53:
42:Temporal range:
34:
6043:
6042:
6038:
6037:
6036:
6034:
6033:
6032:
5963:
5962:
5961:
5956:
5948:
5943:
5935:
5930:
5921:
5920:
5915:
5902:
5894:
5889:
5881:
5876:
5867:
5866:
5861:
5848:
5838:
5829:
5820:Sauropod hiatus
5806:
5791:
5753:
5690:Campylodoniscus
5653:
5574:
5500:
5453:
5423:Nemegtosauridae
5417:
5379:
5316:Argentinosaurus
5297:
5235:
5175:
5093:
5069:
5052:
5012:Atsinganosaurus
4992:Lirainosaurinae
4986:
4973:Xianshanosaurus
4858:Normanniasaurus
4844:Narambuenatitan
4727:Dongyangosaurus
4595:
4582:
4565:
4558:
4550:
4542:
4522:
4498:
4478:Sarmientosaurus
4445:
4398:Phuwiangosaurus
4317:
4130:
4045:Brachiosauridae
4039:
3972:
3959:
3932:
3859:
3846:
3829:
3795:
3725:
3673:
3581:Dicraeosauridae
3572:Flagellicaudata
3558:
3503:
3459:
3433:Nopcsaspondylus
3425:Maraapunisaurus
3354:
3352:Diplodocimorpha
3345:
3328:
3315:
3298:
3280:
3272:
3256:
3180:
3164:
3139:
3112:Tehuelchesaurus
3030:
2990:Tienshanosaurus
2874:
2823:
2811:Tehuelchesaurus
2701:
2688:
2667:Vulcanodontidae
2661:
2602:
2589:
2548:
2498:Gongxianosaurus
2461:
2448:
2431:
2424:
2375:
2355:
2261:
2220:Jingshanosaurus
2212:Chuxiongosaurus
2199:Sauropodiformes
2193:
2116:Adeopapposaurus
2101:
2074:
2015:
2002:
1947:
1916:
1903:
1869:
1806:
1793:
1727:
1664:
1662:Sauropodomorpha
1651:
1649:Sauropodomorpha
1626:
1612:Sauropodomorpha
1606:
1600:Avemetatarsalia
1585:Avemetatarsalia
1575:
1560:Avemetatarsalia
1534:
1533:
1523:
1521:Sauropodomorpha
1518:
1488:
1487:
1470:
1455:
1454:
1450:
1433:
1408:
1407:
1403:
1357:
1356:
1352:
1335:
1312:
1311:
1307:
1273:
1272:
1268:
1251:
1220:
1219:
1210:
1174:
1169:
1168:
1161:
1123:Proc. R. Soc. B
1115:
1114:
1110:
1092:
1091:
1087:
1055:
1050:
1049:
1045:
985:
984:
963:
958:
940:
935:
933:
930:
851:
846:
837:
828:
819:
810:
740:Argentinosaurus
601:belongs to the
596:
538:Argentinosaurus
529:Gregory S. Paul
492:Vertebral bones
486:
385:
342:
335:
327:
326:
313:
288:
280:
267:
251:
235:
223:Sauropodomorpha
219:
146:
124:
122:
121:
120:
119:
114:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
48:
47:
44:Late Cretaceous
40:
30:
17:
12:
11:
5:
6041:
6039:
6031:
6030:
6025:
6020:
6015:
6010:
6005:
6000:
5995:
5990:
5985:
5983:Santonian life
5980:
5978:Coniacian life
5975:
5973:Lithostrotians
5965:
5964:
5958:
5957:
5955:
5954:
5941:
5928:
5912:
5910:
5904:
5903:
5901:
5900:
5887:
5874:
5858:
5856:
5850:
5849:
5844:
5835:
5834:
5831:
5830:
5828:
5827:
5822:
5817:
5811:
5808:
5807:
5804:
5797:
5796:
5793:
5792:
5768:
5766:
5763:
5762:
5759:
5758:
5755:
5754:
5752:
5751:
5744:
5736:
5729:
5721:
5714:
5707:
5700:
5697:Clasmodosaurus
5693:
5686:
5677:
5675:
5667:
5666:
5663:
5662:
5659:
5658:
5655:
5654:
5652:
5651:
5644:
5637:
5630:
5623:
5616:
5612:Maxakalisaurus
5608:
5601:
5593:
5584:
5582:
5576:
5575:
5573:
5572:
5564:
5557:
5549:
5542:
5534:
5526:
5519:
5510:
5508:
5502:
5501:
5499:
5498:
5494:Quaesitosaurus
5490:
5483:
5475:
5467:
5461:
5455:
5454:
5452:
5451:
5443:
5439:Quaesitosaurus
5435:
5427:
5425:
5416:
5415:
5408:
5401:
5393:
5391:
5389:Saltasauroidea
5385:
5384:
5381:
5380:
5378:
5377:
5370:
5363:
5356:
5349:
5341:
5334:
5326:
5319:
5311:
5309:
5303:
5302:
5299:
5298:
5296:
5295:
5288:
5281:
5274:
5267:
5260:
5253:
5245:
5243:
5237:
5236:
5234:
5233:
5226:
5219:
5212:
5205:
5201:Maxakalisaurus
5197:
5194:Adamantisaurus
5189:
5183:
5177:
5176:
5174:
5173:
5166:
5158:
5151:
5144:
5140:Epachthosaurus
5136:
5129:
5122:
5115:
5112:Antarctosaurus
5107:
5101:
5095:
5094:
5092:
5091:
5084:
5076:
5070:
5067:Eutitanosauria
5065:
5062:
5061:
5058:
5057:
5054:
5053:
5051:
5050:
5043:
5040:Mansourasaurus
5036:
5029:
5022:
5015:
5008:
5000:
4994:
4988:
4987:
4985:
4984:
4976:
4969:
4962:
4955:
4948:
4941:
4934:
4927:
4919:
4915:Tastavinsaurus
4911:
4904:
4897:
4890:
4883:
4880:Petrobrasaurus
4876:
4868:
4861:
4854:
4847:
4840:
4833:
4826:
4822:Malarguesaurus
4818:
4811:
4803:
4796:
4789:
4782:
4774:
4767:
4760:
4753:
4749:Epachthosaurus
4745:
4738:
4730:
4723:
4716:
4709:
4702:
4695:
4687:
4679:
4672:
4665:
4662:Austroposeidon
4658:
4651:
4644:
4637:
4629:
4622:
4614:
4606:
4596:
4591:
4584:
4583:
4578:
4571:
4570:
4567:
4566:
4541:
4539:
4536:
4535:
4532:
4531:
4528:
4527:
4524:
4523:
4521:
4520:
4519:
4518:
4512:
4510:
4504:
4503:
4500:
4499:
4497:
4496:
4488:
4481:
4474:
4467:
4459:
4453:
4447:
4446:
4444:
4443:
4436:
4429:
4426:Tangvayosaurus
4422:
4415:
4408:
4401:
4394:
4386:
4379:
4372:
4365:
4358:
4351:
4344:
4336:
4327:
4325:
4319:
4318:
4316:
4315:
4308:
4305:Tastavinsaurus
4301:
4294:
4287:
4280:
4277:Pukyongosaurus
4273:
4266:
4259:
4256:Liubangosaurus
4252:
4244:
4241:Liaoningotitan
4237:
4230:
4222:
4215:
4208:
4201:
4194:
4187:
4180:
4173:
4166:
4163:Astrophocaudia
4159:
4152:
4144:
4138:
4136:Somphospondyli
4132:
4131:
4129:
4128:
4121:
4114:
4107:
4100:
4093:
4086:
4078:
4071:
4064:
4057:
4049:
4047:
4041:
4040:
4038:
4037:
4029:
4022:
4015:
4008:
4001:
3994:
3987:
3979:
3973:
3968:
3965:
3964:
3961:
3960:
3958:
3957:
3954:Lourinhasaurus
3950:
3942:
3940:
3938:Camarasauridae
3934:
3933:
3931:
3930:
3923:
3915:
3908:
3904:Dashanpusaurus
3900:
3893:
3885:
3878:
3870:
3860:
3855:
3848:
3847:
3842:
3835:
3834:
3831:
3830:
3818:
3816:
3813:
3812:
3809:
3808:
3805:
3804:
3801:
3800:
3797:
3796:
3794:
3793:
3786:
3779:
3772:
3764:
3757:
3750:
3747:Dinheirosaurus
3743:
3735:
3733:
3727:
3726:
3724:
3723:
3716:
3708:
3701:
3692:
3690:
3681:
3675:
3674:
3672:
3671:
3664:
3657:
3650:
3643:
3636:
3629:
3621:
3614:
3607:
3600:
3593:
3585:
3583:
3574:
3568:
3567:
3564:
3563:
3560:
3559:
3557:
3556:
3549:
3546:Rebbachisaurus
3542:
3535:
3528:
3521:
3513:
3511:
3505:
3504:
3502:
3501:
3493:
3486:
3478:
3476:
3467:
3461:
3460:
3458:
3457:
3450:
3443:
3436:
3429:
3421:
3418:Lavocatisaurus
3414:
3407:
3399:
3392:
3385:
3378:
3370:
3364:
3355:
3350:
3347:
3346:
3344:
3343:
3335:
3329:
3324:
3317:
3316:
3311:
3304:
3303:
3300:
3299:
3271:
3269:
3266:
3265:
3262:
3261:
3258:
3257:
3255:
3254:
3247:
3244:Qinlingosaurus
3240:
3233:
3226:
3219:
3212:
3204:
3202:
3194:
3193:
3190:
3189:
3186:
3185:
3182:
3181:
3179:
3178:
3177:(see below ↓ )
3174:
3172:
3166:
3165:
3163:
3162:
3161:(see below ↓ )
3158:
3156:
3147:
3141:
3140:
3138:
3137:
3130:
3123:
3116:
3108:
3100:
3093:
3086:
3079:
3072:
3064:
3056:
3048:
3040:
3038:
3032:
3031:
3029:
3028:
3021:
3014:
3007:
3000:
2993:
2986:
2983:Rhomaleopakhus
2979:
2972:
2965:
2962:Mamenchisaurus
2958:
2950:
2943:
2936:
2929:
2922:
2914:
2911:Chuanjiesaurus
2907:
2903:Cetiosauriscus
2899:
2892:
2884:
2882:
2876:
2875:
2873:
2872:
2864:
2856:
2848:
2841:
2833:
2831:
2825:
2824:
2822:
2821:
2814:
2807:
2800:
2793:
2786:
2779:
2772:
2765:
2758:
2755:Dashanpusaurus
2751:
2748:Cetiosauriscus
2744:
2737:
2730:
2723:
2716:
2708:
2702:
2697:
2694:
2693:
2690:
2689:
2687:
2686:
2679:
2671:
2669:
2663:
2662:
2660:
2659:
2655:Zizhongosaurus
2651:
2643:
2636:
2629:
2622:
2613:
2603:
2598:
2595:
2594:
2591:
2590:
2588:
2587:
2580:
2573:
2566:
2558:
2556:
2554:Lessemsauridae
2550:
2549:
2547:
2546:
2542:Tuebingosaurus
2538:
2531:
2524:
2516:
2508:
2501:
2494:
2487:
2480:
2472:
2462:
2457:
2450:
2449:
2444:
2437:
2436:
2433:
2432:
2406:
2404:
2401:
2400:
2397:
2396:
2393:
2392:
2389:
2388:
2385:
2384:
2381:
2380:
2377:
2376:
2374:
2373:
2372:
2371:
2365:
2363:
2357:
2356:
2354:
2353:
2350:Sefapanosaurus
2346:
2339:
2332:
2325:
2322:Melanorosaurus
2318:
2311:
2304:
2297:
2290:
2283:
2275:
2269:
2263:
2262:
2260:
2259:
2252:
2245:
2237:
2230:
2223:
2216:
2207:
2201:
2195:
2194:
2192:
2191:
2183:
2176:
2169:
2162:
2159:Massospondylus
2155:
2148:
2141:
2133:
2130:Glacialisaurus
2126:
2123:Coloradisaurus
2119:
2111:
2109:
2103:
2102:
2100:
2099:
2092:
2084:
2082:
2076:
2075:
2073:
2072:
2069:Tuebingosaurus
2065:
2058:
2051:
2044:
2037:
2030:
2022:
2016:
2011:
2008:
2007:
2004:
2003:
2001:
2000:
1993:
1986:
1983:Pachysauriscus
1979:
1972:
1969:Gresslyosaurus
1965:
1957:
1955:
1953:Plateosauridae
1946:
1945:
1938:
1931:
1928:Plateosauravus
1923:
1917:
1912:
1909:
1908:
1905:
1904:
1902:
1901:
1894:
1887:
1879:
1877:
1871:
1870:
1868:
1867:
1860:
1853:
1846:
1839:
1832:
1825:
1817:
1807:
1802:
1799:
1798:
1795:
1794:
1792:
1791:
1784:
1776:
1768:
1761:
1758:Chromogisaurus
1754:
1746:
1737:
1735:
1729:
1728:
1726:
1725:
1718:
1711:
1704:
1697:
1690:
1683:
1675:
1665:
1660:
1653:
1652:
1647:
1640:
1639:
1636:
1635:
1632:
1631:
1628:
1627:
1625:
1624:
1623:
1622:
1616:
1614:
1608:
1607:
1605:
1604:
1603:
1602:
1593:
1587:
1577:
1576:
1572:
1571:
1562:
1553:
1547:
1541:
1532:
1525:
1524:
1519:
1517:
1516:
1509:
1502:
1494:
1486:
1485:
1448:
1421:(1): 147–156.
1401:
1350:
1319:Geologica Acta
1305:
1266:
1208:
1187:(1): 136–181.
1159:
1108:
1085:
1066:(4): 335–358.
1043:
960:
959:
957:
954:
953:
952:
946:
945:
929:
926:
906:Antarctosaurus
899:Petrobrasaurus
850:
847:
843:
842:
839:
838:
834:
833:
830:
829:
825:
824:
821:
820:
816:
815:
812:
811:
807:
806:
803:
802:
793:
790:
789:
786:
785:
776:
773:
772:
769:
768:
759:
754:
752:
749:
748:
745:
744:
735:
730:
728:
725:
724:
721:
720:
711:
706:
704:
701:
700:
697:
696:
687:
682:
680:
672:
658:The following
619:sister species
595:
592:
563:autapomorphies
485:
482:
384:
381:
344:
343:
337:González Riga
336:
322:
321:
315:
314:
306:
304:
300:
299:
290:González Riga
278:
274:
273:
265:
258:
257:
249:
242:
241:
233:
226:
225:
217:
210:
209:
204:
197:
196:
191:
184:
183:
178:
174:
173:
168:
164:
163:
158:
154:
153:
140:
139:
135:
134:
126:
125:
117:
116:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
54:
41:
15:
13:
10:
9:
6:
4:
3:
2:
6040:
6029:
6026:
6024:
6021:
6019:
6016:
6014:
6011:
6009:
6006:
6004:
6003:Neuquén Basin
6001:
5999:
5996:
5994:
5991:
5989:
5986:
5984:
5981:
5979:
5976:
5974:
5971:
5970:
5968:
5951:
5946:
5942:
5938:
5933:
5929:
5924:
5918:
5914:
5913:
5911:
5909:
5905:
5897:
5892:
5888:
5884:
5879:
5875:
5870:
5864:
5860:
5859:
5857:
5855:
5851:
5847:
5842:
5826:
5823:
5821:
5818:
5816:
5813:
5812:
5809:
5802:
5798:
5789:
5785:
5783:
5778:
5772:
5750:
5749:
5745:
5742:
5741:
5740:Mongolosaurus
5737:
5735:
5734:
5730:
5727:
5726:
5725:Macrurosaurus
5722:
5720:
5719:
5715:
5713:
5712:
5708:
5706:
5705:
5704:Hypselosaurus
5701:
5699:
5698:
5694:
5692:
5691:
5687:
5684:
5683:
5679:
5678:
5676:
5672:
5668:
5650:
5649:
5645:
5643:
5642:
5638:
5636:
5635:
5631:
5629:
5628:
5624:
5622:
5621:
5620:Neuquensaurus
5617:
5614:
5613:
5609:
5607:
5606:
5602:
5599:
5598:
5594:
5591:
5590:
5586:
5585:
5583:
5581:
5580:Saltasaurinae
5577:
5570:
5569:
5568:Zhuchengtitan
5565:
5563:
5562:
5558:
5555:
5554:
5550:
5548:
5547:
5543:
5540:
5539:
5538:Dreadnoughtus
5535:
5532:
5531:
5527:
5525:
5524:
5520:
5517:
5516:
5512:
5511:
5509:
5507:
5503:
5496:
5495:
5491:
5489:
5488:
5487:Qingxiusaurus
5484:
5481:
5480:
5476:
5474:
5473:
5469:
5468:
5465:
5462:
5460:
5459:Saltasauridae
5456:
5449:
5448:
5444:
5441:
5440:
5436:
5434:
5433:
5432:Nemegtosaurus
5429:
5428:
5426:
5424:
5420:
5414:
5413:
5409:
5407:
5406:
5402:
5400:
5399:
5395:
5394:
5392:
5390:
5386:
5376:
5375:
5371:
5369:
5368:
5364:
5362:
5361:
5357:
5355:
5354:
5353:Mendozasaurus
5350:
5347:
5346:
5342:
5340:
5339:
5335:
5332:
5331:
5330:Dzharatitanis
5327:
5325:
5324:
5320:
5318:
5317:
5313:
5312:
5310:
5308:
5304:
5294:
5293:
5289:
5287:
5286:
5282:
5280:
5279:
5278:Gondwanatitan
5275:
5273:
5272:
5268:
5266:
5265:
5261:
5259:
5258:
5254:
5252:
5251:
5247:
5246:
5244:
5242:
5238:
5232:
5231:
5227:
5225:
5224:
5220:
5218:
5217:
5213:
5211:
5210:
5209:Muyelensaurus
5206:
5203:
5202:
5198:
5196:
5195:
5191:
5190:
5187:
5184:
5182:
5178:
5172:
5171:
5167:
5164:
5163:
5159:
5157:
5156:
5152:
5150:
5149:
5145:
5142:
5141:
5137:
5135:
5134:
5133:Chucarosaurus
5130:
5128:
5127:
5123:
5121:
5120:
5116:
5114:
5113:
5109:
5108:
5105:
5102:
5100:
5099:Colossosauria
5096:
5090:
5089:
5088:Menucocelsior
5085:
5083:
5082:
5078:
5077:
5074:
5071:
5068:
5063:
5049:
5048:
5044:
5042:
5041:
5037:
5035:
5034:
5030:
5028:
5027:
5026:Lirainosaurus
5023:
5021:
5020:
5016:
5014:
5013:
5009:
5007:
5006:
5002:
5001:
4998:
4995:
4993:
4989:
4982:
4981:
4977:
4975:
4974:
4970:
4968:
4967:
4963:
4961:
4960:
4956:
4954:
4953:
4949:
4947:
4946:
4945:Trigonosaurus
4942:
4940:
4939:
4935:
4933:
4932:
4928:
4925:
4924:
4920:
4917:
4916:
4912:
4910:
4909:
4908:Ruyangosaurus
4905:
4903:
4902:
4898:
4896:
4895:
4891:
4889:
4888:
4887:Pitekunsaurus
4884:
4882:
4881:
4877:
4874:
4873:
4869:
4867:
4866:
4862:
4860:
4859:
4855:
4853:
4852:
4848:
4846:
4845:
4841:
4839:
4838:
4837:Mnyamawamtuka
4834:
4832:
4831:
4827:
4824:
4823:
4819:
4817:
4816:
4815:Magyarosaurus
4812:
4809:
4808:
4807:Ligabuesaurus
4804:
4802:
4801:
4800:Laplatasaurus
4797:
4795:
4794:
4793:Karongasaurus
4790:
4788:
4787:
4783:
4780:
4779:
4775:
4773:
4772:
4768:
4766:
4765:
4761:
4759:
4758:
4754:
4751:
4750:
4746:
4744:
4743:
4739:
4736:
4735:
4734:Dreadnoughtus
4731:
4729:
4728:
4724:
4722:
4721:
4717:
4715:
4714:
4710:
4708:
4707:
4703:
4701:
4700:
4699:Borealosaurus
4696:
4693:
4692:
4688:
4685:
4684:
4680:
4678:
4677:
4676:Barrosasaurus
4673:
4671:
4670:
4666:
4664:
4663:
4659:
4657:
4656:
4652:
4650:
4649:
4645:
4643:
4642:
4638:
4635:
4634:
4630:
4628:
4627:
4623:
4620:
4619:
4618:Aegyptosaurus
4615:
4613:
4612:
4608:
4607:
4604:
4600:
4597:
4594:
4589:
4585:
4581:
4576:
4572:
4562:
4557:
4554:
4546:
4516:
4515:
4514:
4513:
4511:
4509:
4505:
4494:
4493:
4489:
4487:
4486:
4485:Savannasaurus
4482:
4480:
4479:
4475:
4473:
4472:
4468:
4466:
4465:
4464:Australotitan
4461:
4460:
4457:
4454:
4452:
4448:
4442:
4441:
4437:
4435:
4434:
4430:
4428:
4427:
4423:
4421:
4420:
4416:
4414:
4413:
4409:
4407:
4406:
4402:
4400:
4399:
4395:
4392:
4391:
4387:
4385:
4384:
4380:
4378:
4377:
4373:
4371:
4370:
4366:
4364:
4363:
4359:
4357:
4356:
4352:
4350:
4349:
4345:
4342:
4341:
4337:
4334:
4333:
4332:Australodocus
4329:
4328:
4326:
4324:
4323:Euhelopodidae
4320:
4314:
4313:
4312:Triunfosaurus
4309:
4307:
4306:
4302:
4300:
4299:
4295:
4293:
4292:
4291:Sauroposeidon
4288:
4286:
4285:
4281:
4279:
4278:
4274:
4272:
4271:
4270:Padillasaurus
4267:
4265:
4264:
4260:
4258:
4257:
4253:
4250:
4249:
4248:Ligabuesaurus
4245:
4243:
4242:
4238:
4236:
4235:
4231:
4228:
4227:
4223:
4221:
4220:
4216:
4214:
4213:
4209:
4207:
4206:
4202:
4200:
4199:
4195:
4193:
4192:
4191:Chubutisaurus
4188:
4186:
4185:
4181:
4179:
4178:
4174:
4172:
4171:
4167:
4165:
4164:
4160:
4158:
4157:
4153:
4151:
4150:
4146:
4145:
4142:
4139:
4137:
4133:
4127:
4126:
4122:
4120:
4119:
4115:
4113:
4112:
4108:
4106:
4105:
4101:
4099:
4098:
4094:
4092:
4091:
4087:
4084:
4083:
4079:
4077:
4076:
4072:
4070:
4069:
4068:Brachiosaurus
4065:
4063:
4062:
4058:
4056:
4055:
4051:
4050:
4048:
4046:
4042:
4035:
4034:
4030:
4028:
4027:
4023:
4021:
4020:
4016:
4014:
4013:
4009:
4007:
4006:
4005:Fushanosaurus
4002:
4000:
3999:
3995:
3993:
3992:
3988:
3986:
3985:
3981:
3980:
3977:
3974:
3971:
3966:
3956:
3955:
3951:
3949:
3948:
3944:
3943:
3941:
3939:
3935:
3929:
3928:
3924:
3921:
3920:
3916:
3914:
3913:
3909:
3906:
3905:
3901:
3899:
3898:
3894:
3891:
3890:
3889:Bashunosaurus
3886:
3884:
3883:
3879:
3877:
3876:
3872:
3871:
3868:
3864:
3861:
3858:
3853:
3849:
3845:
3840:
3836:
3827:
3822:
3792:
3791:
3787:
3785:
3784:
3780:
3778:
3777:
3773:
3770:
3769:
3765:
3763:
3762:
3758:
3756:
3755:
3751:
3749:
3748:
3744:
3742:
3741:
3737:
3736:
3734:
3732:
3728:
3722:
3721:
3717:
3714:
3713:
3712:Atlantosaurus
3709:
3707:
3706:
3702:
3699:
3698:
3694:
3693:
3691:
3689:
3688:Apatosaurinae
3685:
3682:
3680:
3676:
3670:
3669:
3665:
3663:
3662:
3658:
3656:
3655:
3654:Smitanosaurus
3651:
3649:
3648:
3644:
3642:
3641:
3637:
3635:
3634:
3630:
3627:
3626:
3625:Dyslocosaurus
3622:
3620:
3619:
3618:Dicraeosaurus
3615:
3613:
3612:
3608:
3606:
3605:
3601:
3599:
3598:
3597:Amargatitanis
3594:
3592:
3591:
3587:
3586:
3584:
3582:
3578:
3575:
3573:
3569:
3555:
3554:
3550:
3548:
3547:
3543:
3541:
3540:
3536:
3534:
3533:
3532:Katepensaurus
3529:
3527:
3526:
3525:Itapeuasaurus
3522:
3520:
3519:
3518:Demandasaurus
3515:
3514:
3512:
3510:
3506:
3499:
3498:
3494:
3492:
3491:
3487:
3485:
3484:
3483:Cathartesaura
3480:
3479:
3477:
3475:
3474:Limaysaurinae
3471:
3468:
3466:
3462:
3456:
3455:
3451:
3449:
3448:
3444:
3442:
3441:
3437:
3435:
3434:
3430:
3427:
3426:
3422:
3420:
3419:
3415:
3413:
3412:
3411:Histriasaurus
3408:
3405:
3404:
3403:Dzharatitanis
3400:
3398:
3397:
3396:Comahuesaurus
3393:
3391:
3390:
3386:
3384:
3383:
3379:
3377:
3376:
3372:
3371:
3368:
3365:
3363:
3359:
3356:
3353:
3348:
3342:
3341:
3337:
3336:
3333:
3330:
3327:
3326:Diplodocoidea
3322:
3318:
3314:
3313:Diplodocoidea
3309:
3305:
3296:
3292:
3290:
3284:
3276:
3253:
3252:
3248:
3246:
3245:
3241:
3239:
3238:
3234:
3232:
3231:
3230:Gigantosaurus
3227:
3225:
3224:
3220:
3218:
3217:
3213:
3211:
3210:
3206:
3205:
3203:
3199:
3195:
3176:
3175:
3173:
3171:
3167:
3160:
3159:
3157:
3155:
3154:Diplodocoidea
3151:
3148:
3146:
3142:
3136:
3135:
3131:
3129:
3128:
3124:
3122:
3121:
3117:
3114:
3113:
3109:
3106:
3105:
3101:
3099:
3098:
3097:Narindasaurus
3094:
3092:
3091:
3087:
3085:
3084:
3080:
3078:
3077:
3076:Losillasaurus
3073:
3070:
3069:
3065:
3062:
3061:
3057:
3054:
3053:
3049:
3047:
3046:
3042:
3041:
3039:
3037:
3033:
3027:
3026:
3025:Zigongosaurus
3022:
3020:
3019:
3018:Yuanmousaurus
3015:
3013:
3012:
3011:Xinjiangtitan
3008:
3006:
3005:
3004:Wamweracaudia
3001:
2999:
2998:
2997:Tonganosaurus
2994:
2992:
2991:
2987:
2985:
2984:
2980:
2978:
2977:
2973:
2971:
2970:
2966:
2964:
2963:
2959:
2956:
2955:
2954:Klamelisaurus
2951:
2949:
2948:
2944:
2942:
2941:
2937:
2935:
2934:
2933:Huangshanlong
2930:
2928:
2927:
2923:
2920:
2919:
2915:
2913:
2912:
2908:
2905:
2904:
2900:
2898:
2897:
2893:
2891:
2890:
2886:
2885:
2883:
2881:
2877:
2870:
2869:
2865:
2862:
2861:
2857:
2854:
2853:
2852:Ferganasaurus
2849:
2847:
2846:
2842:
2840:
2839:
2835:
2834:
2832:
2830:
2829:Cetiosauridae
2826:
2820:
2819:
2815:
2813:
2812:
2808:
2806:
2805:
2801:
2799:
2798:
2794:
2792:
2791:
2787:
2785:
2784:
2780:
2778:
2777:
2773:
2771:
2770:
2766:
2764:
2763:
2759:
2757:
2756:
2752:
2750:
2749:
2745:
2743:
2742:
2738:
2736:
2735:
2731:
2729:
2728:
2724:
2722:
2721:
2717:
2715:
2714:
2710:
2709:
2706:
2703:
2700:
2695:
2685:
2684:
2680:
2678:
2677:
2676:Tazoudasaurus
2673:
2672:
2670:
2668:
2664:
2657:
2656:
2652:
2649:
2648:
2644:
2642:
2641:
2637:
2635:
2634:
2633:Ohmdenosaurus
2630:
2628:
2627:
2623:
2620:
2619:
2615:
2614:
2611:
2607:
2604:
2601:
2596:
2586:
2585:
2581:
2579:
2578:
2574:
2572:
2571:
2567:
2565:
2564:
2560:
2559:
2557:
2555:
2551:
2544:
2543:
2539:
2537:
2536:
2535:Schleitheimia
2532:
2530:
2529:
2525:
2522:
2521:
2517:
2514:
2513:
2512:Leonerasaurus
2509:
2507:
2506:
2502:
2500:
2499:
2495:
2493:
2492:
2488:
2486:
2485:
2484:Blikanasaurus
2481:
2479:
2478:
2474:
2473:
2470:
2466:
2463:
2460:
2455:
2451:
2447:
2442:
2438:
2428:
2423:
2421:
2416:
2410:
2369:
2368:
2367:
2366:
2364:
2362:
2358:
2352:
2351:
2347:
2345:
2344:
2340:
2338:
2337:
2333:
2331:
2330:
2326:
2324:
2323:
2319:
2317:
2316:
2315:Leonerasaurus
2312:
2310:
2309:
2308:Lamplughsaura
2305:
2303:
2302:
2298:
2296:
2295:
2291:
2289:
2288:
2284:
2282:
2281:
2277:
2276:
2273:
2270:
2268:
2264:
2258:
2257:
2256:Yunnanosaurus
2253:
2251:
2250:
2246:
2243:
2242:
2238:
2236:
2235:
2231:
2229:
2228:
2224:
2222:
2221:
2217:
2214:
2213:
2209:
2208:
2205:
2202:
2200:
2196:
2189:
2188:
2184:
2182:
2181:
2177:
2175:
2174:
2170:
2168:
2167:
2163:
2161:
2160:
2156:
2154:
2153:
2152:Lufengosaurus
2149:
2147:
2146:
2142:
2139:
2138:
2134:
2132:
2131:
2127:
2125:
2124:
2120:
2118:
2117:
2113:
2112:
2110:
2108:
2104:
2098:
2097:
2093:
2091:
2090:
2089:Eucnemesaurus
2086:
2085:
2083:
2081:
2080:Riojasauridae
2077:
2071:
2070:
2066:
2064:
2063:
2059:
2057:
2056:
2052:
2050:
2049:
2045:
2043:
2042:
2038:
2036:
2035:
2031:
2029:
2028:
2024:
2023:
2020:
2017:
2014:
2009:
1999:
1998:
1994:
1992:
1991:
1987:
1985:
1984:
1980:
1978:
1977:
1973:
1971:
1970:
1966:
1964:
1963:
1962:Euskelosaurus
1959:
1958:
1956:
1954:
1950:
1944:
1943:
1939:
1937:
1936:
1932:
1930:
1929:
1925:
1924:
1921:
1918:
1915:
1910:
1900:
1899:
1895:
1893:
1892:
1888:
1886:
1885:
1881:
1880:
1878:
1876:
1872:
1866:
1865:
1861:
1859:
1858:
1854:
1852:
1851:
1847:
1845:
1844:
1840:
1838:
1837:
1836:Bagualosaurus
1833:
1831:
1830:
1826:
1824:
1823:
1819:
1818:
1815:
1811:
1808:
1805:
1804:Bagualosauria
1800:
1790:
1789:
1785:
1782:
1781:
1777:
1774:
1773:
1772:Pampadromaeus
1769:
1767:
1766:
1762:
1760:
1759:
1755:
1752:
1751:
1747:
1744:
1743:
1739:
1738:
1736:
1734:
1733:Saturnaliidae
1730:
1724:
1723:
1719:
1717:
1716:
1715:Pampadromaeus
1712:
1710:
1709:
1705:
1703:
1702:
1698:
1696:
1695:
1694:Gigantoscelus
1691:
1689:
1688:
1684:
1682:
1681:
1677:
1676:
1673:
1669:
1666:
1663:
1658:
1654:
1650:
1645:
1641:
1620:
1619:
1618:
1617:
1615:
1613:
1609:
1601:
1597:
1596:
1595:
1594:
1591:
1588:
1586:
1582:
1578:
1570:
1566:
1563:
1561:
1557:
1554:
1552:
1548:
1546:
1542:
1540:
1536:
1535:
1530:
1526:
1522:
1515:
1510:
1508:
1503:
1501:
1496:
1495:
1492:
1481:
1475:
1468:(4): 485–504.
1467:
1463:
1459:
1452:
1449:
1444:
1438:
1429:
1424:
1420:
1416:
1412:
1405:
1402:
1397:
1393:
1388:
1383:
1378:
1373:
1369:
1365:
1361:
1354:
1351:
1346:
1340:
1332:
1328:
1324:
1320:
1316:
1309:
1306:
1301:
1297:
1293:
1289:
1285:
1281:
1277:
1270:
1267:
1262:
1256:
1248:
1244:
1240:
1236:
1232:
1228:
1224:
1217:
1215:
1213:
1209:
1203:
1202:10044/1/53967
1198:
1194:
1190:
1186:
1182:
1181:
1173:
1166:
1164:
1160:
1155:
1151:
1146:
1141:
1137:
1133:
1129:
1125:
1124:
1119:
1112:
1109:
1104:
1100:
1096:
1089:
1086:
1081:
1077:
1073:
1069:
1065:
1061:
1054:
1047:
1044:
1039:
1035:
1030:
1025:
1021:
1017:
1013:
1009:
1005:
1001:
997:
993:
989:
982:
980:
978:
976:
974:
972:
970:
968:
966:
962:
955:
951:
948:
947:
943:
932:
927:
925:
923:
919:
918:
917:Muyelensaurus
913:
909:
907:
901:
900:
895:
891:
887:
883:
879:
875:
871:
867:
863:
862:Neuquén Group
859:
855:
848:
841:
840:
832:
831:
823:
822:
814:
813:
805:
804:
801:
800:
799:
792:
791:
788:
787:
784:
783:
782:
775:
774:
771:
770:
767:
766:
765:
758:
757:
751:
750:
747:
746:
743:
742:
741:
734:
733:
727:
726:
723:
722:
719:
718:
717:
710:
709:
703:
702:
699:
698:
695:
694:
693:
692:Mendozasaurus
686:
685:
679:
676:
675:
671:
669:
665:
661:
656:
654:
650:
649:
644:
643:
638:
634:
630:
626:
625:
624:Dreadnoughtus
620:
616:
612:
608:
604:
600:
593:
591:
589:
585:
581:
577:
572:
570:
569:
564:
560:
553:
548:
544:
541:
539:
534:
530:
525:
521:
520:
515:
514:
509:
508:
507:Dreadnoughtus
503:
499:
490:
483:
481:
479:
475:
471:
467:
463:
462:UNCUYO-LD 301
459:
454:
452:
448:
447:specific name
444:
440:
436:
432:
428:
424:
420:
416:
413:
410:In 2016, the
408:
406:
402:
398:
389:
382:
380:
378:
374:
370:
366:
363:
360:
359:titanosaurian
356:
352:
351:
340:
334:
332:
323:
320:
319:Binomial name
316:
312:
311:
305:
302:
301:
297:
293:
287:
286:
279:
276:
275:
272:
266:
263:
260:
259:
256:
250:
247:
244:
243:
240:
234:
231:
228:
227:
224:
218:
215:
212:
211:
208:
205:
202:
199:
198:
195:
192:
189:
186:
185:
182:
179:
176:
175:
172:
169:
166:
165:
162:
159:
156:
155:
150:
145:
141:
136:
132:
127:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
52:
45:
39:
35:
32:
28:
27:
22:
5907:
5854:Notocolossus
5853:
5774:
5748:Titanosaurus
5746:
5738:
5731:
5723:
5718:Loricosaurus
5716:
5711:Iuticosaurus
5709:
5702:
5695:
5688:
5680:
5648:Yamanasaurus
5646:
5639:
5632:
5625:
5618:
5610:
5603:
5595:
5589:Abditosaurus
5587:
5566:
5559:
5551:
5544:
5536:
5528:
5521:
5515:Abditosaurus
5513:
5492:
5485:
5477:
5470:
5447:Tapuiasaurus
5445:
5437:
5430:
5410:
5405:Titanomachya
5403:
5396:
5374:Puertasaurus
5372:
5365:
5360:Notocolossus
5359:
5358:
5351:
5345:Jiangxititan
5343:
5336:
5328:
5323:Drusilasaura
5321:
5314:
5307:Lognkosauria
5290:
5283:
5276:
5269:
5262:
5255:
5248:
5241:Aeolosaurini
5230:Rinconsaurus
5228:
5221:
5214:
5207:
5199:
5192:
5181:Rinconsauria
5168:
5162:Tengrisaurus
5160:
5155:Quetecsaurus
5153:
5146:
5138:
5131:
5124:
5117:
5110:
5086:
5079:
5045:
5038:
5033:Lohuecotitan
5031:
5024:
5019:Garrigatitan
5017:
5010:
5005:Ampelosaurus
5003:
4978:
4971:
4964:
4959:Uberabatitan
4957:
4952:Sonidosaurus
4950:
4943:
4936:
4929:
4923:Tengrisaurus
4921:
4913:
4906:
4899:
4894:Rapetosaurus
4892:
4885:
4878:
4870:
4863:
4856:
4849:
4842:
4835:
4830:Malawisaurus
4828:
4820:
4813:
4805:
4798:
4791:
4784:
4776:
4771:Huabeisaurus
4769:
4762:
4755:
4747:
4740:
4732:
4725:
4718:
4713:Choconsaurus
4711:
4706:Brasilotitan
4704:
4697:
4689:
4681:
4674:
4667:
4660:
4655:Atacamatitan
4653:
4648:Argyrosaurus
4646:
4639:
4631:
4624:
4616:
4611:Abdarainurus
4609:
4593:Titanosauria
4580:Titanosauria
4549:
4508:Titanosauria
4492:Wintonotitan
4490:
4483:
4476:
4469:
4462:
4438:
4433:Yongjinglong
4431:
4424:
4419:Tambatitanis
4417:
4410:
4403:
4396:
4388:
4383:Huanghetitan
4381:
4376:Huabeisaurus
4374:
4367:
4362:Gannansaurus
4360:
4353:
4346:
4340:Chiayusaurus
4338:
4330:
4310:
4303:
4296:
4289:
4282:
4275:
4268:
4261:
4254:
4246:
4239:
4234:Jiutaisaurus
4232:
4226:Jiangxititan
4224:
4219:Garumbatitan
4217:
4210:
4203:
4198:Dongbeititan
4196:
4189:
4182:
4175:
4170:Austrosaurus
4168:
4161:
4154:
4147:
4123:
4118:Venenosaurus
4116:
4109:
4104:Sonorasaurus
4102:
4095:
4090:Giraffatitan
4088:
4080:
4075:Cedarosaurus
4073:
4066:
4059:
4052:
4031:
4026:Pelorosaurus
4024:
4017:
4010:
4003:
3996:
3989:
3982:
3952:
3947:Camarasaurus
3945:
3925:
3919:Haestasaurus
3917:
3912:Europasaurus
3910:
3902:
3895:
3887:
3880:
3873:
3788:
3781:
3774:
3766:
3759:
3752:
3745:
3738:
3731:Diplodocinae
3720:Brontosaurus
3718:
3710:
3703:
3697:Amphicoelias
3695:
3679:Diplodocidae
3666:
3659:
3652:
3645:
3638:
3631:
3623:
3616:
3609:
3604:Bajadasaurus
3602:
3595:
3590:Amargasaurus
3588:
3551:
3544:
3537:
3530:
3523:
3516:
3497:Rayososaurus
3495:
3488:
3481:
3454:Zapalasaurus
3452:
3447:Xenoposeidon
3445:
3438:
3431:
3423:
3416:
3409:
3401:
3394:
3389:Campananeyen
3387:
3382:Amazonsaurus
3380:
3373:
3338:
3279:
3249:
3242:
3235:
3228:
3221:
3214:
3207:
3145:Neosauropoda
3132:
3125:
3118:
3110:
3102:
3095:
3088:
3081:
3074:
3066:
3060:Haestasaurus
3058:
3050:
3043:
3023:
3016:
3009:
3002:
2995:
2988:
2981:
2974:
2967:
2960:
2952:
2945:
2938:
2931:
2924:
2916:
2909:
2901:
2894:
2887:
2868:Patagosaurus
2866:
2858:
2850:
2843:
2836:
2818:Volkheimeria
2816:
2809:
2802:
2795:
2790:Perijasaurus
2788:
2783:Nebulasaurus
2781:
2774:
2767:
2762:Dystrophaeus
2760:
2753:
2746:
2739:
2734:Barapasaurus
2732:
2725:
2720:Asiatosaurus
2718:
2711:
2681:
2674:
2653:
2645:
2640:Rhoetosaurus
2638:
2631:
2624:
2616:
2584:Lessemsaurus
2582:
2575:
2568:
2563:Antetonitrus
2561:
2540:
2533:
2526:
2518:
2510:
2503:
2496:
2489:
2482:
2475:
2412:
2348:
2341:
2334:
2327:
2320:
2313:
2306:
2299:
2292:
2285:
2278:
2254:
2249:Yizhousaurus
2247:
2239:
2232:
2225:
2218:
2210:
2185:
2178:
2171:
2164:
2157:
2150:
2143:
2137:Ignavusaurus
2135:
2128:
2121:
2114:
2094:
2087:
2067:
2060:
2053:
2046:
2041:Ignavusaurus
2039:
2032:
2025:
1997:Yimenosaurus
1995:
1990:Plateosaurus
1988:
1981:
1974:
1967:
1960:
1942:Xixiposaurus
1940:
1933:
1926:
1914:Plateosauria
1896:
1889:
1882:
1875:Unaysauridae
1862:
1855:
1848:
1841:
1834:
1827:
1820:
1786:
1778:
1770:
1763:
1756:
1748:
1740:
1720:
1713:
1706:
1701:Guaibasaurus
1699:
1692:
1685:
1678:
1564:
1555:
1474:cite journal
1465:
1461:
1451:
1437:cite journal
1418:
1414:
1404:
1377:10.26879/352
1367:
1363:
1353:
1339:cite journal
1322:
1318:
1308:
1286:(1): 61–87.
1283:
1279:
1269:
1255:cite journal
1233:(1): 75–84.
1230:
1226:
1184:
1178:
1127:
1121:
1111:
1094:
1088:
1063:
1059:
1046:
995:
991:
915:
912:Aeolosaurini
903:
897:
894:Notocolossus
893:
854:Notocolossus
853:
852:
849:Paleoecology
798:Puertasaurus
796:
795:
779:
778:
764:Notocolossus
763:
762:
761:
738:
737:
714:
713:
690:
689:
678:Lognkosauria
668:Lognkosauria
664:Notocolossus
663:
657:
653:Lithostrotia
648:Puertasaurus
646:
640:
633:Lognkosauria
629:Lognkosauria
622:
615:Lithostrotia
611:Notocolossus
610:
599:Notocolossus
598:
597:
580:Notocolossus
579:
574:The foot or
573:
566:
559:Notocolossus
558:
556:
551:
536:
533:Notocolossus
532:
524:Notocolossus
523:
517:
511:
505:
498:Notocolossus
497:
495:
478:Notocolossus
477:
461:
455:
438:
434:
414:
412:type species
409:
394:
350:Notocolossus
349:
348:
347:
338:
330:
325:
309:
308:
291:
285:Notocolossus
284:
283:
271:Titanosauria
261:
245:
229:
213:
200:
187:
38:Notocolossus
37:
31:
24:
5733:Microcoelus
5641:Saltasaurus
5627:Paralititan
5523:Alamosaurus
5412:Udelartitan
5367:Patagotitan
5264:Bravasaurus
5257:Arrudatitan
5250:Aeolosaurus
5216:Overosaurus
5148:Jainosaurus
5126:Bonitasaura
5047:Paludititan
4938:Traukutitan
4633:Angolatitan
4440:Yunmenglong
4405:Qiaowanlong
4298:Sibirotitan
4263:Oceanotitan
4205:Europatitan
4184:Brontomerus
4177:Brohisaurus
4149:Angolatitan
4082:Galvesaurus
4054:Abydosaurus
4019:Ornithopsis
4012:Fusuisaurus
3998:Eucamerotus
3897:Daanosaurus
3882:Aragosaurus
3783:Supersaurus
3705:Apatosaurus
3668:Tharosaurus
3539:Nigersaurus
3490:Limaysaurus
3251:Ultrasaurus
3127:Turiasaurus
3090:Moabosaurus
3083:Mierasaurus
3052:Bellusaurus
3036:Turiasauria
2976:Qijianglong
2940:Hudiesaurus
2918:Datousaurus
2838:Cetiosaurus
2797:Shunosaurus
2741:Bellusaurus
2713:Algoasaurus
2699:Eusauropoda
2647:Sanpasaurus
2600:Gravisauria
2520:Pulanesaura
2505:Isanosaurus
2477:Amygdalodon
2343:Pulanesaura
2301:Irisosaurus
2287:Anchisaurus
2267:Anchisauria
2241:Xingxiulong
2187:Xingxiulong
2180:Sarahsaurus
2096:Riojasaurus
2062:Sarahsaurus
2048:Kholumolumo
1891:Macrocollum
1829:Asylosaurus
1765:Nhandumirim
1742:Buriolestes
1708:Mbiresaurus
1680:Buriolestes
1387:11336/13790
1370:(1): 1–24.
1325:(1): 1–12.
890:coelurosaur
781:Patagotitan
642:Patagotitan
519:Paralititan
502:titanosaurs
484:Description
26:Patagotitan
5967:Categories
5634:Rocasaurus
5561:Qunkasaura
5530:Baurutitan
5292:Shingopana
5119:Baalsaurus
4966:Volgatitan
4901:Rukwatitan
4865:Nullotitan
4851:Ninjatitan
4786:Kaijutitan
4757:Gandititan
4720:Daxiatitan
4683:Baurutitan
4626:Andesaurus
4517:see below↓
4212:Fukuititan
4156:Arkharavia
4111:Soriatitan
4061:Atlasaurus
4033:Rugocaudia
3991:Duriatitan
3927:Yuzhoulong
3875:Abrosaurus
3857:Macronaria
3844:Macronaria
3768:Kaatedocus
3761:Galeamopus
3754:Diplodocus
3740:Barosaurus
3647:Pilmatueia
3640:Lingwulong
3633:Kaatedocus
3553:Tataouinea
3465:Khebbashia
3440:Sidersaura
3170:Macronaria
3120:Tendaguria
3104:Oplosaurus
3068:Janenschia
2969:Omeisaurus
2845:Chebsaurus
2769:Janenschia
2683:Vulcanodon
2626:Kotasaurus
2577:Ledumahadi
2370:see below↓
2329:Meroktenos
2145:Leyesaurus
2034:Gyposaurus
1898:Unaysaurus
1857:Pantydraco
1822:Arcusaurus
1788:Saturnalia
1621:see below↓
1569:Dinosauria
1551:Sauropsida
956:References
896:, include
888:, a large
866:claystones
584:metatarsus
391:Limb bones
371:strata of
369:Cretaceous
367:from late
255:Macronaria
207:Saurischia
194:Dinosauria
21:titanosaur
5923:Q22226944
5869:Q23768183
5597:Bonatitan
5479:Isisaurus
5285:Punatitan
4764:Hamititan
4742:Elaltitan
4691:Bonatitan
4412:Silutitan
4369:Gobititan
4355:Euhelopus
4097:Lusotitan
3790:Tornieria
3776:Leinkupal
3661:Suuwassea
3375:Agustinia
3223:Dinodocus
3216:Cardiodon
2947:Jingiella
2896:Anhuilong
2459:Sauropoda
2446:Sauropoda
2361:Sauropoda
2336:Mussaurus
2294:Camelotia
2173:Pradhania
2013:Massopoda
1780:Panphagia
1722:Panphagia
1537:Kingdom:
1396:1094-8074
1080:210840060
1020:2045-2322
998:: 19165.
882:Tetrapods
878:argillite
870:sandstone
660:cladogram
607:cladistic
594:Phylogeny
474:Santonian
470:Coniacian
377:Argentina
303:Species:
239:Sauropoda
167:Kingdom:
161:Eukaryota
5937:47458367
5917:Wikidata
5883:47458366
5863:Wikidata
5605:Ibirania
5271:Caieiria
5081:Inawentu
4284:Ruixinia
4125:Vouivria
3984:Astrodon
3237:Neosodon
2727:Bagualia
2570:Ingentia
2280:Aardonyx
2227:Qianlong
2055:Musankwa
2027:Gryponyx
1935:Ruehleia
1850:Nambalia
1843:Efraasia
1750:Eoraptor
1687:Eoraptor
1545:Chordata
1543:Phylum:
1539:Animalia
1154:28794222
1038:26777391
928:See also
637:polytomy
458:holotype
449:honours
439:kolossos
397:sauropod
365:dinosaur
362:sauropod
181:Chordata
177:Phylum:
171:Animalia
157:Domain:
49:86
4641:Arackar
3045:Amanzia
2889:Analong
2776:Jobaria
2234:Seitaad
1549:Class:
1288:Bibcode
1235:Bibcode
1145:5563814
1099:Bibcode
1029:4725985
1000:Bibcode
886:mammals
860:of the
635:, in a
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