1764:
122:
1754:
1749:
1759:
285:
in 1984 on the basis of GNM CO 1–90, a partial skeleton including the incomplete skull, mandible, 43 vertebrae and both forelimbs (humeri, radii, ulnae, a few carpals, metacarpals, and phalanges) with scapulae. The type specimen was found in
Otradnaya, in the northwestern Russian Caucasus Mchedlidze
343:
in having a very short ascending process of the maxilla, a short lateral process of the maxilla, an anterior end of nasal located anterior to the rostrum base, and a supraorbital process of the frontal bone directed perpendicular to the anteroposterior axis of the skull. Differences from
294:, at the time a wastebasket for baleen-bearing mysticetes that did belong to any crown-group mysticete group (followed by McKenna and Bell 1997). Later recognition of Cetotheriidae sensu McKenna and Bell (1997) as paraphyletic/polyphyletic rendered the systematic position of
382:
Riabinin AI. 1934. New materials on the osteology of
Cetotherium mayeri Brandt from the Upper Sarmatian of the Northern Caucasus. Trudy Vsesoyuznogo Geologorazvedochnogo Ob’edineniya SSSR 350:1-15
391:
Gol’din P, Steeman ME (2015) From
Problem Taxa to Problem Solver: A New Miocene Family, Tranatocetidae, Brings Perspective on Baleen Whale Evolution. PLoS ONE 10(9): e0135500.
437:
348:
include a supraorbital process of the frontal bone not elongated at the lateral end and a robust medially bent coronoid process of the mandible.
1863:
1763:
1858:
1853:
1848:
430:
364:
Mchedlidze GA. 1984. A Fossil whale from the
Miocene deposits of environs of Otradnaya. Metsniereba. Tbilisi: Metsniereba .
337:
is a medium-size cetothere 4-5 meters in length. It differs from other members of
Cetotherioidea and is most similar to
121:
1843:
423:
1753:
1783:
1748:
1815:
1548:
1576:
896:
820:
773:
1415:
1230:
1165:
1019:
906:
732:
203:
1711:
1640:
1346:
1310:
871:
415:
406:
Gol'din P. (2018) New
Paratethyan dwarf baleen whales mark the origin of cetotheres. PeerJ 6:e5800
1142:
1119:
1000:
580:
1758:
1691:
1661:
1613:
1565:
1538:
1290:
1095:
478:
322:
by
Riabinin (1934) and also found in the Middle Miocene of the Russian Caucasus, was referred to
116:
1671:
1172:
1151:
1105:
1045:
570:
549:
529:
1820:
1202:
1035:
970:
591:
559:
1681:
1499:
1369:
373:
M. C. McKenna and S. K. Bell. 1997. Classification of
Mammals Above the Species Level 1-640.
282:
602:
1651:
1596:
1526:
1390:
841:
705:
298:, and the cladistic analyses of Gol'din & Steeman (2015) and Gol'din (2018) recovered
270:
from the middle
Miocene (late Burdigalian to early Serravallian) of the Russian Caucasus.
263:
1721:
1515:
1209:
800:
786:
339:
1837:
1376:
1280:
1264:
990:
661:
654:
291:
68:
1603:
1555:
1484:
1444:
1383:
1355:
1253:
813:
675:
617:
512:
499:
447:
267:
240:
193:
1806:
1701:
1454:
1362:
1321:
1186:
980:
848:
633:
173:
43:
392:
1623:
1474:
1464:
1434:
1404:
1195:
1158:
1128:
1085:
943:
923:
880:
721:
714:
691:
684:
640:
626:
88:
53:
1630:
1586:
1246:
1239:
1216:
1075:
1065:
960:
950:
933:
834:
827:
752:
539:
133:
93:
37:
1800:
1397:
1332:
1055:
759:
745:
466:
153:
83:
78:
63:
58:
48:
1424:
913:
484:
260:
183:
98:
73:
30:
647:
472:
460:
407:
163:
143:
1777:
314:, and Neobalaenidae. The juvenile specimen VSEGEI 2401, referred to
1781:
497:
419:
306:
1717:
1707:
1697:
1687:
1677:
1667:
1657:
1647:
1636:
1619:
1609:
1592:
1582:
1572:
1561:
1544:
1534:
1511:
1480:
1470:
1460:
1450:
1440:
1430:
1343:
1328:
1317:
1306:
1296:
1286:
1276:
1227:
1183:
1139:
1116:
1101:
1091:
1081:
1071:
1061:
1051:
1041:
1031:
996:
986:
976:
966:
956:
939:
929:
919:
902:
892:
797:
770:
702:
672:
614:
598:
587:
576:
566:
555:
545:
535:
525:
1790:
1525:
1498:
1414:
1342:
1263:
1226:
1182:
1138:
1115:
1018:
879:
870:
796:
769:
731:
701:
671:
613:
510:
431:
281:was described by the Georgian paleontologist
8:
393:https://doi.org/10.1371/journal.pone.0135500
402:
400:
1778:
1504:
1269:
1024:
885:
876:
805:
778:
737:
518:
507:
494:
438:
424:
416:
20:
357:
7:
408:https://doi.org/10.7717/peerj.5800
14:
1762:
1757:
1752:
1747:
120:
1:
1864:Fossil taxa described in 1984
1859:Prehistoric cetacean genera
1880:
1854:Miocene genus extinctions
1849:Miocene mammals of Europe
1745:
1507:
1272:
1027:
888:
808:
781:
740:
521:
506:
493:
455:
233:
228:
117:Scientific classification
115:
23:
450:and their extinct allies
16:Extinct genus of whales
304:Parietobalaena securis
1816:Paleobiology Database
326:by Gol'din (2018).
1692:Plesiobalaenoptera
1539:Archaebalaenoptera
1844:Miocene cetaceans
1831:
1830:
1784:Taxon identifiers
1775:
1774:
1771:
1770:
1743:
1742:
1739:
1738:
1735:
1734:
1731:
1730:
1549:Archaeschrichtius
1494:
1493:
1014:
1013:
1010:
1009:
866:
865:
862:
861:
858:
857:
252:
251:
246:
224:
1871:
1824:
1823:
1811:
1810:
1809:
1779:
1766:
1761:
1756:
1751:
1719:
1709:
1699:
1689:
1682:Parabalaenoptera
1679:
1669:
1659:
1649:
1638:
1621:
1611:
1594:
1584:
1574:
1563:
1546:
1536:
1513:
1505:
1500:Balaenopteroidea
1482:
1472:
1462:
1452:
1442:
1432:
1345:
1330:
1319:
1308:
1298:
1288:
1278:
1270:
1229:
1185:
1141:
1118:
1103:
1093:
1083:
1073:
1063:
1053:
1043:
1033:
1025:
998:
988:
978:
968:
958:
941:
931:
921:
904:
894:
886:
877:
806:
799:
779:
772:
738:
704:
674:
616:
600:
589:
578:
568:
557:
547:
537:
527:
519:
508:
495:
440:
433:
426:
417:
410:
404:
395:
389:
383:
380:
374:
371:
365:
362:
283:Guram Mchedlidze
244:
223:Mchedlidze, 1984
222:
215:
125:
124:
110:
40:
29:Temporal range:
21:
1879:
1878:
1874:
1873:
1872:
1870:
1869:
1868:
1834:
1833:
1832:
1827:
1819:
1814:
1805:
1804:
1799:
1786:
1776:
1767:
1727:
1652:Miobalaenoptera
1597:Eschrichtioides
1577:Cetoteriophanes
1527:Balaenopteridae
1521:
1490:
1410:
1391:Mithridatocetus
1338:
1259:
1222:
1178:
1134:
1111:
1006:
897:Antwerpibalaena
854:
821:Micromysticetus
792:
774:Eomysticetoidea
765:
727:
706:Mammalodontidae
697:
667:
609:
515:
502:
489:
451:
444:
414:
413:
405:
398:
390:
386:
381:
377:
372:
368:
363:
359:
354:
332:
276:
245:Mchedlize, 1984
221:
213:
119:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
36:16.2–15.3
35:
34:
27:
17:
12:
11:
5:
1877:
1875:
1867:
1866:
1861:
1856:
1851:
1846:
1836:
1835:
1829:
1828:
1826:
1825:
1812:
1796:
1794:
1788:
1787:
1782:
1773:
1772:
1769:
1768:
1746:
1744:
1741:
1740:
1737:
1736:
1733:
1732:
1729:
1728:
1726:
1725:
1722:Protororqualus
1715:
1705:
1695:
1685:
1675:
1665:
1655:
1645:
1634:
1627:
1617:
1607:
1600:
1590:
1580:
1570:
1559:
1552:
1542:
1531:
1529:
1523:
1522:
1520:
1519:
1516:Eobalaenoptera
1508:
1502:
1496:
1495:
1492:
1491:
1489:
1488:
1478:
1468:
1458:
1448:
1438:
1428:
1420:
1418:
1416:Herpetocetinae
1412:
1411:
1409:
1408:
1401:
1394:
1387:
1380:
1373:
1366:
1359:
1351:
1349:
1340:
1339:
1337:
1336:
1326:
1315:
1304:
1294:
1284:
1273:
1267:
1261:
1260:
1258:
1257:
1250:
1243:
1235:
1233:
1231:Tranatocetidae
1224:
1223:
1221:
1220:
1213:
1210:Parietobalaena
1206:
1199:
1191:
1189:
1180:
1179:
1177:
1176:
1169:
1166:Plesiocetopsis
1162:
1155:
1147:
1145:
1136:
1135:
1133:
1132:
1124:
1122:
1113:
1112:
1110:
1109:
1099:
1089:
1079:
1069:
1059:
1049:
1039:
1028:
1022:
1020:Thalassotherii
1016:
1015:
1012:
1011:
1008:
1007:
1005:
1004:
994:
984:
974:
964:
954:
947:
937:
927:
917:
910:
907:Archaeobalaena
900:
889:
883:
874:
868:
867:
864:
863:
860:
859:
856:
855:
853:
852:
845:
838:
831:
824:
817:
809:
803:
801:Eomysticetidae
794:
793:
791:
790:
787:Cetotheriopsis
782:
776:
767:
766:
764:
763:
756:
749:
741:
735:
733:Chaeomysticeti
729:
728:
726:
725:
718:
710:
708:
699:
698:
696:
695:
688:
680:
678:
669:
668:
666:
665:
658:
651:
644:
637:
630:
622:
620:
611:
610:
608:
607:
596:
585:
574:
564:
553:
543:
533:
522:
516:
511:
504:
503:
498:
491:
490:
488:
487:
481:
475:
469:
463:
456:
453:
452:
445:
443:
442:
435:
428:
420:
412:
411:
396:
384:
375:
366:
356:
355:
353:
350:
346:Parietobalaena
340:Parietobalaena
331:
328:
290:to the family
275:
272:
250:
249:
248:
247:
231:
230:
226:
225:
211:
207:
206:
204:Cetotherioidea
201:
197:
196:
191:
187:
186:
181:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
141:
137:
136:
131:
127:
126:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
41:
28:
15:
13:
10:
9:
6:
4:
3:
2:
1876:
1865:
1862:
1860:
1857:
1855:
1852:
1850:
1847:
1845:
1842:
1841:
1839:
1822:
1817:
1813:
1808:
1802:
1798:
1797:
1795:
1793:
1789:
1785:
1780:
1765:
1760:
1755:
1750:
1724:
1723:
1716:
1714:
1713:
1712:Praemegaptera
1706:
1704:
1703:
1696:
1694:
1693:
1686:
1684:
1683:
1676:
1674:
1673:
1666:
1664:
1663:
1656:
1654:
1653:
1646:
1643:
1642:
1641:Megapteropsis
1635:
1633:
1632:
1628:
1626:
1625:
1618:
1616:
1615:
1608:
1606:
1605:
1601:
1599:
1598:
1591:
1589:
1588:
1581:
1579:
1578:
1571:
1568:
1567:
1560:
1558:
1557:
1553:
1551:
1550:
1543:
1541:
1540:
1533:
1532:
1530:
1528:
1524:
1518:
1517:
1510:
1509:
1506:
1503:
1501:
1497:
1487:
1486:
1479:
1477:
1476:
1469:
1467:
1466:
1459:
1457:
1456:
1449:
1447:
1446:
1439:
1437:
1436:
1429:
1427:
1426:
1422:
1421:
1419:
1417:
1413:
1407:
1406:
1402:
1400:
1399:
1395:
1393:
1392:
1388:
1386:
1385:
1381:
1379:
1378:
1377:Eucetotherium
1374:
1372:
1371:
1367:
1365:
1364:
1360:
1358:
1357:
1353:
1352:
1350:
1348:
1347:Cetotheriinae
1341:
1335:
1334:
1327:
1324:
1323:
1316:
1313:
1312:
1311:Palaeobalaena
1305:
1303:
1302:
1295:
1293:
1292:
1285:
1283:
1282:
1281:Cephalotropis
1275:
1274:
1271:
1268:
1266:
1265:Cetotheriidae
1262:
1256:
1255:
1251:
1249:
1248:
1244:
1242:
1241:
1237:
1236:
1234:
1232:
1225:
1219:
1218:
1214:
1212:
1211:
1207:
1205:
1204:
1200:
1198:
1197:
1193:
1192:
1190:
1188:
1181:
1175:
1174:
1170:
1168:
1167:
1163:
1161:
1160:
1156:
1154:
1153:
1149:
1148:
1146:
1144:
1137:
1131:
1130:
1126:
1125:
1123:
1121:
1114:
1108:
1107:
1100:
1098:
1097:
1090:
1088:
1087:
1080:
1078:
1077:
1070:
1068:
1067:
1060:
1058:
1057:
1050:
1048:
1047:
1040:
1038:
1037:
1030:
1029:
1026:
1023:
1021:
1017:
1003:
1002:
995:
993:
992:
991:Peripolocetus
985:
983:
982:
975:
973:
972:
965:
963:
962:
955:
953:
952:
948:
946:
945:
938:
936:
935:
928:
926:
925:
918:
916:
915:
911:
909:
908:
901:
899:
898:
891:
890:
887:
884:
882:
878:
875:
873:
872:Balaenomorpha
869:
851:
850:
846:
844:
843:
839:
837:
836:
832:
830:
829:
825:
823:
822:
818:
816:
815:
811:
810:
807:
804:
802:
795:
789:
788:
784:
783:
780:
777:
775:
768:
762:
761:
757:
755:
754:
750:
748:
747:
743:
742:
739:
736:
734:
730:
724:
723:
719:
717:
716:
712:
711:
709:
707:
700:
694:
693:
689:
687:
686:
682:
681:
679:
677:
670:
664:
663:
662:Willungacetus
659:
657:
656:
655:Morawanocetus
652:
650:
649:
645:
643:
642:
638:
636:
635:
631:
629:
628:
624:
623:
621:
619:
612:
605:
604:
597:
594:
593:
586:
583:
582:
575:
573:
572:
565:
562:
561:
554:
552:
551:
544:
542:
541:
534:
532:
531:
524:
523:
520:
517:
514:
509:
505:
501:
496:
492:
486:
482:
480:
476:
474:
470:
468:
464:
462:
458:
457:
454:
449:
448:baleen whales
441:
436:
434:
429:
427:
422:
421:
418:
409:
403:
401:
397:
394:
388:
385:
379:
376:
370:
367:
361:
358:
351:
349:
347:
342:
341:
336:
329:
327:
325:
321:
317:
313:
312:Cephalotropis
309:
305:
302:as sister to
301:
297:
293:
292:Cetotheriidae
289:
284:
280:
273:
271:
269:
265:
262:
258:
257:
242:
238:
235:
234:
232:
227:
220:
219:
212:
209:
208:
205:
202:
200:Superfamily:
199:
198:
195:
192:
189:
188:
185:
182:
179:
178:
175:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
145:
142:
139:
138:
135:
132:
129:
128:
123:
118:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
1792:Otradnocetus
1791:
1720:
1710:
1700:
1690:
1680:
1670:
1660:
1650:
1639:
1629:
1622:
1612:
1604:Eschrichtius
1602:
1595:
1585:
1575:
1564:
1556:Balaenoptera
1554:
1547:
1537:
1514:
1485:Piscobalaena
1483:
1473:
1463:
1453:
1445:Herpetocetus
1443:
1433:
1423:
1403:
1396:
1389:
1384:Kurdalagonus
1382:
1375:
1368:
1361:
1356:Brandtocetus
1354:
1331:
1320:
1309:
1301:Otradnocetus
1300:
1299:
1289:
1279:
1254:Tranatocetus
1252:
1245:
1238:
1215:
1208:
1201:
1194:
1171:
1164:
1157:
1150:
1143:Diorocetidae
1127:
1120:Aglaocetidae
1104:
1094:
1084:
1074:
1064:
1054:
1044:
1034:
1001:Protobalaena
999:
989:
979:
969:
959:
949:
942:
932:
922:
912:
905:
895:
847:
840:
833:
826:
819:
814:Eomysticetus
812:
785:
758:
751:
744:
720:
713:
690:
683:
676:Llanocetidae
660:
653:
646:
639:
632:
625:
618:Aetiocetidae
601:
590:
581:Siphonocetus
579:
569:
558:
548:
538:
528:
483:Infraorder:
479:Artiodactyla
387:
378:
369:
360:
345:
338:
335:Otradnocetus
334:
333:
324:Otradnocetus
323:
319:
315:
311:
307:
303:
300:Otradnocetus
299:
296:Otradnocetus
295:
288:Otradnocetus
287:
279:Otradnocetus
278:
277:
268:baleen whale
256:Otradnocetus
255:
254:
253:
236:
218:Otradnocetus
217:
216:
180:Infraorder:
174:Artiodactyla
25:Otradnocetus
24:
18:
1702:Plesiocetus
1662:Nehalaennia
1614:Gricetoides
1566:Burtinopsis
1455:Metopocetus
1363:Cetotherium
1322:Titanocetus
1291:Journocetus
1187:Pelocetidae
1096:Tiphyocetus
981:Morenocetus
849:Yamatocetus
634:Ashorocetus
330:Description
316:Cetotherium
237:O. virodovi
190:Parvorder:
1838:Categories
1672:Notiocetus
1624:Incakujira
1475:Nannocetus
1465:Miocaperea
1435:Herentalia
1405:Zygiocetus
1196:Cophocetus
1173:Thinocetus
1159:Diorocetus
1152:Amphicetus
1129:Aglaocetus
1106:Uranocetus
1086:Taikicetus
1046:Isanacetus
944:Balaenotus
924:Balaenella
881:Balaenidae
722:Mammalodon
715:Janjucetus
692:Mystacodon
685:Llanocetus
641:Chonecetus
627:Aetiocetus
571:Piscocetus
550:Imerocetus
530:Borealodon
446:Genera of
352:References
1807:Q60791091
1631:Megaptera
1587:Diunatans
1370:Ciuciulea
1247:Mixocetus
1240:Mesocetus
1217:Pelocetus
1203:Halicetus
1076:Pinocetus
1066:Mauicetus
1036:Hibacetus
971:Mesoteras
961:Idiocetus
951:Eubalaena
934:Balaenula
835:Tokarahia
828:Tohoraata
753:Sitsqwayk
592:Tretulias
540:Coronodon
513:Mysticeti
500:Mysticeti
459:Kingdom:
308:Pinocetus
286:assigned
194:Mysticeti
140:Kingdom:
134:Eukaryota
1801:Wikidata
1398:Vampalus
1333:Tiucetus
1056:Isocetus
760:Whakakai
746:Horopeta
473:Mammalia
467:Chordata
465:Phylum:
461:Animalia
274:Taxonomy
229:Species
164:Mammalia
154:Chordata
150:Phylum:
144:Animalia
130:Domain:
1425:Caperea
914:Balaena
842:Waharoa
560:Mioceta
485:Cetacea
477:Order:
471:Class:
261:extinct
210:Genus:
184:Cetacea
170:Order:
160:Class:
105:↓
31:Miocene
648:Fucaia
320:mayeri
259:is an
1821:63928
603:Ulias
318:aff.
264:genus
241:type
44:PreꞒ
266:of
1840::
1818::
1803::
399:^
310:,
94:Pg
38:Ma
33:,
1718:†
1708:†
1698:†
1688:†
1678:†
1668:†
1658:†
1648:†
1644:?
1637:†
1620:†
1610:†
1593:†
1583:†
1573:†
1569:?
1562:†
1545:†
1535:†
1512:†
1481:†
1471:†
1461:†
1451:†
1441:†
1431:†
1344:†
1329:†
1325:?
1318:†
1314:?
1307:†
1297:†
1287:†
1277:†
1228:†
1184:†
1140:†
1117:†
1102:†
1092:†
1082:†
1072:†
1062:†
1052:†
1042:†
1032:†
997:†
987:†
977:†
967:†
957:†
940:†
930:†
920:†
903:†
893:†
798:†
771:†
703:†
673:†
615:†
606:?
599:†
595:?
588:†
584:?
577:†
567:†
563:?
556:†
546:†
536:†
526:†
439:e
432:t
425:v
243:)
239:(
214:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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