470:
492:
86:
681:
33:
57:
947:
227:
596:, though they may emerge during the day. They are swift runners, darting away to their burrows at the slightest sign of danger, even when the intruder sighted is still 30 ft (9.1 m) away. However, they can be approached much more closely at night, though they may still try to escape if illuminated.
537:(appendages modified into sexual organs) of the male is stem-like. Its cross-section has three sides on the base ending in a slightly curving tip. A broad and flat palp is present. The covering (operculum) of the female genital opening is oriented lengthwise with a thick straight rim slanting backwards.
603:
exhibit elaborate territorial displays. Like other ghost crabs, they produce sound and vibration by rapping their larger claws against the ground. They may also display "dances" on the approach of another ghost crab. Beginning with a rearing posture, the males conduct increasingly complex sideways
308:
reaches a length of 41 mm (1.6 in) and a width of 50 mm (2.0 in). They are a mottled brown to yellow in coloration. Like other ghost crabs, one of their claws is much larger than the other. Their eyestalks are large and elongated, tipped with prolongations at the tip known as
487:
The eyestalks are large and swollen with the cornea occupying most of the bottom half. The eyestalks exhibit prolongations (styles) on the tips like some other members of the genus. The styles may be absent or much shorter in juvenile specimens, as it only starts growing when the crab is around
465:
in shape, with the rear end distinctly narrower than the front. It is wider than it is long and covered with scattered rough bumps (tubercles). The upper margins of the eye sockets slant forwards on the inner half and backwards for the outer half. The edges of the eye sockets are rectangular.
521:
is much bigger than the other. The palm of the larger cheliped is long with a rough bumpy texture on the upper surface. The upper edges of the palm are covered with small bumps while the bottom edge is serrated. The inner surface of the palm of the larger claw in both sexes features
1049:
544:
are a mottled yellow to brown in coloration. Males have been observed to display brighter yellow colors, especially on their legs. Juveniles have almost perfect
1062:
1149:
1134:
1023:
1144:
1075:
896:
461:
in adults ranges from 24 to 41 mm (0.94 to 1.61 in) in length, and 26 to 50 mm (1.0 to 2.0 in) in width. It is almost
1139:
1101:
811:
951:
469:
530:, the stridulating ridge is composed of a row of 23 to 28 tubercles. The smaller cheliped tapers towards a pointed end.
997:
1067:
526:(sound-producing) ridges, which is important for identifying different species within the subfamily Ocypodinae. In
85:
491:
965:
641:
359:
348:
226:
1002:
888:
267:
200:
768:
1129:
1106:
799:
368:
343:. In 1852, he again described the same species from adult specimens recovered from the same area as
738:
636:
433:
328:
216:
686:
647:
571:
401:, due to the fact that both species possess elongations of their eyestalks (styles). The variety
351:
244:
80:
371:
for the entirety of the 20th century. And most literature concerning the species refer to it as
1054:
1088:
1010:
892:
665:
659:
653:
410:
1093:
882:
379:
Katsushi Sakai and German carcinologist
Michael Türkay discovered that the type specimens of
925:
862:
A. K. Nagabhushanam (1972). "Studies on the marine intertidal ecology of the Orissa coast".
440:
56:
722:
632:
567:
310:
301:
73:
1123:
499:
376:
332:
309:
styles. They are common inhabitants of open sandy beaches, living in burrows in the
1015:
620:
616:
523:
495:
297:
293:
127:
32:
680:
988:
1036:
336:
147:
548:
camouflage, making it very difficult to pick them out from their surroundings.
929:
839:
K. G. Raja Bai Naidu (1951). "Some stages in the development and bionomics of
676:
604:
movements ending by running around the intruder in circles until it retreats.
557:
443:
436:
304:. They are relatively large ghost crabs with a somewhat trapezoidal body. The
289:
167:
387:
belonged to the same species. They restored the validity of the earlier name
695:
628:
593:
575:
514:
462:
325:
117:
97:
946:
982:
586:
510:
458:
305:
137:
65:
1028:
578:
545:
534:
477:
428:
285:
177:
44:
1041:
582:
107:
959:
624:
490:
481:
468:
424:
340:
48:
1080:
157:
963:
912:
David
Clayton (2008). "Singing and dancing in the ghost crab
884:
Ocracoke Wild: A Naturalist's Year on an Outer Banks Island
64:
An 1882 illustration of the eyestalks and the front of the
881:
Pat Garber (2006). "Phantoms in the Surf: Ghost Crabs".
845:
Proceedings of the Indian
Academy of Sciences, Section B
635:. They are one of six ghost crab species found in the
509:
Like other ghost crabs, one of the claw appendages (
972:
864:Proceedings of the Indian National Science Academy
335:consisted only of two juvenile individuals from
767:Sakai, Katsushi & Türkay, Michael (2013).
8:
457:are large ghost crabs with deep bodies. The
960:
225:
55:
31:
20:
781:Memoirs of the Queensland Museum – Nature
727:in the collections of the British Museum"
834:
832:
830:
828:
762:
760:
758:
756:
754:
752:
750:
748:
716:
714:
712:
708:
397:has also been frequently confused with
731:Annals and Magazine of Natural History
773:with the description of a new genus,
488:12 mm (0.47 in) in length.
7:
916:(Crustacea, Decapoda, Ocypodidae)".
409:described by the American zoologist
347:. In 1880, the American zoologist
324:was first described by the French
14:
1150:Taxa named by Henri Milne-Edwards
777:(Crustacea: Decapoda: Brachyura)"
945:
812:World Register of Marine Species
679:
570:of open sandy beaches. They are
498:ridge on the palm of the larger
413:is now known to be a synonym of
84:
1135:Crustaceans of the Indian Ocean
566:live in deep burrows near the
1:
1145:Crustaceans described in 1837
619:. They can be found from the
1166:
918:Journal of Natural History
555:
930:10.1080/00222930701840530
250:
243:
233:
224:
206:
199:
81:Scientific classification
79:
63:
54:
39:
30:
23:
1140:Crustaceans of Sri Lanka
721:Edward J. Miers (1882).
562:Like other ghost crabs,
375:. In 2013, the Japanese
804:H. Milne Edwards, 1837"
769:"Revision of the genus
642:Ocypode ceratophthalma
615:are restricted to the
506:
484:
415:Ocypode ceratophthalma
399:Ocypode ceratophthalma
360:Ocypode ceratophthalma
349:John Sterling Kingsley
331:in 1837. However, his
258:H. Milne-Edwards, 1852
556:Further information:
494:
472:
956:at Wikimedia Commons
798:Peter Davie (2013).
699:- the semaphore crab
363:. As a result, only
974:Ocypode brevicornis
953:Ocypode brevicornis
914:Ocypode platytarsus
841:Ocypoda platytarsis
802:Ocypode brevicornis
723:"On the species of
639:, the others being
637:Indian subcontinent
613:Ocypode brevicornis
601:Ocypode brevicornis
592:They are primarily
564:Ocypode brevicornis
542:Ocypode brevicornis
528:Ocypode brevicornis
519:Ocypode brevicornis
504:Ocypode brevicornis
474:Ocypode brevicornis
455:Ocypode brevicornis
421:Ocypode brevicornis
403:Ocypode brevicornis
395:Ocypode brevicornis
389:Ocypode brevicornis
385:Ocypode platytarsis
381:Ocypode brevicornis
373:Ocypode platytarsis
365:Ocypode platytarsis
355:Ocypode brevicornis
345:Ocypode platytarsis
329:Henri Milne-Edwards
322:Ocypode brevicornis
281:Ocypode brevicornis
254:Ocypode platytarsis
235:Ocypode brevicornis
210:Ocypode brevicornis
192:O. brevicornis
70:Ocypode brevicornis
41:Ocypode brevicornis
25:Ocypode brevicornis
891:. pp. 94–98.
889:Parkway Publishers
687:Crustaceans portal
648:Ocypode cordimanus
589:on small animals.
507:
485:
432:of the ghost crab
237:distribution range
1117:
1116:
1089:Open Tree of Life
966:Taxon identifiers
950:Media related to
898:978-1-933251-31-8
666:Ocypode rotundata
660:Ocypode pallidula
654:Ocypode macrocera
411:James Dwight Dana
277:
276:
271:
259:
1157:
1110:
1109:
1097:
1096:
1084:
1083:
1071:
1070:
1058:
1057:
1045:
1044:
1032:
1031:
1019:
1018:
1006:
1005:
993:
992:
991:
961:
949:
934:
933:
924:(3–4): 141–155.
909:
903:
902:
878:
872:
871:
859:
853:
852:
836:
823:
822:
820:
818:
795:
789:
788:
764:
743:
742:
718:
689:
684:
683:
367:was regarded as
266:
263:Ocypode neglecta
257:
229:
217:H. Milne-Edwards
212:
89:
88:
59:
35:
21:
1165:
1164:
1160:
1159:
1158:
1156:
1155:
1154:
1120:
1119:
1118:
1113:
1105:
1100:
1092:
1087:
1079:
1074:
1066:
1061:
1053:
1048:
1040:
1035:
1027:
1022:
1014:
1009:
1001:
996:
987:
986:
981:
968:
942:
937:
911:
910:
906:
899:
880:
879:
875:
870:(3–4): 308–315.
861:
860:
856:
838:
837:
826:
816:
814:
797:
796:
792:
766:
765:
746:
720:
719:
710:
706:
685:
678:
675:
633:Nicobar Islands
610:
568:intertidal zone
560:
554:
452:
423:belongs to the
319:
311:intertidal zone
302:Nicobar Islands
265:
256:
238:
220:
214:
208:
195:
83:
74:Edward J. Miers
17:
16:Species of crab
12:
11:
5:
1163:
1161:
1153:
1152:
1147:
1142:
1137:
1132:
1122:
1121:
1115:
1114:
1112:
1111:
1098:
1085:
1072:
1059:
1046:
1033:
1020:
1007:
994:
978:
976:
970:
969:
964:
958:
957:
941:
940:External links
938:
936:
935:
904:
897:
873:
854:
824:
790:
744:
707:
705:
702:
701:
700:
691:
690:
674:
671:
609:
606:
553:
550:
451:
448:
333:type specimens
318:
315:
292:native to the
275:
274:
273:
272:
260:
248:
247:
241:
240:
231:
230:
222:
221:
215:
204:
203:
197:
196:
189:
187:
183:
182:
175:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
135:
131:
130:
125:
121:
120:
115:
111:
110:
105:
101:
100:
95:
91:
90:
77:
76:
61:
60:
52:
51:
37:
36:
28:
27:
15:
13:
10:
9:
6:
4:
3:
2:
1162:
1151:
1148:
1146:
1143:
1141:
1138:
1136:
1133:
1131:
1128:
1127:
1125:
1108:
1103:
1099:
1095:
1090:
1086:
1082:
1077:
1073:
1069:
1064:
1060:
1056:
1051:
1047:
1043:
1038:
1034:
1030:
1025:
1021:
1017:
1012:
1008:
1004:
999:
995:
990:
984:
980:
979:
977:
975:
971:
967:
962:
955:
954:
948:
944:
943:
939:
931:
927:
923:
919:
915:
908:
905:
900:
894:
890:
886:
885:
877:
874:
869:
865:
858:
855:
850:
846:
842:
835:
833:
831:
829:
825:
813:
809:
805:
803:
794:
791:
787:(2): 665–793.
786:
782:
778:
776:
772:
763:
761:
759:
757:
755:
753:
751:
749:
745:
740:
736:
732:
728:
726:
717:
715:
713:
709:
703:
698:
697:
693:
692:
688:
682:
677:
672:
670:
668:
667:
662:
661:
656:
655:
650:
649:
644:
643:
638:
634:
630:
626:
622:
618:
614:
607:
605:
602:
597:
595:
590:
588:
584:
580:
577:
573:
569:
565:
559:
551:
549:
547:
543:
538:
536:
531:
529:
525:
520:
516:
512:
505:
501:
497:
493:
489:
483:
479:
475:
471:
467:
464:
460:
456:
449:
447:
445:
442:
438:
435:
431:
430:
426:
422:
418:
416:
412:
408:
404:
400:
396:
392:
390:
386:
382:
378:
377:carcinologist
374:
370:
366:
362:
361:
356:
353:
350:
346:
342:
338:
334:
330:
327:
323:
316:
314:
312:
307:
303:
299:
295:
291:
287:
283:
282:
269:
264:
261:
255:
252:
251:
249:
246:
242:
236:
232:
228:
223:
218:
213:
211:
205:
202:
201:Binomial name
198:
194:
193:
188:
185:
184:
181:
180:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
139:
136:
133:
132:
129:
126:
123:
122:
119:
116:
113:
112:
109:
106:
103:
102:
99:
96:
93:
92:
87:
82:
78:
75:
71:
67:
62:
58:
53:
50:
46:
42:
38:
34:
29:
26:
22:
19:
973:
952:
921:
917:
913:
907:
883:
876:
867:
863:
857:
848:
844:
840:
817:November 11,
815:. Retrieved
807:
801:
793:
784:
780:
774:
770:
734:
730:
724:
694:
664:
658:
652:
646:
640:
621:Gulf of Oman
617:Indian Ocean
612:
611:
608:Distribution
600:
598:
591:
563:
561:
541:
539:
532:
527:
524:stridulating
518:
513:, the first
508:
503:
496:Stridulating
486:
473:
454:
453:
427:
420:
419:
414:
407:longicornuta
406:
402:
398:
394:
393:
388:
384:
380:
372:
364:
358:
354:
344:
321:
320:
298:Gulf of Oman
294:Indian Ocean
280:
279:
278:
262:
253:
234:
209:
207:
191:
190:
178:
154:Infraorder:
128:Malacostraca
69:
40:
24:
18:
1130:Ocypodoidea
1037:iNaturalist
851:(1): 32–40.
775:Hoplocypode
585:as well as
572:generalists
463:trapezoidal
450:Description
352:synonymized
337:Pondicherry
296:, from the
239:(in green)
148:Pleocyemata
1124:Categories
704:References
631:, and the
576:scavenging
558:Ghost crab
533:The first
444:Ocypodidae
437:Ocypodinae
290:ghost crab
168:Ocypodidae
144:Suborder:
118:Arthropoda
989:Q13908723
696:Heloecius
629:Sri Lanka
599:Males of
594:nocturnal
517:pair) of
515:pereiopod
511:chelipeds
434:subfamily
326:zoologist
186:Species:
158:Brachyura
104:Kingdom:
98:Eukaryota
1055:11878907
983:Wikidata
673:See also
459:carapace
317:Taxonomy
306:carapace
245:Synonyms
164:Family:
138:Decapoda
114:Phylum:
108:Animalia
94:Domain:
66:carapace
1094:2981541
1068:1977103
1029:5971892
771:Ocypode
725:Ocypoda
587:preying
579:carrion
552:Ecology
546:cryptic
535:gonopod
478:Chennai
439:in the
429:Ocypode
300:to the
286:species
270:, 1894a
268:Ortmann
179:Ocypode
174:Genus:
134:Order:
124:Class:
45:Chennai
1107:444983
1081:444983
1042:361850
1003:934188
895:
663:, and
583:debris
540:Adult
441:family
219:, 1837
1102:WoRMS
1050:IRMNG
1016:48LW2
808:WoRMS
741:–317.
737:(5):
625:India
482:India
425:genus
405:var.
369:valid
357:with
341:India
284:is a
49:India
43:from
1076:OBIS
1063:NCBI
1024:GBIF
998:BOLD
893:ISBN
819:2013
581:and
500:claw
383:and
1011:CoL
926:doi
868:38B
843:".
739:304
623:to
502:of
476:in
288:of
72:by
68:of
1126::
1104::
1091::
1078::
1065::
1052::
1039::
1026::
1013::
1000::
985::
922:42
920:.
887:.
866:.
849:33
847:.
827:^
810:.
806:.
785:56
783:.
779:.
747:^
733:.
729:.
711:^
669:.
657:,
651:,
645:,
627:,
574:,
480:,
446:.
417:.
391:.
339:,
313:.
47:,
932:.
928::
901:.
821:.
800:"
735:5
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
